deciduous, monoecious. Stipules present, free
, often deciduous, rarely persistent
. Leaves alternate, simple
, usually doubly serrate, rarely simply serrate, lobulate
, or entire; veins pinnate. Flowers unisexual
. Male inflorescence precocious
, pendulous, with numerous
overlapping bracts; each bract usually subtending
a small dichasium with 1-3 male flowers; stamens as many as and opposite sepals or, if sepals obsolete
, then stamens of inflorescence to 20; filaments
very short, connate
or nearly so; anthers
2-loculed, thecae connate or separate, opening by longitudinal
slits. Female inflorescence pendulous or erect
, with numerous overlapping bracts; each bract subtending a small dichasium with 2 or 3 flowers; calyx with 1-6 scalelike lobes
, or obsolete; petals absent; ovary inferior, 2-loculed; styles 2, free; ovules 2, or 1 by abortion
, pendulous from near apex of each locule. Fruit a nut or nutlet
or not. Seed 1, with straight embryo and flat or thickened cotyledons, without endosperm.
Six genera and 150-200 species: mainly in Asia, Europe, and North and South America; six genera (one endemic) and 89 species (56 endemic) in China.
Because of evolutionary divergence within the Betulaceae, the family has often been divided into tribes (i.e. , Betuleae, Carpineae, and Coryleae) or more recently into subfamilies (J. J. Furlow, J. Arnold Arbor . 71: 1-67. 1990) .
, 8--25 m
usually 1, branching mostly deliquescent, trunk and branches irregularly longitudinally ridged
of trunk and branches bluish to brownish gray, thin, smooth
, close [thicker, broken
generally inconspicuous. Wood
nearly white to light brown, very hard and heavy, texture
fine. Branches, branchlets
, and twigs
conspicuously 2-ranked; young twigs differentiated into long and short shoots
. Winter buds
, 4-angled in cross
, apex acute; scales
many, imbricate, smooth. Leaves on long and short shoots, 2-ranked. Leaf blade
to ovate, elliptic
, or obovate
with 10 or more pairs of lateral
veins, 3--12 × 3--6 cm, thin, margins
doubly serrate to serrulate
; surfaces abaxially glabrous
, sometimes covered with small glands
. Inflorescences: staminate
catkins solitary or in small racemose clusters
, lateral, formed previous growing season
and enclosed [exposed] in buds during winter, expanding with leaves; pistillate
catkins distal to staminate on short, leafy new growth, solitary, ± erect
; bracts and flowers uncrowded. Staminate flowers
in catkins 3 per scale, crowded together on pilose
receptacle; stamens 3(--6), short; filaments
part way to base
into 2 parts, each 1-locular, apex pilose, Pistillate flowers 2 per bract. Infructescences
racemose clusters of paired
bracts, clusters pendulous, elongate; paired bracts deciduous with fruit, expanded, (1--) 3-lobed, variously toothed
, each bract subtending 1 fruit. Fruits small nutlets
, deltoid, longitudinally ribbed
, often crowned with persistent
sepals and styles. x
Species ca. 25: mostly north temperate zone; Europe, Asia (s to India, Iran).
In the flora , Carpinus consists of a single native species , C. caroliniana, which is composed of two fairly distinctive geographic races (J. J. Furlow 1987, 1987b), treated here as subspecies . Worldwide it includes about 25 species, some of which become large trees. The European C. betulus is frequently planted in North America and persists long after other signs of human development have vanished. It seldom escapes , however, and it has not become naturalized . In the mountains of Mexico and Central America, the larger C. tropicalis (Donnell Smith) Lundell is widespread in the temperate forest zone.
Closely related to Ostrya, Carpinus is easily recognized by its smooth, gray, often fluted stems and racemose infructescences consisting of pairs of uncrowded, foliaceous, 3-lobed bracts, each subtending a small triangular nutlet . The staminate (but not the pistillate) catkins develop in the autumn and are enclosed within buds throughout the winter prior to anthesis (in Ostrya, these are exposed during the winter). The pistillate catkins are produced on the first new growth in the spring .
Of relatively minor economic importance, Carpinus has limited use for its very hard wood, especially in Europe, where it is used for making mallet heads , tool handles, levers, and other small, hard, wooden objects.
Size: over 40' tall.
- Whittaker & Margulis,1978
- Haeckel, 1866
- Cavalier-Smith, 1981
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Kenrick & Crane, 1997
- Brongniart, 1843
- Takhtajan, 1967
- (Engler, 1892) Takhtajan, 1997
- Engler, 1892
- Gray, 1821
- Birch Family
- Subfamily: Coryloideae ()
- Family: Betulaceae () - Gray, 1821 - Birch Family
- Order: Fagales () - Engler, 1892
- Superorder: Faganae () - (Engler, 1892) Takhtajan, 1997
- Subclass: Rosidae () - Takhtajan, 1967
- Class: Spermatopsida () - Brongniart, 1843
- Infraphylum: Radiatopses () - Kenrick & Crane, 1997
- Subphylum: Euphyllophytina ()
- Phylum: Tracheophyta () - Sinnott, 1935 ex Cavalier-Smith, 1998 - Vascular Plants
- Subkingdom: Viridaeplantae () - Cavalier-Smith, 1981
- Kingdom: Plantae () - Haeckel, 1866 - Plants
Carpinus coreensis Koidz. • Carpinus fauriei Nakai • Carpinus yedoensis Maxim.
