- Amaranthus pumilus is a globally and federally threatened species (Global Heritage Status Rank G2; National Heritage Status Rank N2) originally restricted to dunes and barrier island beaches along the Atlantic Ocean from southern Massachusetts to South Carolina. It has been eliminated from two-thirds of its former range , being last reported from Massachusetts in 1849, from Rhode Island in 1856, from New Jersey in 1913, and from Virginia in 1972 (S. Ramsey et al. 2000). As of fall 2003, extant populations are known only from Long Island, New York, Delaware, Maryland, North Carolina, and South Carolia. Many threats exist, including construction of sea walls and dune fencing, development, heavy recreational use, and off-road vehicle traffic. It is difficult to afford protection because of the dynamic nature of the habitat and the fugitive nature of the biology of the species. "Fugitive" refers to the fact that the species does not necessarily occur throughout its potential range at any given time (S. E. Clemants 1992). [source]
- Amaranthus pumilus is in the Center for Plant Conservations National Collection of Endangered Plants . [source]
Common Names in English:
Coast Amaranth, Seabeach Amaranth, Seabeach Amaranthus, Seabeach Pigweed, Seaside Amaranth
, or lianas. Leaves alternate or opposite, entire, exstipulate
. Flowers small, bisexual
, or sterile
and reduced, subtended by 1 membranous bract and 2 bracteoles, solitary or aggregated in cymes. Inflorescences elongated or condensed spikes (heads
, or thyrsoid
structures of varying complexity. Bracteoles membranous or scarious
. Tepals 3-5, membranous, scarious or subleathery, 1-, 3-, 5-, or 7(-23) -veined. Stamens as many as tepals and opposite these, rarely fewer than tepals; filaments
into a cup
or ± entirely into a tube
, filament lobes present or absent, pseudostaminodes present or absent; anthers
(1- or) 2-loculed, dorsifixed
, introrsely dehiscent
. Ovary superior, 1-loculed; ovules 1 to many; style persistent
, short and indistinct or long and slender; stigma capitate, penicillate
, 2-lobed or forming 2 filiform
branches. Fruit a dry utricle or a fleshy
capsule, indehiscent, irregularly bursting, or circumscissile. Seeds lenticular
, subglobose, or shortly cylindric
About 70 genera and 900 species: worldwide; 15 genera (one introduced ) and 44 species (three endemic, 14 introduced) in China.
Morphology of the androecium, perianth (tepals), and the inflorescence has traditionally been used to circumscribe genera and tribes . Pseudostaminodia are interstaminal appendages with variously shaped apices. Filament appendages are the lateral appendages of filaments (one on each side) . The basic structure of the inflorescence is the cyme (branchlets arising from the bracteole axils, the bracteoles serving as bracts for upper flowers), which can be reduced to one flower with two bracteoles and a bract. Units of dispersal vary considerably (capsules opening with lower part persistent, flower and bracteoles falling together, or cymose partial inflorescences breaking off above bract) and can be characteristic for genera. Several genera possess long trichomes serving dispersal at the base of the tepals.
Herbs, usually annual
, rarely perennial
, monoecious (subg. Amaranthus and Albersia) or dioecious (subg. Acnida), glabrous
. Stems erect
, or prostrate
, usually branched, occasionally simple
or nearly so; without nodal
spines (except in A. spinosus ). Leaves alternate, petiolate
, lanceolate, oblanceolate
, or orbiculate to linear
to narrowly cuneate, margins
usually entire, usually plane
, slightly undulate
, or crispate
, rarely undulate-erose, apex acute, obtuse
, or emarginate
, usually mucronulate
. Inflorescences terminal
or exclusively terminal, compound
dichasia arranged in spikes, thyrses
, panicles, or glomerules
of terminal inflorescences often subtended by reduced leaves (pseudobracts), each dichasium unit
subtended by persistent
bracts. Bracts ovate, lanceolate, linear, subulate
, deltate, or broadly triangular (in A. acanthochiton), or proximal
bracts modified into spines (in A. spinosus) ; bracts of pistillate
flowers not keeled
(keeled in A. scleropoides and A. crassipes) ; bracteoles absent or 1-2. Flowers unisexual
. Pistillate flowers: tepals absent or (1-) 3-5, distinct
in proximal 1/3 in A. polygonoides, equal or outer tepals larger than inner ones, usually membranaceous
, sometimes scarious
at maturity; stamens absent [rudimentary
]; pistil 1; ovule 1; style 0.1-1 mm, or absent; stigmas 2-3(-5), slender. Staminate flowers
: tepals 3-5, equal or subequal
; stamens 3-5, filaments
4-locular, pseudostaminodes absent; pistils absent or rudimentary. Utricles loosely enclosed by inner tepals, occasionally conspicuously 3(-5) -veined, usually globose
, or elongate-ovoid, thin walled, membranaceous, rugose
, glabrous, dehiscence regularly circumscissile, irregularly dehiscent
, or indehiscent. Seeds 1, subglobose or lenticular
, usually smooth
, shiny, sometimes indistinctly puncticulate
; embryo annular
. x = 16, 17.
