Overview
Interesting Facts
Common Names
Click on the language to view common names.
Common Names in Afrikaans:
Kasuarisboom
Common Names in English:
Cunningham's Casuarina, Beefwood, Cunningham's Beefwood, River She-Oak, River Sheoak, River-Oak
Common Names in Portuguese:
Casuarina, Pinheiro-Australiano
Description
Family Casuarinaceae
Trees
or shrubs
evergreen
, monoecious or dioecious. Ultimate
branchlets
jointed
, with several short, basal articles and 1-numerous elongated articles, slender, ridged
, often pubescent
at least when young. Leaves small, toothlike, in whorls of 4 to ca.
20 (equal to number of ridges
on branchlets) ; stipules absent. Inflorescences spikelike or headlike, with alternating whorls of toothlike bracts; within each bract a single flower, with 2 lateral
scalelike bracteoles, persistent
(rarely deciduous in male Allocasuarina) ; parts sometimes pubescent when young, mostly glabrous
at maturity. Flowers unisexual
, without pedicels. Male inflorescences spikes, terminal
or lateral, terete
, slender. Female inflorescences headlike, globose
or ellipsoid
, usually terminal on short, lateral branches. Male flowers: tepals 1 or 2, scalelike, deciduous, hooded
; stamen 1; anthers
2-loculed, longitudinally dehiscent
, basifixed
. Female flowers: perianth none; carpels 2, fused; only anterior carpel fertile, posterior carpel usually ± reduced or obsolete
; ovules 2 (rarely 4), parietal
, paired
at base
of carpel, chalazogamous; style short; stigmas 2, red, linear
. Infructescences
± woody, conelike, with dense whorls of fruit, with 2 bracteoles of each flower enlarged as valves
, persistent. Samaras flat, (usually) winged
at apex, initially enclosed by 2 bracteoles, which separate at maturity to release
samara. Seed 1; cotyledons large; endosperm absent; embryo straight, often more than 1.
Four genera and 97 species: mainly in Australia, extending to SE Asia, Malesia, Pacific Islands; commonly introduced
and occasionally naturalized
elsewhere; one genus and three species (all introduced) in China.[1]
Genus Casuarina
Young persistent
branchlets
distinguished from deciduous branchlets by shorter segments and differences in shape
or size of leaves; furrows
deep and closed
, concealing stomates. Infructescences
pedunculate
, pubescent
at least when immature
; bracts thin in exposed portion, not vertically expanded; bracteoles ± protruding from surface of infructescence, never greatly thickened, always lacking dorsal protuberance
. Samaras pale
yellow-brown or grayish, dull
, glabrous
. x
= 9.
Species 17: almost throughout range
of family
.
Hybrids are frequent in cultivation; in the flora
, hybrids are known between all combinations
of the three species.[2]
Physical Description
Habit: Tree • Growth Form: Single Stem • Shape and Orientation: Erect
Flowers: Bloom Period: Mid Spring • Flower Color: Red • Flower Conspicuous: No
Seeds: Seed per Pound: 700000 • Seed Spread Rate: Rapid • Seedling Vigor: High • Fruit/Seed Abundance: High • Fruit/Seed Color: Brown • Fruit/Seed Conspicuous: No • Cold Stratification Required: No
Foliage: Foliage Color: Green • Foliage Porosity Summer: Dense • Foliage Porosity Winter: Dense • Foliage Texture: Fine • Fall Conspicuous: No • Leaf Retention: Yes
Size/Age/Growth
Active Growth Period: Spring , Summer, Fall • Growth Rate: Rapid • Mature Height (feet): 100.0 • Maximum Height at 20 Years (feet): 50 • Size: over 40' tall. • Vegetative Spread Rate: None • Lifespan: Lifespan
Habitat
Typically found at an altitude of 0 to 3,868 meters (0 to 12,690 feet).[3]
Biology
Reproduction
Duration: Perennial • Coppice Potential: Yes • Progagated by Bulbs: No • Propagated by Bare Root: Yes • Propagated by Container: Yes • Propagated by Corms: No • Propagated by Cuttings: Yes • Propagated by Seed: Yes • Propagated by Sod: No • Propagated by Sprigs: No • Propagated by Tubers: No • Fruit/Seed Period Begin: Summer • Fruit/Seed Period End: Fall • Fruit/Seed Persistence: No
Growth
Culture: Space 30-40' apart.
Soil: Adapted to Medium Textured: Adapted to Medium Textured Soils • Adapted to Coarse Textured Soils: Yes • Anaerobic Tolerance: Medium • Salinity Tolerance: Low • CaCO3 Tolerance: Medium • Minimum pH: 6.5 • Maximum pH: 7.5 • Fertility Requirement: Low
Sunlight: Sun Exposure: Full Sun . • Shade Tolerance: Intolerant
Moisture: Drought Tolerance: Low • Minimum Precipitation: 25 • Maximum Precipitation: 60 • Moisture Use: Medium
Temperature: Minimum Temperature (F): 22 • Minimum Frost Free Days: 300 • Cold Hardiness: 9a, 9b, 10a, 10b, 11. (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Rosidae
(
)
- Takhtajan, 1967
- Superorder:
Casuarinanae
(
)
- (Lindley, 1833) Takhtajan, 1997 Ex Reveal & Doweld, 1999
- Order:
Fagales
(
)
- Engler, 1892
- Family:
Casuarinaceae
(
)
- R. Brown, in Flinders, 1814
- She-Oak Family
- Genus:
Casuarina
(
)
- Rumphius ex Linnaeus, 1759
- She-oak [Neo-Latin casuarius, cassowary, from resemblance of drooping branchlets to feathers of the cassowary]
- Specific epithet:
cunninghamiana
- Miq.
