Interesting Facts
Description
Genus Brodiaea
Herbs, perennial
, scapose
, from fibrous-coated corms. Leaves 1-6, basal; blade
linear
, crescent-shaped in cross
section
. Scape solitary, cylindrical, usually slender, occasionally stout, rigid
. Inflorescences umbellate
, open, bracteate
; bracts scarious
, not enclosing flower buds. Flowers: perianth 6-tepaled, distinctly connate
proximally into tube
, shiny, abaxial
perianth usually bluish purple, tube narrowly campanulate
or funnelform
, outer 3 lobes
narrower than inner 3; stamens 3, epitepalous, opposite inner perianth lobes, alternating with 3 staminodia (staminodia absent in B
. orcuttii) opposite outer perianth lobes; filaments
adnate
to perianth tube, linear, base
sometimes dilated
to form triangular flap, or sometimes with abaxial wings or appendages
; anthers
basifixed
, appressed
to style; pistil 3-carpellate; ovary superior, green (purple in B. jolonensis), sessile, 3-locular, ovules several; style erect; stigma 3-lobed, lobes distinctly spreading
and recurved; pedicel erect
, articulate
at base. Fruits capsular
, ovoid
, dehiscence loculicidal. Seeds black, rounded
to flattened, coat
with crust with longitudinal
surface striations
. x
= 6, 8, 12, 16, 18, 20, or 24.
Species 14: w North America including Mexico (Baja California).
Two schools of thought have existed regarding generic
limits
within the complex
of species recognized under Brodiaea, Triteleia, and Dichelostemma: a single large genus (Brodiaea s.l.) with three subgenera
(S. Watson 1879; W. L. Jepson 1923-1925; P. A. Munz 1959), or three separate genera (E. L. Greene 1886; R. F. Hoover 1939; G. Keator 1967, 1989, 1993; T. F. Niehaus 1971, 1980). Recent molecular, anatomical, and developmental evidence supports
neither of these views
. Bloomeria is related to Triteleia, and Brodiaea is closely related to Dichelostemma, with the only hexandrous
species, D. capitatum, being sister to the rest of the three-staminate Brodiaea/Dichelostemma clade (R. Y. Berg
1978, 1996; J. C.
Pires 2000). This recent evidence also suggests that the sections presently established
within Brodiaea are in need of revision
; thus a sectional classification is not utilized in this treatment (R. F. Hoover 1939b; T. F. Niehaus 1971; J. C. Pires 2000).
Polyploidy and ecological specialization to serpentine and other unique substrates is common in Brodiaea, resulting in several rare and endangered species. Eleven of the fourteen species are restricted
to California, where the flowering date is highly dependent
on the amount of moisture in the early spring
. Several species are exceedingly variable. Corms of some species were eaten by native
Americans. Among the most important diagnostic characters within Brodiaea are features of the androecium, particularly the size and shape
of the staminodia and apical filament appendages. These characters are easily seen with a hand lens
in the field
. When collecting flowering specimens, one should make a point
of mounting a few dissected
flowers in a manner that displays these critical characters.[1]
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Liliidae
(
)
- Takhtajan, 1967
- Superorder:
Lilianae
(
)
- Takhtajan, 1967
- Order:
Asparagales
(
)
- Bromhead, 1838
- Suborder:
Asparagineae
(
)
-
- Family:
Asparagaceae
(
)
- A.L. de Jussieu, 1789, nom. cons.
- Family:
Asparagaceae
(
- Suborder:
Asparagineae
(
- Order:
Asparagales
(
- Superorder:
Lilianae
(
- Subclass:
Liliidae
(
- Class:
Spermatopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Similar Species
Members of the genus Brodiaea
ZipcodeZoo has pages for 24 species, subspecies, varieties, forms, and cultivars in this genus:
B. appendiculata (Appendage Brodiaea) · B. californica (California Brodiaea) · B. californica var. californica (California Brodiaea) · B. californica var. leptandra (California Brodiaea) · B. coronaria (Cascade Lily) · B. coronaria coronaria (Crown Brodiaea) · B. coronaria rosea (Indian Valley Brodiaea) · B. coronaria subsp. rosea (Indian Valley Brodiaea) · B. elegans (Elegant Brodiaea) · B. elegans hooveri (Harvest Brodiaea) · B. elegans subsp. hooveri (Hoover's Brodiaea) · B. filifolia (Thread-Leaved Brodiaea) · B. insignis (Kaweah Brodiaea) · B. jolonensis (Chaparral Brodiaea) · B. kinkiensis (San Clemente Island Brodiaea) · B. minor (Dwarf Brodiaea) · B. orcuttii (Orcutt's Brodiaea) · B. pallida (Chinese Camp Brodiaea) · B. purdyi (Sierra Brodiaea) · B. stellaris (Star-Flower Brodiaea) · B. terrestris (Crown Brodiaea) · B. terrestris kernensis (Dwarf Brodiaea) · B. terrestris terrestris (Dwarf Brodiaea) · B. terrestris subsp. kernensis (Kern Brodiaea)
More Info
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Further Reading
- A flora of California, by Willis Linn Jepson. San Francisco, Calif., Cunningham, Curtis & Welch, 1909- url , p. 282, p. 289, p. 289, p. 290, p. 291.
- An introduction to California plant life Berkeley: University of California Press, [c1974] url p. 77.
- California fish and game. [San Francisco, etc.]: State of California, Resources Agency, Dept. of Fish and Game. url p. 250.
- California native plantsman: UC Berkeley Botanical Garden, Tilden Botanic Garden: with an introduction by Walter Knight; interviews conducted by Suzanne B. Riess in 1990. url p. 145.
- Catalog of hymenoptera in America north of Mexico / prepared cooperatively by specialists on the various groups of Hymenoptera under the direction of Karl V. Krombein. .. [et al.]. Washington: Smithsonian Institution Press, 1979- url p. 2046.
- Great Basin naturalist memoirs. 1987 [Provo, Utah]Brigham Young University, 1976-1992. url p. 806.
- Hand-list of herbaceous plants cultivated in the Royal Botanic Gardens. London, Printed for H. M. Stationery Off. by Darling, 1902. url p. 201.
- Journal of ethnobiology. 17 1997 Flagstaff, Ariz.: Center for Western Studies, 1981- url p. 162.
- Leaflets of western botany. San Fransisco:[J. T. Howell], 1932-1966. url .
- The Great Basin naturalist. 41 1981 Provo, Utah: M.L. Bean Life Science Museum, Brigham Young University, 1939-1999. url p. 26, p. 404, p. 85.
- Niehaus, T. F. 1971. A biosystematic study of the genus Brodiaea (Amaryllidaceae). Univ. Calif. Publ. Bot. 60.
Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 10, 2012.
Identifiers
- Biodiversity Heritage Library NamebankID: 2777510
- Zipcode Zoo Species Identifier: 4024495
Footnotes
- J. Chris Pires "Brodiaea". in Flora of North America Vol. 26 Page 20, 53, 55, 321, 326, 328, 331, 332, 336, 3. Oxford University Press. Online at EFloras.org. [back]
