Interesting Facts
Common Names
Common Names in English:
Mountain Sandwort
Description
Family Caryophyllaceae
Herbs annual
or perennial
, rarely subshrubs
or shrubs
. Stems and branches usually swollen at nodes. Leaves opposite, decussate, rarely alternate or verticillate
, simple
, entire, usually connate
at base
; stipules scarious
, bristly
, or often absent. Inflorescence of cymes or cymose
panicles, rarely flowers solitary or few in racemes
, capitula, pseudoverticillasters, or umbels. Flowers actinomorphic
, bisexual
, rarely unisexual
, occasionally cleistogamous
. Sepals (4 or) 5, free
, imbricate, or connate into a tube
, leaflike or scarious, persistent
, sometimes bracteate
below calyx. Petals (4 or) 5, rarely absent, free, often comprising claw
and limb; limb entire or split, usually with coronal scales
at juncture of claw and limb. Stamens (2--) 5--10, in 1 or 2 series. Pistil 1; carpels 2--5, united
into a compound
ovary. Ovary superior, 1-loculed or basally imperfectly 2--5-loculed. Gynophore
present or absent. Placentation free, central, rarely basal; ovules (1 or) few or numerous
, campylotropous. Styles (1 or) 2--5, sometimes united at base. Fruit usually a capsule, with pericarp crustaceous
, scarious, or papery
, dehiscing by teeth or valves
1 or 2 × as many as styles, rarely berrylike with irregular dehiscence or an achene. Seeds 1 to numerous, reniform
, ovoid
, or rarely dorsiventrally compressed
, abaxially grooved
, blunt
, or sharply pointed
, rarely fimbriate-pectinate; testa granular
, striate
or tuberculate
, rarely smooth
or spongy
; embryo strongly curved
and surrounding perisperm
or straight but eccentric
; perisperm mealy.
Between 75 and 80 genera and ca.
2000 species: widespread but mainly of temperate
or warm-temperate occurrence in the N hemisphere, with principal centers of distribution in the Mediterranean region and W Asia to W China and the Himalayas, fewer species in Africa S of the Sahara, America, and Oceania; 30 genera (two endemic) and 390 species (193 endemic) in China.
Arenaria, Silene, and Stellaria contain over half the species in the family
in China. They are mostly concentrated in the Qinghai-Xizang plateau
, and are especially rich from the Hengduan Mountains to the Himalayas. The main uses of this family are medicinal and ornamental
. Dianthus superbus, Pseudostellaria heterophylla, Stellaria dichotoma var. lanceolata, and Vaccaria hispanica are commonly used in traditional Chinese medicine
. Some species of Arenaria, Dianthus, Gypsophila, Psammosilene, and Silene are used as medicinal herbs among the people or are habitually used in local Chinese medicine. Many species of Dianthus, Gypsophila, Lychnis, Saponaria, and Silene are grown as ornamentals. Atocion armeria (Linnaeus) Rafinesque ( Silene armeria Linnaeus), native
to Russia and Europe, is also cultivated in China. It differs from Silene in having a corymbose
inflorescence and obscure
calyx veins. Wu Cheng-yih, Ke Ping, Zhou Li-hua, Tang
Chang-lin & Lu De-quan. 1996. Caryophyllaceae. In: Tang Chang-lin, ed., Fl.
Reipubl. Popularis Sin. 26: 47–449.[1]
Genus Arenaria
Herbs annual
or perennial
, rarely biennial. Stems erect
or rarely creeping
, often caespitose
or pulvinate
. Leaves opposite, rarely whorled
; leaf blade
linear
to elliptic
, ovate
, or orbicular
, usually flat, margin
entire. Flowers solitary or numerous
in cymes, actinomorphic
. Sepals 4 or 5, apex entire, rarely emarginate
. Petals 4 or 5, sometimes absent, apex entire to toothed
, 2-cleft, or fimbriate. Stamens (2--5 or 8 or) 10. Ovary 1-loculed; ovules numerous. Styles 2 or 3(--5) . Capsule ovoid
, obovoid
, or globose
, usually shorter than persistent
sepals, rarely equaling or longer
than them, 3- or 6-valved. Seeds reniform
or subovoid, flattened, smooth
, tuberculate
, or narrowly winged
.
More than 300 species: N temperate
to arctic
regions; 102 species (79 endemic) in China.
Arenaria has been subdivided into ten subgenera
, several of which were further divided
into sections
and series (McNeill, Notes
Roy. Bot. Gard. Edinburgh 24: 79 155. 1962) . Six subgenera are represented in China, one of which ( A. subgen.
Odontostemma ) contains more than half the species.
