Interesting Facts
Common Names
Common Names in English:
Beet
Description
Family Amaranthaceae
Herbs, clambering
subshrubs
, shrubs
, or lianas. Leaves alternate or opposite, entire, exstipulate
. Flowers small, bisexual
or unisexual
, or sterile
and reduced, subtended by 1 membranous bract and 2 bracteoles, solitary or aggregated in cymes. Inflorescences elongated or condensed spikes (heads
), racemes
, or thyrsoid
structures of varying complexity. Bracteoles membranous or scarious
. Tepals 3-5, membranous, scarious or subleathery, 1-, 3-, 5-, or 7(-23) -veined. Stamens as many as tepals and opposite these, rarely fewer than tepals; filaments
free
, united
into a cup
at base
or ± entirely into a tube
, filament lobes present or absent, pseudostaminodes present or absent; anthers
(1- or) 2-loculed, dorsifixed
, introrsely dehiscent
. Ovary superior, 1-loculed; ovules 1 to many; style persistent
, short and indistinct or long and slender; stigma capitate, penicillate
, 2-lobed or forming 2 filiform
branches. Fruit a dry utricle or a fleshy
capsule, indehiscent, irregularly bursting, or circumscissile. Seeds lenticular
, reniform
, subglobose, or shortly cylindric
, smooth
or verruculose
.
About 70 genera and 900 species: worldwide; 15 genera (one introduced
) and 44 species (three endemic, 14 introduced) in China.
Morphology of the androecium, perianth (tepals), and the inflorescence has traditionally been used to circumscribe genera and tribes
. Pseudostaminodia are interstaminal appendages
with variously shaped apices. Filament appendages are the lateral
appendages of filaments (one on each side) . The basic structure of the inflorescence is the cyme (branchlets
arising from the bracteole axils, the bracteoles serving as bracts for upper flowers), which can be reduced to one flower with two bracteoles and a bract. Units
of dispersal
vary considerably (capsules opening with lower part persistent, flower and bracteoles falling together, or cymose
partial inflorescences breaking off above bract) and can be characteristic for genera. Several genera possess long trichomes
serving dispersal at the base of the tepals.[1]
Genus Beta
Herbs, annual
, biennial, or perennial
, often with fleshy
, thickened roots
, glabrous
throughout. Stems erect
or procumbent
, not jointed
, not armed
, not fleshy. Leaves alternate, petiolate
or sessile; blade
ovate-cordate to rhombic-cuneate, margins
± entire, apex obtuse
. Inflorescences spikelike cymes or glomerules
, ebracteate
at least in distal 1/2. Flowers bisexual
, bracteate
; perianth segments 3-5, distinct
, sometimes petaloid
, rounded
or keeled
abaxially, wings
and spines absent; stamens 5; ovary semi-inferior; stigmas usually 2-3(-5), connate
basally. Fruiting structures achenes, connate with receptacle, often enclosed by swollen perianth. Seeds horizontal, orbicular
or reniform
; seed coat
dark brown, smooth
; embryo ± annular
, perisperm
copious
. x = 9.
Species ca.
6: introduced
; Eurasia
.
Beta is widely distributed and is known especially for the economically important Beta vulgaris subsp.
vulgaris, the commonly cultivated beet. The forms of the beets introduced in North America and established
in the wild occupy both inland and maritime habitats
.
The taxonomy of the genus is complicated by a long history of cultivation in which selective breeding
has caused a bewildering array of diverse
morphologies. In looking at the differences between the two forms of Beta that occasionally become established in waste
places in North America, it is tempting to segregate
the clearly distinct forms as different species. However, a number of researchers documenting the variation
within the complex
(B
. V. Ford-Lloyd and J. T. Williams 1975; H. Van Kijk and B. Desplanque 1999) agree that the range
of variation stretches along a continuum between the two extreme types defined by the maritima and vulgaris groups. The morphological lineage
assigned here to subsp. maritima is usually considered ancestral to the cultivated forms of beet included
within subsp. vulgaris.[2]
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Subclass:
Caryophyllidae
(
)
- Takhtajan, 1967
- Superorder:
Caryophyllanae
(
)
- Takhtajan, 1967
- Order:
Caryophyllales
(
)
- Perleb, 1826
- Suborder:
Chenopodiineae
(
)
- Family:
Amaranthaceae
(
)
- Adanson, 1763 ex A.L. de Jussieu, 1789, nom. cons.
- amaranthes, pigweed
- Subfamily:
Chenopodioideae
(
)
- Subfamily:
Chenopodioideae
(
- Family:
Amaranthaceae
(
- Suborder:
Chenopodiineae
(
- Order:
Caryophyllales
(
- Superorder:
Caryophyllanae
(
- Subclass:
Caryophyllidae
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Notes
Publishing author
: Fisch. & C.A.Mey. Publication
: Bull
. Soc.
Imp. Naturalistes Moscou (1838) 360.
Name
Status: Accepted Name
.