Species ca. 70 (38 in the flora , including cultivated species) : mostly tropical , subtropical , and warm-temperate zones, some species in temperate zones; some taxa are at present almost worldwide as introduced and naturalized weeds .
Some segregate genera of Amaranthus, in the broad sense, have been proposed and sometimes recognized (see synonymy ). In the present treatment, Amaranthus is accepted in its broad sense. Three subgenera are currently recognized (S. L. Mosyakin and K . R. Robertson 1996) : subg. Acnida, subg. Amaranthus, and subg. Albersia.
Morphologic terminology in Amaranthus, as used in different floristic and taxonomic treatments, is rather confusing, especially regarding the terms applied to inflorescences and flowers. In the present treatment, we follow the traditional inflorescence terminology only for brevity and convenience; see T. A. Fedorova (1997) for a more complex scheme. A flower is subtended by a bract, often termed a "bracteole," and 0-2 lateral bracts, the true bracteoles. Structures that are clearly reduced green leaves subtending portions of the inflorescence are sometimes incorrectly called bracts.
Specimens of Amaranthus are often difficult to identify by someone not familiar with the group. When using the key , look closely at the tips of pistillate inflorescence branches for staminate flowers to determine whether the plant is monoecious or dioecious; this is especially important for some monoecious species that produce few staminate flowers. Also, pistillate plants of dioecious species are usually required for positive identification. Descriptions and measurements of floral parts are given in more detail for pistillate flowers, unless noted otherwise.
Determining the exact distribution of some species of Amaranthus in North America requires additional floristic and taxonomic studies. Because of the weedy life strategies of some Amaranthus species, they may occasionally occur as naturalized weeds or waifs very far from their original areas of distribution. Some of such isolated populations exist only as long as conditions are favorable and may eventually disappear or, vice versa, become expansive and invasive. These factors , together with frequent misidentifications in herbaria and the literature, obscure the distribution patterns of some Amaranthus species in North America. Weedy and introduced species of Amaranthus are often neglected or misidentified by collectors . Consequently, some taxa are known only from scattered localities in various regions of the flora, and their actual distribution may be much wider than present data indicate. Some species have been reported for the flora only as rare, casual , non-naturalized aliens , e.g. , on ballast , or as grain immigrants or wool contaminants, and may not now be present in North America. Because of all these factors, the maps and distribution statements in the treatment show the generalized distribution and may not properly reflect the actual changing distribution patterns of some species, especially those that have expanded their ranges over the decades due to various anthropic factors. In addition to the taxa discussed below, some other South American or Old World species may be found in North America in the future as introduced weeds.
Species of Amaranthus occasionally form interspecific hybrids. Such hybridization seems to be especially important and widespread in cultivated grain-amaranths, in wild representatives of the A. hybridus aggregate, between species of sect. Amaranthus, and between A. tuberculatus and species of sect. Amaranthus. The degree and scope of hybridization in Amaranthus are often overestimated, especially by European authors , and some taxa described as putative hybrids are in fact nonhybrid infraspecific forms of morphologically variable species. Hybrids between more distantly related species, if they occur at all, are usually highly sterile , such as hybrids between taxa of the subgenera Amaranthus and Acnida, or at least show much decreased fertility . There are no verified records of hybrids between representatives of the subgenera Amaranthus and Albersia.
Some species of Amaranthus are cultivated as pseudocereal and leaf-vegetable crops , or as ornamental or fodder plants (J. D. Sauer 1967; D. M. Brenner 1990; J. T. Williams and D. M. Brenner 1995; S. Cheatham et al. 1995). The most commonly cultivated taxa are A. caudatus Linnaeus, A. hypochondriacus Linnaeus, and A. cruentus Linnaeus of American origin , and south Asian A. tricolor Linnaeus. The cultivated species may occur occasionally as escapes near places of cultivation; they cannot be regarded as truly naturalized.
