Interesting Facts
Common Names
Common Names in English:
Undulate Alkanet
Description
Family Boraginaceae
Herbs perennial
, biennial, or annual
, less often lianas, shrubs
, or trees
, usually bristly
or scabrous-pubescent. Leaves simple
, exstipulate
, alternate, rarely opposite, entire or serrate at margin
. Inflorescences often double
scorpioid cymes, rarely solitary; bracts present or absent. Flowers bisexual
, actinomorphic
, rarely zygomorphic. Calyx usually 5-parted or lobed
, mostly persistent
. Corolla tubular
, campanulate
, rotate, funnelform
, or salverform
; tube
appendages
5, rarely more, mostly trapeziform, rarely absent, sometimes a ring
of hairs
present; limb usually 5-parted; lobes
overlapping, rarely twisted in bud. Stamens 5, inserted
on corolla tube or rarely at throat
, included
or rarely exserted; anthers
introrse
, 2-loculed, usually dorsifixed
at base
, less often medifixed
, dehiscence longitudinal
. Nectaries at base of corolla tube or on disc below ovary. Ovary superior, 2-carpellate; locules 2 and each with 2 ovules, or 4 and each with 1 ovule; ovules nearly atropous
, semianatropous, or anatropous
. Style terminal
or gynobasic
, branched or not. Gynobase
flat, fastigiate
, or subulate
. Fruit 1-4-seeded drupes or nutlets
(mericarps) ; nutlets mostly dry, often ornamented with wings
, prickles and/or glochids (stiff bristles
with barbed
or anchorlike tips
) . Seeds vertical
or oblique
, coat
membranous; embryo straight, less often curved
; cotyledons flat, fleshy
.
About 156 genera and 2500 species: temperate
and tropical regions
, centered in the Mediterranean region; 47 genera and 294 species in China, of which four genera and 156 species are endemic.[1]
Genus Anchusa
Herbs annual
or perennial
, sparsely strigose
or hispid
, rarely soft appressed
pubescent
. Stems erect
or spreading
. Leaves alternate. Cymes terminal
, widely spaced in fruit, scorpioid; bracts lanceolate. Calyx 5-parted nearly to base
or less; lobes
equal or unequal, linear
to triangular, often slightly enlarged in fruit. Corolla blue-purple or yellowish, regular or slightly zygomorphic; tube
usually longer
than calyx, straight or arcuate
or geniculate
curved
; throat
appendages
scaly
or tuberculate
and short pubescent; limb campanulate
; lobes 5, equal or unequal, apex obtuse
. Stamens inserted
at or below middle
of corolla tube, included
; filaments
short, filiform
; anthers
ovate-oblong, apex obtuse. Ovary 4-divided. Style included in corolla tube; stigma capitate, 2-cleft. Gynobase
flat. Nutlets
straight, reniform
, or oblique-ovoid, reticulate-wrinkled; attachment scar
at or near base, margin
ringlike, thickened, hardened.
About 50 species: N Africa, C and W Asia, Europe; one species in China.[2]
Physical Description
Flowers: Bloom Period: June, July, August, September. • Flower Color: medium blue
Size/Age/Growth
Size: 4-6' tall.
Habitat
Typically found at an altitude of 0 to 991 meters (0 to 3,251 feet).[3]
Biology
Growth
Culture: Space 18-24" apart.
Soil: Minimum pH: 5.6 • Maximum pH: 7.5
Sunlight: Sun Exposure: Full Sun .
Temperature: Cold Hardiness: 3a, 3b, 4a, 4b, 5a, 5b, 6a, 6b, 7a, 7b, 8a, 8b, 9a, 9b. (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Asteridae
(
)
- Takhtajan, 1967
- Superorder:
Solananae
(
)
- R. Dahlgren Ex Reveal, 1992
- Family:
Boraginaceae
(
)
- A.L. de Jussieu, 1789, nom. cons.
- borage, bourraches
- Subfamily:
Boraginoideae
(
)
- Tribe:
Boragineae
(
)
- Genus:
Anchusa
(
)
- C. Linnaeus, 1753
- Bugloss
- Specific epithet:
undulata
- Pall.
- Botanical name: - Anchusa undulata Pall.
- Specific epithet:
undulata
- Pall.
- Genus:
Anchusa
(
- Tribe:
Boragineae
(
- Subfamily:
Boraginoideae
(
- Family:
Boraginaceae
(
- Superorder:
Solananae
(
- Subclass:
Asteridae
(
- Class:
Spermatopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Notes
Publishing author
: B
.Valdés ex
M
.Laínz Publication
: in An. Jard.
Bot. Madrid, 38(2): 531 (1981 publ. 1982), contrary to Art.
45.l
& type collected after descr.
publ. 1982
Publishing author: Vahl ex DC. Publication: Prodr. (DC.) 10: 42 1846
[8 Apr 1846]
An accepted name
in the RHS
Horticultural Database.