Last scrutiny: 15-Mar-2000
Similar Species
Members of the genus Beta
ZipcodeZoo has pages for 144 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them:
B. adanensis (Beet) · B. atriplicifolia (Beet) · B. juncea var. japonica 'Mustard Lettuce' (Mustard) · B. lomatogona (Beet) · B. nana (Beet) · B. oleracea var. acephala 'Blue Armor' (Kale) · B. patellaris (Beet) · B. patula (Beet) · B. procumbens (Cultivated Beet) · B. trigyna (Beet) · B. trojana (Trojan Beet) · B. vulgaris (Beet) · B. vulgaris cicla (Chard) · B. vulgaris macrocarpa (Common Beet) · B. vulgaris maritima (Spinach Beet (As B Bengalensis - India)) · B. vulgaris var. cicla 'Argentata' (Swiss Chard) · B. vulgaris var. cicla 'Bright Lights' (Seakale Beet) · B. vulgaris var. cicla 'Bright Yellow' (Swiss Chard) · B. vulgaris var. cicla 'Broadstem Green' (Swiss Chard) · B. vulgaris var. cicla 'Canary' (Swiss Chard) · B. vulgaris var. cicla 'Cardinal' (Swiss Chard) · B. vulgaris var. cicla 'Charlotte' (Ruby Chard) · B. vulgaris var. cicla 'Costa Bianca' (Swiss Chard) · B. vulgaris var. cicla 'Flamingo' (Swiss Chard) · B. vulgaris var. cicla 'Fordhook Giant' (Swiss Chard) · B. vulgaris var. cicla 'French' (Swiss Chard) · B. vulgaris var. cicla 'Golden Sunrise' (Swiss Chard) · B. vulgaris var. cicla 'Japanese' (Swiss Chard) · B. vulgaris var. cicla 'Large White Ribbed' (Seakale Beet) · B. vulgaris var. cicla 'Leaf Beet' (Swiss Chard) · B. vulgaris var. cicla 'Liscia Verde Da Taglio' (Swiss Chard) · B. vulgaris var. cicla 'Lucullus' (Swiss Chard) · B. vulgaris var. cicla 'Lyons' (Swiss Chard) · B. vulgaris var. cicla 'Magenta Sunset' (Swiss Chard) · B. vulgaris var. cicla 'Oriole' (Swiss Chard) · B. vulgaris var. cicla 'Parrot' (Swiss Chard) · B. vulgaris var. cicla 'Pink Passion' (Swiss Chard) · B. vulgaris var. cicla 'Rainbow' (Seakale Beet) · B. vulgaris var. cicla 'Rhubarb Chard' (Swiss Chard) · B. vulgaris var. cicla 'Ruby Red' (Swiss Chard) · B. vulgaris var. cicla 'Scarlet Charlotte' (Swiss Chard) · B. vulgaris var. cicla 'Sibella' (Swiss Chard) · B. vulgaris var. cicla 'Silverado' (Swiss Chard) · B. vulgaris var. cicla 'Virgo' (Swiss Chard) · B. vulgaris var. cicla 'Vulcan' (Swiss Chard) · B. vulgaris var. cicla 'White King' (Swiss Chard) · B. vulgaris var. cicla 'White Silver 2' (Swiss Chard) · B. vulgaris var. cicla 'Witerbi Mangold' (Swiss Chard) · B. vulgaris var. conditiva (Beetroot) · B. vulgaris vulgaris (Yellow-Fleshed Beetroot) · B. vulgaris vulgaris var. vulgaris (Sicilian Broad-Rib Beet) · B. vulgaris vulgaris 'Arcangeli' (Sea Beet) · B. vulgaris 'Action' (Beet) · B. vulgaris 'Albina Vereduna' (Beet) · B. vulgaris 'Albino' (Beet) · B. vulgaris 'Alvro Mono' (Beet) · B. vulgaris 'Always Tender' (Beet) · B. vulgaris 'Asgrow Wonder' (Beet) · B. vulgaris 'Baby Ball' (Beet) · B. vulgaris 'Baby Gladiator' (Beet) · B. vulgaris 'Big Top' (Beet) · B. vulgaris 'Bikores' (Beet) · B. vulgaris 'Blankoma White' (Beet) · B. vulgaris 'Bloody Mary' (Beet) · B. vulgaris 'Bolivar' (Beet) · B. vulgaris 'Boltardy' (Beet) · B. vulgaris 'Bonel' (Beet) · B. vulgaris 'Boro' (Beet) · B. vulgaris 'Bull's Blood' (Bull's Blood Giant Ornamental Beet) · B. vulgaris 'Burpee's Golden' (Burpee's Golden Beet) · B. vulgaris 'Burpee's White' (Beet) · B. vulgaris 'Centurion' (Beet) · B. vulgaris 'Chariot' (Beet) · B. vulgaris 'Cheltenham Green Top' (Beet) · B. vulgaris 'Chicago Red' (Beet) · B. vulgaris 'Chioggia' (Beet) · B. vulgaris 'Crosby's Extra Early Egyptian' (Beet) · B. vulgaris 'Cylindra' (Beet) · B. vulgaris 'Detroit' (Beet) · B. vulgaris 'Detroit 2 - Nero' (Beet) · B. vulgaris 'Detroit Dark Red' (Beet) · B. vulgaris 'Detroit Supreme' (Beet) · B. vulgaris 'Dewings Early Blood Turnip' (Beet) · B. vulgaris 'Early Egyptian' (Beet) · B. vulgaris 'Early Wonder' (Beet) · B. vulgaris 'Egyptian Flat Tf 68' (Beet) · B. vulgaris 'Excalibur' (Beet) · B. vulgaris 'First Crop' (Beet) · B. vulgaris 'Formanova' (Beet) · B. vulgaris 'Forono' (Beet) · B. vulgaris 'Globe 2' (Beet) · B. vulgaris 'Golden' (Beet) · B. vulgaris 'Golden Surprise' (Beet) · B. vulgaris 'Green Top Bunching' (Beet) · B. vulgaris 'Improved Blood Turnip' (Beet) · B. vulgaris 'Kestrel' (Beet) · B. vulgaris 'Lalraj' (Beet) · B. vulgaris 'Little Chicago' (Beet) · B. vulgaris 'Lutz Green Leaf' (Beet) · B. vulgaris 'Macgregor's Favourite' (Beet)
More Info
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Further Reading
- Buttler, K. P. 1977. Revision von Beta section Corallinae (Chenopodiaceae) I. Selbststerile Basisarten. Mitt. Bot. Staatssamml. München 13:283.
