Common Names
Click on the language to view common names.
Common Names in English:
Side-Oats Grama, Sideoats Grama, Tall Grama Grass
Common Names in Portuguese:
Grama-Azul
Common Names in Spanish:
Navajita Banderilla
Description
Family Poaceae
Annual
or perennial
herbs, or tall woody bamboos
. Flowering stems (culms
) jointed
, internodes hollow or solid; branches arising singly from nodes and subtended by a leaf sheath
and 2-keeled prophyll, often fascicled in bamboos. Leaves arranged alternately in 2 ranks
, differentiated into sheath, blade
, and an adaxial
erect
appendage
at sheath/blade junction (ligule) ; leaf sheath surrounding and supporting culm-internode, split to base
or infrequently tubular
with partially or completely fused margins
, modified with reduced blade in bamboos (culm sheaths) ; leaf blades divergent, usually long, narrow and flat, but varying from inrolled
and filiform
to ovate
, veins parallel, sometimes with cross-connecting veinlets
(especially in bamboos) ; ligule membranous or a line
of hairs
. Inflorescence terminal
or axillary
, an open, contracted
, or spikelike panicle, or composed of lax
to spikelike racemes
arranged along an elongate
central axis, or digitate, paired
, or occasionally solitary; axillary inflorescences often many, subtended by spatheoles
(specialized bladeless leaf sheaths) and gathered into a leafy compound
panicle; spikelets
often aggregated into complex
clusters
in bamboos. Spikelets composed of distichous bracts arranged along a slender axis (rachilla) ; typically 2 lowest bracts (glumes
) empty, subtending
1 to many florets
; glumes often poorly differentiated from accompanying bracts in bamboos. Florets composed of 2 opposing bracts enclosing a single small flower, outer bract (lemma) clasping
the more delicate, usually 2-keeled inner bract (palea) ; base of floret often with thickened prolongation articulated with rachilla (callus) ; lemma often with apical or dorsal bristle
(awn
), glumes also sometimes awned
. Flowers bisexual
or unisexual
; lodicules (small scales
representing perianth) 2, rarely 3 or absent, 3 to many in bamboos, hyaline
or fleshy
; stamens 3 rarely 1, 2, 6, or more in some bamboos, hypogynous, filaments
capillary
, anthers
versatile; ovary 1-celled, styles (1 or) 2(rarely 3), free
or united
at base, topped by feathery stigmas, exserted from sides or apex of floret. Fruit normally a dry indehiscent caryopsis with thin pericarp firmly adherent
to seed, pericarp rarely free, fleshy in some bamboos; embryo small or large; hilum
punctate
to linear
.
About 700 genera and 11,000 species: widely distributed in all regions of the world.[1]
Genus Bouteloua
Annuals
or perennials
. Culms
mostly tufted
. Leaf blades
narrow; ligule a line
of hairs
. Inflorescence of racemes
inserted
singly along an axis; racemes unilateral
, 1-80, short, deciduous or persistent
; spikelets
sessile, few to numerous
, biseriate
, sometimes pectinate
; rachis narrow, flat, ending in a straight or forked
point
. Spikelets subterete or laterally compressed
, fertile
floret 1, usually 2nd sterile
floret present, rarely this reduced to a rachilla extension
; glumes
unequal, narrow, membranous, 1-veined, keeled
, acuminate to awn-pointed; lemma of fertile floret ± as long as upper glume, rounded
or keeled on back, thinly leathery, 3-veined, veins excurrent into 3 short awns
, central awn sometimes flanked by 2 teeth, less often apex simply acute; palea veins sometimes excurrent; sterile floret variable within a species or even an individual specimen, usually lemma body reduced and prominently awned
. Caryopsis ellipsoid
. x = 7, 10.
About 40 species: Canada to Argentina, centered on Mexico; two species (both introduced
) in China.[2]
Physical Description
Species Bouteloua curtipendula
Perennial
with short, slender, scaly
rhizomes. Culms
tufted
, erect
,
30-100 cm tall. Leaf sheaths
glabrous
or nearly so; leaf blades
flat
or slightly involute
, 20-30 cm, 1-5 mm wide, both surfaces and margins
scabrous
, base
pubescent
; ligule ca.
1 mm.
Inflorescence axis
15-25
cm; racemes
10-50, 1-2 cm, purplish, secund
along axis, usually nodding
,
with 3-6 (-10) appressed
or ascending
spikelets
, falling entire.
