Interesting Facts
Common Names
Common Names in English:
Flexible Ceanothus
Description
Family Rhamnaceae
Deciduous or evergreen
, often thorny trees
, shrubs
, woody climbers
, or lianas, rarely herbs. Leaves simple
, petiolate
, alternate or opposite, pinnately veined or 3-5-veined, entire to serrate, sometimes much reduced; stipules small, caducous
or persistent
, sometimes transformed into spines. Flowers yellowish to greenish, rarely brightly colored
, small, bisexual
or unisexual
, rarely polygamous, (4 or) 5-merous, hypogynous to epigynous
, in mostly axillary
, sessile or pedunculate
cymes, or reduced to few in fascicles. Calyx tube
patelliform
or hemispherical to tubular
, sometimes absent, at rim
with calyx, corolla, and stamens; sepals 4 or 5, valvate
in bud, triangular, erect
or ± recurved during anthesis
, adaxially often distinctly keeled
, alternate with petals. Petals 4 or 5, rarely absent, usually smaller than sepals, concave
or hooded
, rarely nearly flat, often shortly clawed. Stamens 4 or 5, antepetalous
and often ± enclosed by petals; filaments
thin, adnate
to bases
of petals; anthers
minute, versatile or not, 2(or 4) -celled, dehiscing by longitudinal
slits, usually introrse
. Disk intrastaminal
, nectariferous
, thin to ± fleshy
, entire or lobed
, glabrous
or rarely pubescent
, free
from ovary or tightly surrounding it, or adnate to calyx tube. Ovary superior to inferior, (1 or) 2-4-loculed, with 1(or 2) ovules per locule; ovules anatropous
, basal and erect; styles simple or ± deeply 3-lobed or 3-cleft. Fruit either an indehiscent, rarely explosively dehiscent
, sometimes winged
, schizocarpic capsule, or a ± fleshy drupe with 1-4 indehiscent, rarely dehiscent, pyrenes (stones
) . Seeds with thin, oily albumen, sometimes exalbuminous
; embryo large, oily, straight or rarely bent.
About 50 genera and more than 900 species: almost cosmopolitan
, mainly in subtropical
to tropical
areas; 13 genera and 137 species (82 endemic, one introduced
) in China.
Former classifications usually placed Rhamnaceae in the Rhamnales, together with Vitaceae and Leeaceae (Suessenguth in Engler & Prantl, Nat. Pflanzenfam., ed. 2, 20d. 1953), or together with Elaeagnaceae (Thorne, Bot. Rev. 58: 225-348. 1992) . Orders
such as Celastrales, Urticales, and Euphorbiales have often been considered as closely related groups. Recent analyses of DNA sequences strongly supported including the family
in the Rosales, beside the closest relatives Barbeyaceae and Dirachmaceae (see Angiosperm Phylogeny Group II, Bot. J. Linn. Soc. 141: 399-436. 2003) . Suessenguth (loc. cit.
) grouped the family into five tribes
, mainly characterized by fruit characters. Richardson et al.
(Kew Bull
. 55: 311-340. 2000; Amer. J. Bot. 87: 1309-1324. 2000) revised this tribal classification on the basis of a phylogenetic
analysis using rbcL and trnL-F sequences of the plastid genome. Now 11 tribes are recognized, of which four are represented in the Flora
area.