[2]
Physical Description
Habit: Mat
Flowers: Bloom Period: March, June. • Flower Color: White • Flower Conspicuous: Cup-shaped flowers
Foliage: Prostrate stems with 1/2 to 3/4-inch leaves • Foliage Shape: Linear • Normal foliage color: Green • Underside foliage: Green • Juvenile foliage: Green • Mature foliage: Green • New foliage: Green • Spring foliage: Green • Summer foliage: Green • Fall foliage: Green • Winter foliage: Green
Size/Age/Growth
Growth Rate: Fast Growing • Size: Growing to 2 to 4 in. high, spreads to 2 ft . Flower stalks to 12 inches.
Landscaping
Care: Follow a regular watering schedule during the first growing season to establish a deep, extensive root system . Control weeds until the plants have filled in. Feed with a general purpose fertilizer .
Habitat
Typically found at an altitude of 0 to 1,303 meters (0 to 4,275 feet).[3]
Biology
Growth
Culture: Space 3-6" apart.
Soil: Minimum pH: 5.6 • Maximum pH: 7.5
Sunlight: Sun Exposure: Full to partial sun
Moisture: Water Requirements: Water regularly, when top 3 in. of soil is dry.
Temperature: Heat Zones: High: 5 (>30 to 45 days) Low:1 (< 1 days) (map) • Cold Hardiness: High:5 (-20 to -10 F) Low:3 (-40 to -30 F) (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Caryophyllidae
(
)
- Takhtajan, 1967
- Superorder:
Caryophyllanae
(
)
- Takhtajan, 1967
- Order:
Caryophyllales
(
)
- Perleb, 1826
- Suborder:
Caryophyllineae
(
)
-
- Family:
Caryophyllaceae
(
)
- Durande, 1782 ex A.L. de Jussieu, 1789, nom. cons.
- cariophyllacées, pinks
- Subfamily:
Alsinoideae
(
)
- Subfamily:
Alsinoideae
(
- Family:
Caryophyllaceae
(
- Suborder:
Caryophyllineae
(
- Order:
Caryophyllales
(
- Superorder:
Caryophyllanae
(
- Subclass:
Caryophyllidae
(
- Class:
Spermatopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Notes
Publishing author
: Raf. Publication
: Atl. Journ. 151.
Publishing author: Malagarriga Publication: Las Subesp. y Variac.
Geogr.: 6 (1973)
Basionym
author: (Boiss.)
Publishing author: L. Publication: Cent. i. 12; Amoen. Acad. iv
.
272.
An accepted name
in the RHS
Horticultural Database.
Similar Species
Members of the genus Arenaria
ZipcodeZoo has pages for 89 species, subspecies, varieties, forms, and cultivars in this genus:
A. aberrans (Mt. Dellenbaugh Sandwort) · A. aculeata (Needleleaf Sandwort) · A. balearica (Corsican Sandwort) · A. benthamii (Hilly Sandwort) · A. capillaris (Slender Mountain Sandwort) · A. capillaris americana (Beautiful Sandwort) · A. capillaris americana var. americana (Fescue Sandwort) · A. capillaris capillaris (Slender Mountain Sandwort) · A. capillaris subsp. americana (Fescue Sandwort) · A. capillaris subsp. capillaris (Slender Mountain Sandwort) · A. ciliata (Fringed Sandwort) · A. congesta (Baldhead Sandwort) · A. congesta var. cephaloidea (Sharptip Sandwort) · A. congesta var. charlestonensis (Charleston Sandwort) · A. congesta var. congesta (Ballhead Sandwort) · A. congesta var. crassula (Ballhead Sandwort) · A. congesta var. glandulifera (Gland Sandwort) · A. congesta var. lithophila (Loosehead Sandwort) · A. congesta var. prolifera (Ballhead Sandwort) · A. congesta var. simulans (Ballhead Sandwort) · A. congesta var. subcongesta (Ballhead Sandwort) · A. congesta var. suffrutescens (Suffrutescent Sandwort) · A. congesta var. wheelerensis (Wheeler's Sandwort) · A. eastwoodiae (Eastwood Sandwort) · A. eastwoodiae var. adenophora (Eastwood's Sandwort) · A. eastwoodiae var. eastwoodiae (Eastwood's Sandwort) · A. fendleri (Fendler Sandwort) · A. fendleri brevifolia (Fendler's Sandwort) · A. fendleri var. brevifolia (Fendler Sandwort) · A. fendleri var. fendleri (Fendler Sandwort) · A. fendleri var. porteri (Porter Sandwort) · A. fendleri var. tweedyi (Tweedy Sandwort) · A. franklinii (Franklin Sandwort) · A. franklinii var. franklinii (Franklin's