Spikelets 4.5-10 mm; lower glume
linear-lanceolate, 2.5-4 mm; upper
glume lanceolate, 4(-7) mm; lemma of fertile
floret usually somewhat
exceeding glumes, acuminate, lateral
veins extended into ca. 1 mm
mucros
; palea slightly longer
than lemma; 2nd floret rudimentary,
with long central awn
and 2 shorter laterals, or greatly reduced,
or lacking. Fl.
and fr. summer to autumn. 2n = 28, 35, 40,
42, 56, 70. [source]
This is an American pasture grass
(Side-oats Grama) reported to be
excellent in China for grazing and also for hay. [source]
Habit: Graminoid • Growth Form: Rhizomatous • Shape and Orientation: Erect
Flowers: Bloom Period: Mid Summer • Flower Color: Yellow • Flower Conspicuous: No
Seeds: Seed per Pound: 159200 • Seed Spread Rate: Slow • Seedling Vigor: High • Fruit/Seed Abundance: Medium • Fruit/Seed Color: Brown • Fruit/Seed Conspicuous: No • Cold Stratification Required: No
Foliage: Foliage Color: Green • Foliage Porosity Summer: Dense • Foliage Porosity Winter: Porous • Foliage Texture: Medium • Fall Conspicuous: No • Leaf Retention: No
Size/Age/Growth
Active Growth Period: Summer • Growth Rate: Moderate • After Harvest Regrowth Rate: Moderate • Mature Height (feet): 3.0 • Size: 18-24" tall. • Vegetative Spread Rate: Slow • Lifespan: Lifespan
Habitat
Typically found at an altitude of 0 to 4,848 meters (0 to 15,906 feet).[3]
Biology
Reproduction
Duration: Perennial • Coppice Potential: No • Progagated by Bulbs: No • Propagated by Bare Root: No • Propagated by Container: No • Propagated by Corms: No • Propagated by Cuttings: No • Propagated by Seed: Yes • Propagated by Sod: No • Propagated by Sprigs: No • Propagated by Tubers: No • Fruit/Seed Period Begin: Summer • Fruit/Seed Period End: Fall • Fruit/Seed Persistence: No
Growth
Culture: Space 12-15" apart.
Soil: Adapted to Medium Textured: Adapted to Medium Textured Soils • Adapted to Coarse Textured Soils: Yes • Anaerobic Tolerance: None • Salinity Tolerance: Medium • CaCO3 Tolerance: High • Minimum pH: 5.5 • Maximum pH: 7.8 • Fertility Requirement: Medium
Sunlight: Sun Exposure: Full Sun . • Shade Tolerance: Intolerant
Moisture: Drought Tolerance: Medium • Minimum Precipitation: 6 • Maximum Precipitation: 25 • Moisture Use: Medium
Temperature: Minimum Temperature (F): -33 • Minimum Frost Free Days: 120 • Cold Hardiness: 4a, 4b, 5a, 5b, 6a, 6b, 7a, 7b, 8a, 8b, 9a, 9b. (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 Ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Liliopsida
(
)
- Scopoli, 1760
- Subclass:
Commelinidae
(
)
- Takhtajan, 1967
- Superorder:
Poanae
(
)
- (Small, 1903) Takhtajan, 1997 Ex Reveal & Doweld, 1999
- Order:
Poales
(
)
- Small, 1903
- Family:
Poaceae
(
)
- (R. Brown) Barnhart, 1895
- Grass Family
- Subfamily:
Chloridoideae
(
)
- Tribe:
Cynodonteae
(
)
- Genus:
Bouteloua
(
)
- Lagasca, Varied. Ci. 2: 134. 1805 [Botelua].
- Grama
- Specific epithet:
curtipendula
- (Michaux) Torrey, Explor. Red River Louisiana. 300. 1853.
- Botanical name: - Bouteloua curtipendula (Michx.) Torr.
- Specific epithet:
curtipendula
- (Michaux) Torrey, Explor. Red River Louisiana. 300. 1853.
- Genus:
Bouteloua
(
- Tribe:
Cynodonteae
(
- Subfamily:
Chloridoideae
(
- Family:
Poaceae
(
- Order:
Poales
(
- Superorder:
Poanae
(
- Subclass:
Commelinidae
(
- Class:
Liliopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Unambiguous Synonyms
- Andropogon curtipendulum (Michx.) Spreng. Ex Steud.
- Atheropogon curtipendulus (Michaux) E. Fournier
- Atheropogon curtipendulus (Michx.) E. Fourn.
- Chloris curtipendula Michaux, Fl. Bor. -Amer. 1: 59. 1803
- Chloris curtipendula Michx.
- Cynodon curtipendulus (Michaux) Raspail
- Cynodon curtipendulus (Michx.) Raspail
- Dinebra curtipendula (Michaux) P. Beauvois
- Dinebra curtipendula (Michx.) P. Beauv.
- Eutriana curtipendula (Michaux) Trinius.
Notes
Name
Status: Accepted Name
. Latest taxonomic
scrutiny: 15-Mar-2000
Place of publication
: R. B
. Marcy, Explor. Red River
Louisiana 300. 1853 (W. H. Emory, Not. milit. reconn. 154. 1848, nom. provis.)