- Davis, P. H., ed. 1965–1988. Flora of Turkey and the east Aegean islands. (F Turk)
- Flora Rossica; sive, Enumeratio plantarum in totius Imperii Rossici provinciis Europaeis, Asiaticis et Americanis hucusque observatarum, auctore Carolo Friderico a Ledebour. Stuttgartiae, Sumptibus Librariae E. Schweizerbart, 1842-1853. LAT url p. 690.
- Ford-Lloyd, B. V. & J. T. Williams. 1975. A revision of Beta section Vulgares (Chenopodiaceae), with new light on the origin of cultivated beets. Bot. J. Linn. Soc. 71:92.
- Greuter, W. et al., eds. 1984–. Med-Checklist. (L Medit) 1:296.
- Hegi. 1960. Illustrierte Flora von Mittel-Europa. ed. 1:1906-1931; ed. 2:1936-68; ed. 3:1966- (IllF MEur) 3:568.
- Komarov, V. L. et al., eds. 1934–1964. Flora SSSR. (F USSR)
- Krassochkin, V. T. 1959. Obzor vidov roda Beta. Trudy Prikl. Bot. 32:9–10.
- Kuan Ke-chien. 1979. Amaranthaceae. In: Kung Hsien-wu & Tsien Cho-po, eds., Fl. Reipubl. Popularis Sin. 25(2): 194241.
- Ford-Lloyd, B. V. and J. T. Williams. 1975. A revision of Beta section Vulgares (Chenopodiaceae), with new light on the origin of cultivated herbs. Bot. J. Linn. Soc. 71: 89-102.
Notes
Contributors
- Bisby, F.A., Y.R. Roskov, M.A. Ruggiero, T.M. Orrell, L.E. Paglinawan, P.W. Brewer, N. Bailly, J. van Hertum, eds (2007). Species 2000 & ITIS Catalogue of Life: 2007 Annual Checklist. Species 2000: Reading, U.K.
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 10, 2012.
- Global Biodiversity Information Facility. Accessed February 28, 2008. http://www.gbif.org Mediated distribution data from 2 providers.
- Ruggiero M., Gordon D., Bailly N., Kirk P., Nicolson D. (2011). The Catalogue of Life Taxonomic Classification, Edition 2, Part A. In: Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist (Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D., eds). DVD; Species 2000: Reading, UK.
- The International Plant Names Index. Accessed Dec 27, 2011.
- USDA, ARS, National Genetic Resources Program. Germplasm Resources Information Network - (GRIN) [Online Database]. National Germplasm Resources Laboratory, Beltsville, Maryland. URL (April 26, 2008)
- USDA, NRCS. 2005. The PLANTS Database, Version 3.5 (http://plants.usda.gov). National Plant Data Center, Baton Rouge, LA 70874-4490 USA.
Data Sources
Accessed through GBIF Data Portal February 28, 2008:
- Bioversity International: EURISCO
- Bioversity International: EURISCO, The European Genetic Resources Search Catalogue
- Leibniz Institute of Plant Genetics and Crop Plant Research (IPK): IPK Genebank
Identifiers
- Biodiversity Heritage Library NamebankID: 1
- Catalogue of Life Accepted Name Code: ITS-506573
- Globally Unique Identifier: urn:lsid:ipni.org:names:164473-1
- GRIN Nomen Number: 7049
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 506573
- International Plant Names Index (IPNI) ID: 164473-1
- U.S.D.A. Plant Symbol: BELO
- Zipcode Zoo Species Identifier: 25171
Footnotes
- Bojian Bao, Thomas Borsch & Steven E. Clemants "Amaranthaceae". in Flora of China Vol. 5 Page 415. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Leila M. Shultz "Beta". in Flora of North America Vol. 4 Page 258, 261, 266. Oxford University Press. Online at EFloras.org. [back]