Sandwort) · A. franklinii var. thompsonii (Thompson's Sandwort) · A. holosteoides (False Jagged-Ckickweed) · A. hookeri (Hooker Sandwort) · A. hookeri desertorum (Hooker Sandwort) · A. hookeri pinetorum (Hooker Sandwort) · A. hookeri subsp. desertorum (Hooker's Desert Sandwort) · A. hookeri subsp. pinetorum (Hooker's Sandwort) · A. humifusa (Creeping Sandwort) · A. kingii (King Sandwort) · A. kingii compacta (King's Compact Sandwort) · A. kingii kingii (King's Sandwort) · A. kingii plateauensis (King's Plateau Sandwort) · A. kingii rosea (King's Rosy Sandwort) · A. kingii uintahensis (Uinta Sandwort) · A. kingii subsp. compacta (King's Compact Sandwort) · A. kingii subsp. kingii (King's Sandwort) · A. kingii subsp. plateauensis (King's Plateau Sandwort) · A. kingii subsp. rosea (King's Rosy Sandwort) · A. kingii subsp. uintahensis (Uinta Sandwort) · A. lanuginosa (Spreading Sandwort) · A. lanuginosa lanuginosa var. Lanuginosa (Spreading Sandwort) · A. lanuginosa saxosa (Spreading Sandwort) · A. lanuginosa var. lanuginosa (Spreading Sandwort) · A. lanuginosa var. longipedunculata (Spreading Sandwort) · A. lanuginosa subsp. saxosa (Spreading Sandwort) · A. lithophila (Loosehead Sandwort) · A. livermorensis (Livermore Sandwort) · A. longipedunculata (Longstem Sandwort) · A. ludens (Trans Pecos Sandwort) · A. macradenia (Mojave Sandwort) · A. macradenia ferrisiae (Mojave Sandwort) · A. macradenia macradenia (Forest Camp Sandwort) · A. macradenia macradenia var. Macradenia (Mojave Sandwort) · A. macradenia subsp. ferrisiae (Ferris' Sandwort) · A. macradenia subsp. macradenia (Mojave Sandwort) · A. montana (Mountain Sandwort) · A. montana 'Avalanche' (Mountain Sandwort) · A. montana 'Bloodstone' (Mountain Sandwort) · A. paludicola (Marsh Sandwort) · A. pinetorum (Hooker's Sandwort) · A. pseudoacantholimon (Sandwort) · A. pseudofrigida (Tundra Sandwort) · A. pumicola (Crater Lake Sandwort) · A. pumicola var. pumicola (Crater Lake Sandwort) · A. rosea (King's Rosy Sandwort) · A. serpyllifolia (Sandworts) · A. serpyllifolia leptoclados (Thymeleaf Sandwort) · A. serpyllifolia serpyllifolia (Thyme-Leaf Sandwort) · A. stenomeres (Meadow Valley Sandwort) · A. suffrutescens (Suffrutescent Sandwort) · A. tetraquetra (Spanish Sandwort) · A. tmolea (Sandwort) · A. tweedyi (Tweedy's Sandwort) · A. ursina (Bear Valley Sandwort) · A. verna 'Aurea' (Golden Moss)
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Further Reading
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- A garden diary: September 1899-September 1900 / London: Methuen & Co., 1901. url .
- A practical guide to garden plants, containing descriptions of the hardiest and most beautiful annuals and biennials, hardy herbaceous and bulbous perennials, hardy water and bog plants, flowering and ornamental trees and shrubs, conife London;Longmans, Green, 1901. url p. 90.
- A year's gardening London, T. Werner Laurie, [1912?] url .
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- Alien flora of Britain. London, West, Newman, 1905. url p. 33.
- Alpine flowers and rock gardens illustrated in colour / described by Walter P. Wright; with notes on "Alpine plants at home", by William Graveson. London: Headley Brothers, 1911. url p. 287.
- Alpine flowers for English gardens / by W. Robinson. London: Murray, 1879. url p. 185, p. 401, p. 434.
- Alpine flowers for gardens: rock, wall, marsh plants, and mountain shrubs / by W. Robinson. London: John Murray, 1903. url p. 172.
- Alpine flowers for gardens; rock, wall, marsh plants, and mountain shrubs, by W. Robinson. London, J. Murray, 1910. url p. 172.
- Alpine plants / by A.J. Macself; illustrated with colour photographs by R.A. Malby, water colour drawings by Winifred Walker, line drawings by G.E. Lee. New York: Scribner, 1923. url p. 64.