Name verified on 24-Feb-2004 by ARS Systematic Botanists. Last updated: 24-Feb-2004
Similar Species
Members of the genus Bouteloua
ZipcodeZoo has pages for 159 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them:
B. acuminata · B. alamosana · B. americana (American Grama) · B. annua · B. arenosa · B. aristidioides · B. aristidoides (Needle Gramma) · B. aristidoides var. aristidoides (Arizona Needle Grama) · B. aristidoides var. arizonica (Arizona Needle Grama) · B. aristoides · B. aschenborniana · B. barbata (Six Weeks Gramma) · B. barbata var. barbata · B. barbata var. major · B. barbata var. sonorae · B. bolanderi · B. brachyathera · B. bracteata · B. brasiliensis · B. brevifolia · B. breviseta (Gypsum Grama Grass) · B. bromoides · B. bromoides var. bromoides · B. burkei · B. burkii · B. butuostiga · B. chasei · B. chihuahuana · B. chondrioisoides · B. chondrosioides (Spruce-Top Gramma) · B. chondrosoides · B. ciliata · B. curtipendula (Tall Grama Grass) · B. curtipendula 'El Reno' · B. curtipendula 'Trailway' · B. curtipendula var. aristosa · B. curtipendula var. caespitosa (Sideoats Grama) · B. curtipendula var. curtipendula (Side-Oats Grama) · B. curtipendula var. tenuis · B. dactyloides (Legacy Buffalo Grass) · B. dimorpha · B. distans · B. disticha · B. diversispicula · B. elata · B. elatior · B. eludens (Santa Rita Mountain Grama) · B. erecta · B. eriopoda (Woolly Foot Grama) · B. eriopoda var. eriostachya · B. eriostachya · B. filiformis · B. foena · B. foenea · B. fournierana · B. glandulosa · B. gracilis (Grama-Needlegrass) · B. gracilis 'Hachita' (Blue Grama Grass) · B. gracilis var. gracilis · B. griffithsii · B. havardi · B. havardii · B. heterostega · B. hirsuta (Hairy Gramma) · B. hirsuta minor · B. hirsuta pectinata · B. hirsuta hirsuta · B. hirsuta var. hirsuta (Hairy Grama) · B. hirsuta var. minor · B. hirsuta var. pectinata (Tall Grama) · B. hirta · B. hirticulmis · B. humboldtiana · B. humilis · B. johnstoni · B. johnstonii · B. juncea (Lamilla) · B. juncifolia · B. karwinskii · B. kayi (Kay's Grama) · B. latifolia · B. litigiosa · B. litigosa · B. longiseta · B. lophostachya · B. major · B. media · B. megapotamica · B. melicaeformis · B. melicoides · B. mexicana · B. micrantha · B. micranthaa · B. microstachya · B. microstachys · B. multifida · B. multiseta · B. nervatus · B. oligostachya · B. oligostachya major
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Further Reading
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Notes
Contributors
- Bisby, F.A., Y.R. Roskov, M.A. Ruggiero, T.M. Orrell, L.E. Paglinawan, P.W. Brewer, N. Bailly, J. van Hertum, eds (2007). Species 2000 & ITIS Catalogue of Life: 2007 Annual Checklist. Species 2000: Reading, U.K.
- "Bouteloua curtipendula". in Flora of China Vol. 22 Page 494. Published by Missouri Botanical Garden Press. Online at EFloras.org.
- Brands, S.J. (comp.) 1989-2006. Systema Naturae 2000. The Taxonomicon. Universal Taxonomic Services, Amsterdam, The Netherlands. Accessed October 5, 2006.
- Global Biodiversity Information Facility. Accessed November 21, 2007. http://www.gbif.org Mediated distribution data from 62 providers.
- USDA, ARS, National Genetic Resources Program. Germplasm Resources Information Network - (GRIN) [Online Database]. National Germplasm Resources Laboratory, Beltsville, Maryland. URL (April 26, 2008)
Data Sources
Accessed through GBIF Data Portal November 21, 2007:
- Missouri Botanical Garden, Missouri Botanical Garden
- School of Life Sciences, Arizona State University, Arizona State University Vascular Plant Herbarium
- USDA PLANTS, USDA PLANTS Database
- University of Alabama Biodiversity and Systematics, Herbarium
- Utah Valley State College
- , Utah Valley State College Herbarium
Identifiers
- Biodiversity Heritage Library NamebankID: 2661509
- Catalogue of Life Accepted Name Code: ITS-41500
- Global Biodiversity Information Facility Taxonkey: 13753982
- Globally Unique Identifier: urn:lsid:ipni.org:names:392772-1
- GRIN Nomen Number: 7524
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 41500
- Natural Heritage Network Species Identifier: PMPOA10060
- U.S.D.A. Plant Symbol: BOCU
- Zipcode Zoo Species Identifier: 24792
Footnotes
- Shou-liang Chen, De-Zhu Li, Guanghua Zhu, Zhenlan Wu, Sheng-lian Lu, Liang Liu, Zheng-ping Wang, Bi-xing Sun, Zheng-de Zhu, Nianhe Xia, Liang-zhi Jia, Zhenhua Guo, Wenli Chen, Xiang Chen, Yang Guangyao, Sylvia M. Phillips, Chris Stapleton, Robert J. Soreng, Susan G. Aiken, Nikolai N. Tzvelev, Paul M. Peterson, Stephen A. Renvoize, Marina V. Olonova & Klaus Ammann "Poaceae". in Flora of China Vol. 22. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Bi-xing Sun & Sylvia M. Phillips "Bouteloua". in Flora of China Vol. 22 Page 488, 494. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Mean = 315.950 meters (1,036.581 feet), Standard Deviation = 440.360 based on 9,331 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