The bark
, leaves, and fruit of several species of Rhamnus have been used as laxatives
, notably R. cathartica and R. frangula. Diverse
Old World species of Rhamnus provide yellow and green dyes as well as drugs. Timber of Alphitonia, Colubrina, Hovenia, and Ziziphus species is used for construction, fine furniture, carving, lathework, and musical instruments. Many Ziziphus species yield edible fruit; among them, Z. jujuba (Chinese jujube) and Z. mauritiana (Indian jujube) are cultivated on a commercial
scale. Hovenia dulcis is also grown for its edible, fleshy inflorescence stalks
. Species of Hovenia, Paliurus, and Rhamnus are cultivated as ornamentals
.[1]
Physical Description
Habit: Shrub • Growth Form: Colonizing • Shape and Orientation: Prostrate
Flowers: Bloom Period: Early Summer • Flower Color: Blue • Flower Conspicuous: Yes
Seeds: Seed Spread Rate: None • Seedling Vigor: Low • Fruit/Seed Abundance: Low • Fruit/Seed Color: Brown • Fruit/Seed Conspicuous: No • Cold Stratification Required: No
Foliage: Foliage Color: Dark Green • Foliage Porosity Summer: Dense • Foliage Porosity Winter: Dense • Foliage Texture: Coarse • Fall Conspicuous: Yes • Leaf Retention: Yes
Size/Age/Growth
Active Growth Period: Spring and Summer • Growth Rate: Moderate • Mature Height (feet): 1.0 • Maximum Height at 20 Years (feet): 1 • Vegetative Spread Rate: Moderate • Lifespan: Lifespan
Biology
Reproduction
Duration: Perennial • Coppice Potential: No • Progagated by Bulbs: No • Propagated by Bare Root: Yes • Propagated by Container: Yes • Propagated by Corms: No • Propagated by Cuttings: Yes • Propagated by Seed: No • Propagated by Sod: No • Propagated by Sprigs: No • Propagated by Tubers: No • Fruit/Seed Period Begin: Spring • Fruit/Seed Period End: Summer • Fruit/Seed Persistence: No
Growth
Soil: Adapted to Medium Textured: Adapted to Medium Textured Soils • Adapted to Coarse Textured Soils: Yes • Anaerobic Tolerance: None • Salinity Tolerance: None • CaCO3 Tolerance: None • Minimum pH: 5.9 • Maximum pH: 7.9 • Fertility Requirement: Low
Sunlight: Shade Tolerance: Tolerant
Moisture: Drought Tolerance: High • Minimum Precipitation: 24 • Maximum Precipitation: 35 • Moisture Use: Low
Temperature: Minimum Temperature (F): -23 • Minimum Frost Free Days: 130
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Rosidae
(
)
- Takhtajan, 1967
- Superorder:
Rhamnanae
(
)
- Takhtajan Ex Reveal, 1992
- Order:
Rosales
(
)
- Perleb, 1826
- Family:
Rhamnaceae
(
)
- A.L. de Jussieu, 1789, nom. cons.
- buckthorn, nerpruns
- Genus:
Ceanothus
(
)
- C. Linnaeus, 1753
- Specific epithet:
flexilis
- Greene ex McMinn
- Botanical name: - Ceanothus flexilis Greene ex McMinn
- Specific epithet:
flexilis
- Greene ex McMinn
- Genus:
Ceanothus
(
- Family:
Rhamnaceae
(
- Order:
Rosales
(
- Superorder:
Rhamnanae
(
- Subclass:
Rosidae
(
- Class:
Spermatopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Notes
Name
Status: Accepted Name
.
Last scrutiny: 15-Mar-2000
Similar Species
Members of the genus Ceanothus
ZipcodeZoo has pages for 161 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them:
C. americanus (Ceanothus) · C. arboreus (Catalina Ceanothus) · C. arboreus 'Trewithen Blue' (Feltleaf Ceanothus) · C. arcuatus (Arching Ceanothus) · C. bakeri (Baker's Ceanothus) · C. coeruleus (Azure Ceanothus) · C. confusus (Rincon Ridge Ceanothus) · C. connivens (Trailing Buckbrush) · C. cordulatus (Mountain Whitethorn) · C. crassifolius (Hoary-Leaf Whitethorn) · C. cuneatus (Buckbrush) · C. cuneatus var. cuneatus (Buckbrush) · C. cuneatus var. fascicularis (Buck Brush Ceanothus) · C. cuneatus var. ramulosus (Buckbrush Ceanothus) · C. cuneatus var. rigidus (Monterey Ceanothus) · C. cyaneus (San Diego Buckbrush) · C. dentatus (Cropleaf Ceanothus) · C. divergens (Calistoga Ceanothus) · C. divergens 'Burtonensis' (Crop-Leaf Ceanothus) · C. diversifolius (Pine Mat Ceanothus) · C. fendleri (Ceanothus) · C. ferrisae (Coyote Ceanothus) · C. ferrisiae (Coyote Ceanothus) · C. flexilis (Flexible Ceanothus) · C. foliosus (Wavy-Leaf Whitethorn) · C. foliosus var. foliosus (Wavyleaf Ceanothus) · C. foliosus var. medius (Wavyleaf Buckbrush) · C. foliosus var. vineatus (Vine Hill Ceanothus) · C. fresnensis (Fresno Ceanothus) · C. gloriosus (Holly Ceanothus) · C. gloriosus var. exaltatus (Glory Brush Ceanothus) · C. gloriosus var. exaltatus 'Emily Brown' (Emily Brown Ceanothus) · C. gloriosus var. gloriosus (Point Reyes Ceanothus) · C. gloriosus var. porrectus (Mount Vision Ceanothus) · C. gloriosus 'Anchor Bay' (Holly Ceanothus) · C. greggii (Ceanothus) · C. greggii greggii (Ceanothus) · C. greggii greggii var. greggii (Mohave Desert Ceanothus) · C. greggii var. greggii (Desert Ceanothus) · C. greggii var. lanuginosa (Mohave Desert Ceanothus) · C. greggii var. perplexans (Cupleaf Ceanothus) · C. greggii var. vestitus (Mohave Ceanothus) · C. griseus (California Lilac) · C. griseus var. horizontalis 'Carmel Creeper' (California Lilac) · C. griseus var. horizontalis 'Diamond Heights' (Ceanothus) · C. griseus var. horizontalis 'Santa Ana' (California Lilac) · C. griseus var. horizontalis 'Yankee Point' (Ceanothus) · C. hearstiorum (Hearst Ranch Buckbrush) · C. herbaceus (Jersey Tea) · C. impressus (Santa Barbara Ceanothus) · C. impressus var. impressus (Santa Barbara Ceanothus) · C. impressus var. nipomensis (Santa Barbara Ceanothus) · C. impressus 'Puget Blue' (Santa Barbara Ceanothus) · C. incanus (Coast Whitethorn) · C. infestus (Junco) · C. integerrimus (Deer Brush) · C. integerrimus var. parvifolius (Deerbrush Ceanothus) · C. jepsonii (Jepson Ceanothus) · C. jepsonii var. albiflorus (Ceanothus) · C. jepsonii var. jepsonii (Jepson Ceanothus) · C. lemmoni (Lemmon's Ceanothus) · C. lemmonii (Ceanothus) · C. leucodermis (Chaparral Whitethorn) · C. lobbianus (Lobbian Ceanothus) · C. lorenzenii (Lorenzen's Ceanothus) · C. maritimus (Maritime Ceanothus) · C. maritimus 'Point Sierra' (Maratime Ceanothus) · C. martini (Martin Ceanothus) · C. martinii (Ceanothus) · C. masonii (Bolinas Ceanothus) · C. megacarpus (Big-Pod Ceanothus) · C. megacarpus var. insularis (Island Ceanothus) · C. megacarpus var. megacarpus (Bigpod Ceanothus) · C. mendocinensis (Mendocino Ceanothus) · C. microphyllus (Little-Leaf Whitethorn) · C. oliganthus (Hairy Ceanothus) · C. ophiochilus (Vail Lake Ceanothus) · C. otayensis (Otay Ceanothus) · C. palmeri (Palmer Ceanothus) · C. papillosus (Ceanothus) · C. papillosus roweanus (Wartleaf Ceanothus) · C. papillosus var. papillosus (Wartleaf Ceanothus) · C. papillosus var. roweanus (California Lilac) · C. papillosus var. roweanus 'Julia Phelps' (Ceanothus) · C. parryi (Ceanothus) · C. parvifolius (Ceanothus) · C. pinetorum (Ceanothus) · C. prostratus (Mahala Mats Ceanothus) · C. pumilus (Dwarf Ceanothus) · C. purpureus (Hollyleaf Ceanothus) · C. rigidus 'Snowball' (Monterey Ceanothus) · C. rodericki (Buckthorn) · C. roderickii (Pine Hill Ceanothus) · C. roweanus (Wartleaf Ceanothus) · C. rugosus (Rugose Ceanothus) · C. sanguineus (Oregon-Tea) · C. serpyllifolius (Coastal Plain Buckbrush) · C. serrulatus (Cascade Lake Ceanothus) · C. sonomensis (Sonoma Ceanothus) · C. sorediatus (Jim-Brush)
More Info
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- Search for Scholarly Articles: Google Scholar
- Search using Scientific Name and Vernacular Names: All the Web | AltaVista Canada | AltaVista | Excite | Google | HotBot | Lycos
- Search using Specialized Databases: GenBank | Medline | Scirus | CISTI/CAL | Agricola Periodicals | Agricola Books
Further Reading
- Englert, J. M. et al. 1999–. USDA-NRCS Improved conservation plant materials released by NRCS and cooperators. (NRCS Cons Pl Mat)
- Kartesz, J. T. 1994. A synonymized checklist of the vascular flora of the United States, Canada, and Greenland. (L US Can ed2)
- Munz, P. A. & D. D. Keck. 1959. A California flora. (F CalifMunz)
- Chen Yi-ling and Chou Pan-kai. 1982. Rhamnaceae. In: Chen Yi-ling, ed., Fl. Reipubl. Popularis Sin. 48(1): i-vi, 1-169.
Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 10, 2012.
Identifiers
- Biodiversity Heritage Library NamebankID: 3874838
- Catalogue of Life Accepted Name Code: ITS-28469
- GRIN Nomen Number: 418971
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 28469
- U.S.D.A. Plant Symbol: CEFL4
- Zipcode Zoo Species Identifier: 29003
Footnotes
- Yilin Chen & Carsten Schirarend "Rhamnaceae". in Flora of China Vol. 12 Page 115,355. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