: Maxim. Publication
. Acad. Petersb. xxvii.
Name Status: Accepted Name .
Last scrutiny: 11-Nov-2003
Members of the genus Carpinus
ZipcodeZoo has pages for 21 species, subspecies, varieties, forms, and cultivars in this genus:
C. betulus (European Hornbeam) · C. betulus 'Asplenifolia' (European Hornbeam) · C. betulus 'Fastigiata' (Columnar European Hornbeam) · C. betulus 'Globosa' (European Hornbeam) · C. betulus 'Incisa' (European Hornbeam) · C. betulus 'Pendula' (Hornbeam) · C. betulus 'Quercifolia' (European Hornbeam) · C. betulus 'Streetwise' (Common Hornbeam 'streetwise') · C. betulus 'Vienna Weeping' (Hornbeam) · C. caroliniana (American Hornbeam) · C. caroliniana caroliniana (American Hornbeam) · C. caroliniana var. Crimson Cloud (Crimson Cloud English Hawthorn) · C. caroliniana virginiana (American Hornbeam) · C. cordata (Heartleaf Hornbeam) · C. fangiana (Fang Hornbeam) · C. japonica (Japanese Hornbeam) · C. laxiflora (Hornbeam) · C. orientalis (Eastern Hornbeam) · C. turczaninovii (Korean Hornbeam) · C. turczaninowii (Korean Hornbeam) · C. viminea (Hornbeam)
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- 1997 IUCN red list of threatened plants Cambridge: IUCN, World Conservation Union, 1998 url p. 207.
- Flora of Japan: in English: combined, much revised and extended translation / by the author of his Flora of Japan (1953) and Flora of Japan, Pteridophyta (1957); edited by Frederick G. Meyer and Egbert H. Walker. Washington: Smithsonian Institution, 1965. url p. 371.
- Great Basin naturalist memoirs. 1992 [Provo, Utah]Brigham Young University, 1976-1992. url p. 1259.
- Li Pei-chun & Cheng Sze-hsu. 1979. Betulaceae. In: Kuang Ko-zen & Li Pei-chun, eds., Fl. Reipubl. Popularis Sin. 21: 44-137.
- Furlow, J. J. 1987. The Carpinus caroliniana complex in North America. I. A multivariate analysis of geographical variation. Syst. Bot. 12: 21--40.
- Furlow, J. J. 1987b. The Carpinus caroliniana complex in North America. II. Systematics. Syst. Bot. 12: 416--434.
- Winkler, H. 1914. Neue Revision der Gattung Carpinus. Bot. Jahrb. Syst. 15(suppl.): 488--508.
- Winstead, J. E., B. J. Smith, and G. I. Wardell. 1977. Fruit weight clines in populations of ash, ironwood, cherry, dogwood and maple. Castanea 42: 56--60.
- Bisby, F.A., Y.R. Roskov, M.A. Ruggiero, T.M. Orrell, L.E. Paglinawan, P.W. Brewer, N. Bailly, J. van Hertum, eds (2007). Species 2000 & ITIS Catalogue of Life: 2007 Annual Checklist. Species 2000: Reading, U.K.
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 10, 2012.
- Global Biodiversity Information Facility. Accessed November 15, 2007. http://www.gbif.org Mediated distribution data from 6 providers.
- Ruggiero M., Gordon D., Bailly N., Kirk P., Nicolson D. (2011). The Catalogue of Life Taxonomic Classification, Edition 2, Part A. In: Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist (Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D., eds). DVD; Species 2000: Reading, UK.
- The International Plant Names Index. Accessed Dec 27, 2011.
- World Checklist of Selected Plant Families. Release date: November 27, 2009
Accessed through GBIF Data Portal November 15, 2007:
- Forest Research Institute, Department of Natural Forests, Herbarium
- Missouri Botanical Garden, Missouri Botanical Garden
- National Institute of Genetics, ROIS, Herbarium Specimens of Museum of Nature and Human Activities, Hyogo Pref., Japan
- National Science Museum of Korea, National Science Museum of Korea Plant
- Biodiversity Heritage Library NamebankID: 5828700
- Catalogue of Life Accepted Name Code: Kew-34455
- Global Biodiversity Information Facility Taxonkey: 14251049
- Globally Unique Identifier: urn:lsid:ipni.org:names:295285-1
- International Plant Names Index (IPNI) ID: 295285-1
- Zipcode Zoo Species Identifier: 724572