Species of Amaranthus were widely used by prehistoric and modern Native Americans as food, forage for livestock, medicinal plants, and, occasionally, for some other uses, such as face and body paint, ceremonial items, and fuel (S. Cheatham et al. 1995; D. E. Moerman 1998).
Species Amaranthus pumilus
Plants annual , glabrous . Stems prostrate to ascending (often forming mats), red, much-branched, 0.1-0.4(-0.5) m , fleshy . Leaves clustered near tips of branches; petiole 5-10 mm; blade orbiculate, broadly ovate or obovate , 1-1.5 cm × 1-1.5 cm, fleshy, base broadly cuneate to tapering, margin entire, plane or slightly undulate , apex broadly rounded to obtuse , mucronate . Inflorescences dense axillary glomerules , green. Bracts of pistillate flowers ovate or elliptic , 1.2-2 mm, 1/2 as long as tepals. Pistillate flowers: tepals 5, linear to narrowly oblanceolate , slightly unequal, 2.5-4 mm, margins entire, apex obtuse; style branches erect ; stigmas 3. Staminate flowers intermixed with pistillate; tepals 5; stamens 5. Utricles ovoid , 4-6 mm, exceeding tepals, fleshy, smooth or slightly rugose , longitudinally wrinkled on drying, indehiscent. Seeds dark reddish brown, lenticular , 2.5 mm diam., glossy. Flowering summer-fall. [source]
Maritime sand dunes, beaches, mostly on foredunes and at high tide level; of conservation concern; 0-10 m .
Typically found in the intertidal zone at the water's edge at a mean distance from sea level of 2.50 meters (8.20 feet).
- Chatton, 1925
- Haeckel, 1866
- Cavalier-Smith, 1981
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Kenrick & Crane, 1997, nom. inval.
- Kenrick & Crane, 1997
- Novák ex Takht. (1967)
- Novák ex Takhtajan, 1967
- Takhtajan, 1967
- Juss. ex Bercht. & J. Presl, 1820
- Juss. ex Bercht. & J. Presl (1820)
- Family: Amaranthaceae () - A.L. de Jussieu, 1789, nom. cons. - Amaranth Family
- Suborder: Chenopodiineae () - Juss. ex Bercht. & J. Presl (1820)
- Order: Caryophyllales () - Juss. ex Bercht. & J. Presl, 1820
- Superorder: Caryophyllanae () - Takhtajan, 1967
- Subclass: Magnoliidae () - Novák ex Takhtajan, 1967 - Angiosperms
- Class: Spermatopsida () - Novák ex Takht. (1967)
- Infraphylum: Radiatopses () - Kenrick & Crane, 1997
- Subphylum: Euphyllophytina () - Kenrick & Crane, 1997, nom. inval.
- Phylum: Tracheophyta () - Sinnott, 1935 ex Cavalier-Smith, 1998 - Vascular Plants
- Subkingdom: Viridaeplantae () - Cavalier-Smith, 1981
- Kingdom: Plantae () - Haeckel, 1866 - Plants
Publishing author : Raf. Publication : Med. Repos. ser. 2, 5: 360 1808
Members of the genus Amaranthus
ZipcodeZoo has pages for 94 species, subspecies, varieties, forms, and cultivars in this genus:
A. acanthochiton (Green-Stripe Amaranth) · A. acutilobus (Sharplobe Amaranth) · A. albus (Pigweed) · A. A. caudatus (Love-Lies-Bleeding) · A. arenicola (Sand Amaranth) · A. australis (Giant Amaranth) · A. bigelovii (Bigelow's Amaranth) · A. blitoides (Mat Amaranth) · A. blitum (Bride Malabar) · A. blitum blitum (Purple Amaranth) · A. blitum var. pseudogracilis (Amaranthus Blitum) · A. brownii (Brown's Amaranth) · A. californicus (California Amaranth) · A. cannabinus (Salt-Marsh Water-Hemp) · A. caudatus (Foxtail Amaranth) · A. caudatus caudatus (Love-Lies-Bleeding) · A. caudatus 'Fat Spike' (Love-Lies-Bleeding) · A. caudatus 'Pony Tails' (Love-Lies-Bleeding) · A. caudatus 'Viridis' (Green Tassel Flower) · A. chihuahuensis (Chihuahuan Amaranth) · A. crassipes (Clubfoot