Similar Species
Members of the genus Anchusa
ZipcodeZoo has pages for 11 species, subspecies, varieties, forms, and cultivars in this genus:
A. arvensis (Annual Bugloss) · A. azurea (Italian Alkanet) · A. azurea 'Dropmore' (Italian Alkanet) · A. azurea 'Feltham Pride' (Italian Alkanet) · A. azurea 'Loddon Royalist' (Italian Alkanet) · A. barrelieri (Barrelier's Bugloss) · A. capensis (Cape Bugloss) · A. capensis 'Blue Angel' (Summer Forget-Me-Not) · A. officinalis (Alkanet) · A. tinctoria (Alkanet) · A. undulata (Undulate Alkanet)
More Info
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Further Reading
- A complete dictionary of practical gardening: comprehending all the modern improvements in the art; whether in the raising of the various esculent vegetables, or in the forcing and managing of different sorts of fruits and plants, and that of lay By Alexander McDonald [pseud.] London, 1807. url .
- A manual flora of Egypt / by Reno Muschler. With a preface by Paul Ascherson and Georg Schweinfurth. Berlin, R. Friedlaender, 1912. url p. 794, p. 795.
- Alien flora of Britain. London, West, Newman, 1905. url p. 135.
- An encyclopaedia of gardening; comprising the theory and practice of horticulture, floriculture, arboriculture, and landscape-gardening, including all the latest improvements; a general history of gardening in all countries; and a By J.C. Loudon. London, Printed for Longman, Ross, Orme, Brown, and Green, 1828. url p. 872.
- An encyclopædia of gardening; comprising the theory and practice of horticulture, floriculture, arboriculture, and landscape-gardening, including all the latest improvements; a general history of gardening in all countries; and a By J.C. Loudon. .. illustrated with many hundred engravings on wood by Branston. London: Printed for Longman, Rees, Orme, Brown, and Green, 1827. url p. 872.
- Anales de la Sociedad Española de Historia Natural. Madrid: La Sociedad, url p. 100, p. 100, p. 200, p. 230, p. 246, p. 460, p. 85.
- Curtis's botanical magazine. 47 1820 London; New York [etc.]: Academic Press [etc.] url , .
- Flowering plants of the Riviera: a sescriptive account of 1800 of the more interesting species / by H. Stuart Thompson; with an introduction on Riviera vegetation by A. G. Tansley London; New York: Longmans, Green, 1914. url p. 161.
- Journal of botany, British and foreign. London: Robert Hardwicke, 1863-1942. url p. 14.
- Memoirs of the Literary and Philosophical Society of Manchester. Manchester: The Society, 1785-1879. url p. 153.
- Spicilegium florae Maroccanae / by John Ball. London: Printed by Taylor and Francis, 1878. url p. 571.
- The Annals of Scottish natural history. Edinburgh: David Douglas, 1892-1911. url p. 40.
- Wang Wen-tsai, Liu Yu-lan, Zhu Ge-ling, Lian Yong-shan, Wang Jing-quan & Wang Qing-rui in Kung Xian-wu & Wang Wen-tsai, eds. 1989. Boraginaceae. Fl. Reipubl. Popularis Sin. 64(2): 1-253.
Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 10, 2012.
- Global Biodiversity Information Facility. Accessed February 27, 2008. http://www.gbif.org Mediated distribution data from 4 providers.
- The International Plant Names Index. Accessed Jan 19, 2007.
Data Sources
Accessed through GBIF Data Portal February 27, 2008:
- GBIF-Spain: Botánica, Universidad de León: LEB-Cormo
- GBIF-Spain: Dirección General de Investigación, Desarrollo Tecnológico e Innovación de la Junta de Extremadura(DGIDTI): HSS
- GBIF-Spain: Dpto de Botánica, Ecología y Fisiología Vegetal (herbario_cofc).Facultad de Ciencias.Universidad de Córdoba
- GBIF-Spain: Herbario Universidad de Málaga: MGC-Cormófitos
- GBIF-Spain: Hortus Botanicus Sollerensis Herbarium (FBonafè)
- GBIF-Spain: Institut Botanic de Barcelona, BC
- GBIF-Spain: Jardín Botánico de Córdoba: Herbarium COA
- GBIF-Spain: Real Jardin Botanico (Madrid), Vascular Plant Herbarium (MA)
- GBIF-Spain: Univ. Herbarium SALAMANCA: SALA
- GBIF-Spain: Universidad de Almería, HUAL
- GBIF-Spain: Universidad de Extremadura, UNEX
- GBIF-Spain: Universidad de Oviedo. Departamento de Biología de Organismos y Sistemas: FCO
- GBIF-Sweden: Lund Botanical Museum (LD)
- Israel Nature and Parks Authority
- UK National Biodiversity Network: Botanical Society of the British Isles - Vascular Plants Database
Identifiers
- Biodiversity Heritage Library NamebankID: 5822556
- Global Biodiversity Information Facility Taxonkey: 13809541
- Globally Unique Identifier: urn:lsid:ipni.org:names:113388-1
- International Plant Names Index (IPNI) ID: 113386-1
- Zipcode Zoo Species Identifier: 717969
Footnotes
- Gelin Zhu, Harald Riedl & Rudolf V. Kamelin "Boraginaceae". in Flora of China Vol. 16 Page 329. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- "Anchusa". in Flora of China Vol. 16 Page 358. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Mean = 387.550 meters (1,271.490 feet), Standard Deviation = 294.120 based on 260 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