- Botanical name: - Casuarina cunninghamiana Miq.
- Specific epithet:
cunninghamiana
- Miq.
- Genus:
Casuarina
(
- Family:
Casuarinaceae
(
- Order:
Fagales
(
- Superorder:
Casuarinanae
(
- Subclass:
Rosidae
(
- Class:
Spermatopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Notes
Name
Status: Accepted Name
.
Last scrutiny: 1996
Similar Species
Members of the genus Casuarina
ZipcodeZoo has pages for 6 species, subspecies, varieties, forms, and cultivars in this genus:
C. cristata (Belah) · C. cunninghamiana (Cunningham's Casuarina) · C. equisetifolia (Australian Pine) · C. glauca (Gray Sheoak) · C. lepidophloia (Belah) · C. obesa (Swamp She-Oak)
More Info
- Search for Pictures: images.google.com
- Search for Scholarly Articles: Google Scholar
- Search using Scientific Name and Vernacular Names: All the Web | AltaVista Canada | AltaVista | Excite | Google | HotBot | Lycos
- Search using Specialized Databases: GenBank | Medline | Scirus | CISTI/CAL | Agricola Periodicals | Agricola Books
Further Reading
- Bio-ecology, by Frederic E. Clements. .. and Victor E. Shelford. .. New York, J. Wiley & Sons, inc.;1939. url p. 377.
- Flora Costaricensis. William Burger, editor. Family #40, Casuarinaceae, Family #41, PIperaceae [by] William Burger. 35 1971 Chicago, Ill.: Field Museum of Natural History, 1971. url p. 4.
- Forestry handbook / ed. by R. Dalrymple Hay, director of forests. Sydney: W. A. Gullick, 1915-17. url .
- Forestry handbook. .. Sydney, W. A. Gullick, government printer, 1915-17. url .
- Inventory of seeds and plants imported / U.S. Department of Agriculture, Bureau of Plant Industry. Washington, D.C.: G.P.O., 1914-1924. url p. 20, p. 23, p. 46, p. 75, p. 82, p. 94.
- Irish gardening. Dublin: Pub. Office, 1906-1922 url p. 137.
- Journal of ethnobiology. 8-9 1988-1989 Flagstaff, Ariz.: Center for Western Studies, 1981- url p. 64.
- Plants for California landscapes: a catalog of drought tolerant plants. [Sacramento, Calif.]: State of California, Resources Agency, Dept. of Water Resources, [1979] url p. 114, p. 16.
- Proceedings of the Linnean Society of New South Wales. Sydney, Linnean Society of New South Wales. url p. 124, p. 247, p. 289, p. 380, p. 80, p. 89, p. 99.
- Report on the forests of British East Africa. LondonPrinted for His Majesty's Stationery Off. by Darling1909 url p. 112.
- Standardized plant names; a catalogue of approved scientific and common names of plants in American commerce. Salem, Mass., 1923. url p. 38.
- Chen Te-chao. 1982. Casuarinaceae. In: Tseng Yung-chien, ed., Fl. Reipubl. Popularis Sin. 20(1): 1-5.
Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 10, 2012.
Data Sources
Accessed through GBIF Data Portal November 19, 2007:
- Australian National Herbarium
- , Australian National Herbarium
- Bernice Pauahi Bishop Museum, Bishop Museum Natural History Specimen Data
- GBIF-Spain, Dirección General de Investigación, Desarrollo Tecnológico e Innovación de la Junta de Extremadura(DGIDTI): HSS
- GBIF-Spain, Institut Botanic de Barcelona, BC
- GBIF-Spain, Jardín Botánico de Córdoba: Herbarium COA
- Instituto Nacional de Biodiversidad
- , Biodiversidad de Costa Rica
- International Plant Genetic Resources Institute(IPGRI), The System-wide Information Network for Genetic Resources
- Missouri Botanical Garden, Missouri Botanical Garden
- National Herbarium of New South Wales, NSW herbarium collection
- National Herbarium of New South Wales, Plants of Papua New Guinea
- USDA PLANTS, USDA PLANTS Database
Identifiers
- Biodiversity Heritage Library NamebankID: 2666025
- Catalogue of Life Accepted Name Code: IOP-48625
- Global Biodiversity Information Facility Taxonkey: 13691820
- Globally Unique Identifier: urn:lsid:ipni.org:names:51808-3
- GRIN Nomen Number: 9548
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 501334
- International Plant Names Index (IPNI) ID:
- Natural Heritage Network Species Identifier: PDCAS01020
- U.S.D.A. Plant Symbol: CACU8
- Zipcode Zoo Species Identifier: 28968
Footnotes
- Nianhe Xia, Lawrence A. S. Johnson & Karen L. Wilson "Casuarinaceae". in Flora of China Vol. 4 Page 106. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- "Casuarina". in Flora of North America Vol. 3. Oxford University Press. Online at EFloras.org. [back]
- Mean = 412.700 meters (1,354.003 feet), Standard Deviation = 449.030 based on 284 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