Amaranth) · A. crispus (Crisp-Leave Amaranth) · A. cruentus (Blood Amaranth) · A. cruentus 'Giant Copperhead' (Mexican Grain Amaranth) · A. cruentus 'Golden Giant' (Mexican Grain Amaranth) · A. cruentus 'Hot Biscuits' (Mexican Grain Amaranth) · A. cruentus 'Marvel Bronze' (Mexican Grain Amaranth) · A. cruentus 'Oeschberg' (Mexican Grain Amaranth) · A. cruentus 'Orange Giant' (Mexican Grain Amaranth) · A. cruentus 'Split Personality' (Mexican Grain Amaranth) · A. cruentus 'Towers Green' (Mexican Grain Amaranth) · A. cruentus 'Towers Red' (Mexican Grain Amaranth) · A. cruentus 'Velvet Curtain' (Mexican Grain Amaranth) · A. deflexus (Argentina Amaranth) · A. dubius (Spleen Amaranth) · A. fimbriatus (Fringed Amaranth) · A. fimbriatus var. denticulatus (Fringed Amaranth) · A. fimbriatus var. fimbriatus (Fringed Amaranth) · A. floridanus (Florida Amaranth) · A. gangeticus (Elephant Head Amaranth) · A. gangeticus 'Greek' (Amaranth) · A. graecizans (Mat Amaranth) · A. greggii (Gregg's Amaranth) · A. hybridus (Green Amaranth) · A. hybridus cruentus (Green Pigweed) · A. hypochondriacus (Prince's Feather Amaranth) · A. hypochondriacus 'Burgundy' (Amaranth) · A. hypochondriacus 'Green Thumb' (Prince-Of-Wales Feather) · A. hypochondriacus 'Manna De Montana' (Amaranth) · A. hypochondriacus 'Mercado' (Amaranth) · A. hypochondriacus 'Oscar Blanco' (Amaranth) · A. hypochondriacus 'Pygmy Torch' (Prince-Of-Wales Feather) · A. lineatus (Australian Amaranth) · A. lividus (Purple Amaranth) · A. mangostanus (Chinese Spinach) · A. muricatus (African Amaranth) · A. obcordatus (Trans Pecos Amaranth) · A. palmeri (Careless Weed) · A. polygonoides (Smartweed Amaranth) · A. powellii (Green Amaranth) · A. pringlei (Pringle's Amaranth) · A. pumilus (Coast Amaranth) · A. retroflexus (Careless Weed) · A. retroflexus 'Fotete' (Green Amaranth) · A. rudis (Common Waterhemp) · A. scleropoides (Bone-Bract Amaranth) · A. spinosus (Calaloo) · A. texensis (Amaranthus) · A. thunbergii (Thunberg's Amaranth) · A. torreyi (Torrey's Amaranthus) · A. tricolor (Chinese Spinach) · A. tricolor var. salicifolius (Flaming Fountain) · A. tricolor 'Aurora' (Chinese Spinach) · A. tricolor 'Cinco De Mayo' (Amaranth) · A. tricolor 'Green Leaf' (Chinese Spinach) · A. tricolor 'Illumination' (Chinese Spinach) · A. tricolor 'New Garnet Red' (Chinese Spinach) · A. tricolor 'Perfecta' (Chinese Spinach) · A. tricolor 'Red Leaf' (Chinese Spinach) · A. tricolor 'Splendens' (Chinese Spinach) · A. tuberculatus (Rough-Fruit Amaranth) · A. viridis (Chinese Spinach) · A. viridis 'All Red' (Amaranth) · A. viridis 'Asia Red' (Amaranth) · A. viridis 'Bayam' (Amaranth) · A. viridis 'Green Pointed Leaf' (Amaranth) · A. viridis 'Green Round Leaf' (Amaranth) · A. viridis 'Red Stripe Leaf' (Amaranth) · A. viridis 'Tender Leaf' (Amaranth) · A. viridis 'White Leaf' (Amaranth) · A. watsonii (Torrey's Amaranthus) · A. wrightii (Wright's Amaranth) · A. 'Magic Fountains Mix' (Amaranthus) · A. 'Opopeo' (Love-Lies-Bleeding)
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- 1997 IUCN red list of threatened plants Cambridge: IUCN, World Conservation Union, 1998 url p. 47.
- An illustrated flora of the northern United States, Canada and the British possessions: from Newfoundland to the parallel of the southern boundary of Virginia and from the Atlantic Ocean westward to the 102nd meridian / by Nathaniel Lord Britton and Hon. Addison Brown. New York: Scribner, 1913. url .