- Alpine plants, a practical manual for their culture, with a supplementary essay on The small rock-garden. London, C. Scribner's Sons, 1906, 1907. url .
- Alpines and bog-plants / by Reginald Farrer. London, Arnold, 1908. url p. 98.
- Alpines and bog-plants, London, E. Arnold, 1908. url .
- An encyclopaedia of gardening; comprising the theory and practice of horticulture, floriculture, arboriculture, and landscape-gardening, including all the latest improvements; a general history of gardening in all countries; and a By J.C. Loudon. London, Printed for Longman, Ross, Orme, Brown, and Green, 1828. url p. 882, p. 884.
- An encyclopædia of gardening; comprising the theory and practice of horticulture, floriculture, arboriculture, and landscape-gardening, including all the latest improvements; a general history of gardening in all countries; and a By J.C. Loudon. .. illustrated with many hundred engravings on wood by Branston. London: Printed for Longman, Rees, Orme, Brown, and Green, 1827. url p. 884.
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- Gardening for beginners; a handbook to the garden, by E.T. Cook. [London]"Country Life" [etc., 1914] url , .
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- Standardized plant names; a catalogue of approved scientific and common names of plants in American commerce. Salem, Mass., 1923. url p. 460.
- Studies in gardening, New York, C. Scribner's sons, 1916. url , , , , .
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- Sub-alpine plants: or, Flowers of the Swiss woods and meadows / by H. Stuart Thompson; with 33 coloured plates (168 figures) by George Flemwell. London: Dutton, 1912. url p. 108.
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- The Florists' exchange: a weekly medium of interchange for florists, nurserymen, seedsmen and the trade in general. New York, N.Y.: [A.T. De la Mare Ptg. and Pub. Co., url p. 1095, p. 1330, p. 306, p. 312.
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Notes
Contributors
- Brands, S.J. (comp.) 1989-2007. Systema Naturae 2000. The Taxonomicon. Universal Taxonomic Services, Amsterdam, The Netherlands. Accessed March 26, 2007.
- Global Biodiversity Information Facility. Accessed February 28, 2008. http://www.gbif.org Mediated distribution data from 3 providers.
- The International Plant Names Index. Accessed Jan 19, 2007.
Data Sources
Accessed through GBIF Data Portal February 28, 2008:
- Conservatoire botanique national du Bassin parisien: Observations du Conservatoire botanique national du Bassin parisien.
- European Environment Agency: EUNIS
- GBIF-Spain: Botánica, Universidad de León: LEB-Cormo
- GBIF-Spain: Departamento de Biolog. Veg. II, Facultad de Farmacia, Universidad Complutense, Madrid: MAF
- GBIF-Spain: Dirección General de Investigación, Desarrollo Tecnológico e Innovación de la Junta de Extremadura(DGIDTI): HSS
- GBIF-Spain: Herbario Universidad de Málaga: MGC-Cormófitos
- GBIF-Spain: Jardín Botánico de Córdoba: Herbarium COA
- GBIF-Spain: Real Jardin Botanico (Madrid), Vascular Plant Herbarium (MA)
- GBIF-Spain: Univ. Herbarium SALAMANCA: SALA
- GBIF-Spain: Universidad de Almería, HUAL
- GBIF-Spain: Universidad de Extremadura, UNEX
- GBIF-Spain: Universidad de Oviedo. Departamento de Biología de Organismos y Sistemas: FCO
- UK National Biodiversity Network: Botanical Society of the British Isles - Vascular Plants Database
Identifiers
- Biodiversity Heritage Library NamebankID: 5882029
- Global Biodiversity Information Facility Taxonkey: 13683851
- Globally Unique Identifier: urn:lsid:ipni.org:names:151459-1
- International Plant Names Index (IPNI) ID: 151458-1
- Zipcode Zoo Species Identifier: 645413
Footnotes
- Dequan Lu, Zhengyi Wu, Lihua Zhou, Shilong Chen, Michael G. Gilbert, Magnus Lidén, John McNeill, John K. Morton, Bengt Oxelman, Richard K. Rabeler, Mats Thulin, Nicholas J. Turland & Warren L. Wagner "Caryophyllaceae". in Flora of China Vol. 6 Page 1. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Dequan Lu, Zhengyi Wu, Lihua Zhou, Shilong Chen, Michael G. Gilbert, Magnus Lidén, John McNeill, John K. Morton, Bengt Oxelman, Richard K. Rabeler, Mats Thulin, Nicholas J. Turland & Warren L. Wagner "Arenaria". in Flora of China Vol. 6 Page 40. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Mean = 274.300 meters (899.934 feet), Standard Deviation = 313.060 based on 461 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