- Annotated list of the ferns and flowering plants of New York state, by Homer D. House. Albany, The University of the state of New York, 1924. url p. 302.
- Annual report of the New Jersey State Museum. Trenton, N.J.: MacCrellish & Quigley, url p. 433, p. 54, p. 96.
- Annual report of the State Botanist of the State of New York. Albany: Troy Press, 1889-1897. url .
- Bartonia;proceedings of the Philadelphia botanical club. .. 57 Supplement 1993-1996 Philadelphia, Philadelphia Botanical Club, Academy of Natural Sciences. url p. 12, p. 2, p. 30, p. 35, p. 92.
- Botany of the northern and middle states, or, A description of the plants found in the United States, north of Virginia: arranged according to the natural system: with a synopsis of the genera according to the Linnaean system. .. / Albany: Printed by Webster and Skinners. .., 1833. url p. 295.
- Britton, N. L. (ed.). North American flora. 21 1917 [New York]New York Botanical Garden. url p. 119.
- Bulletin of the Torrey Botanical Club. 51 1924 New York: Torrey Botanical Club, 1870-1996 url p. 191, p. 431, p. 503, p. 55, p. 560.
- Catalogue of the cabinet of natural history of the state of New York, and of the historical and antiquarian collection annexed thereto. Albany, C. Van Benthuysen, printer, 1853. url .
- Contributions from the Botanical Laboratory. Philadelphia: University of Pennsylvania Press, 1892- url .
- Flora of Delaware and the Eastern Shore: an annotated list of the ferns and flowering plants of the peninsula of Delaware, Maryland and Virginia. [Wilmington]: Society of Natural History of Delaware, 1946. url p. 112.
- Flora of Pennsylvania, Boston, Ginn & Company, 1903. url , .
- Flora of Pennsylvania. Edited with the addition of analytical keys by John Kunkel Small. Boston, Ginn & Co., 1903. url p. 122, p. 123.
- Flora of the southeastern United States; being descriptions of the seed-plants, ferns and fern-allies growing naturally in North Carolina, South Carolina, Georgia, Florida, Tennessee, Alabama, Mississippi, Arkansas, Louisiana, and in Oklahom by John Kunkel Small. New York, The author, 1913. url p. 393, p. 393.
- Flora of the vicinity of New York; a contribution to plant geography. [New York]1915 url p. 29.
- Gray's lessons in botany and vegetable physiology, illus. by over 360 wood engravings, from original drawings, by Isaac Sprague. To which is added a copious glossary, or dictionary of botanical terms. By Asa Gray. Chicago, Ivison, Blakeman, Taylor & co. url p. 369.
- Handbook of the flora of Philadelphia and vicinity, containing data relating to the plants within the following radius: eastern Pennsylvania, north to the Blue mountains, and west to the Susquehanna; all of New Jersey except the northern counties; and Philadelphia, Philadelphia Botanical Club, 1905. url .
- List of Pteridophyta and Spermatophyta growing without cultivation in northeastern North America. Prepared by a Committee of the Botanical Club, American Association for the Advancement of Science. New York, 1894. url p. 145.
- Manual of the botany of the northern United States: including Virginia, Kentucky, and all east of the Mississippi / by Asa Gray; (The mosses and liverworts by Wm. S. Sullivant.) New York: Ivison, Phinney, 1863, c1862. url p. 369.
- Manual of the flora of the northern states and Canada / by Nathaniel Lord Britton. New York: Holt, 1905. url p. 379.
- Memoirs of the Torrey Botanical Club. 5 1893 - 18 Durham, N.C.: Published for the Club by the Seeman Printery, 1889- url p. 145.
- Natural area inventory of Pender County, North Carolina / Raleigh: North Carolina Coastal Energy Impact Program, Office of Coastal Management, North Carolina Dept. of Natural Resources and Community Development, [1982?] url p. 139, p. 142.
- Natural history of New York. Albany: D. Appleton: 1842-1894. url p. 144, p. 144, p. 532, p. 532.
- New York State Museum bulletin. Albany, N.Y.: University of the State of New York, 1916-1944. url p. 56, p. 97.
- Phytologia memoirs. Plainfield, N.J.: H.N. Moldenke and A.L. Moldenke, 1980- url p. 251.
- Proceedings of the Boston Society of Natural History. Boston [etc.]Boston Society of Natural History. url , p. 93.
- Report of the Botanist. Albany, N.Y.: James B. Lyon, State Printer, 1867-1924. url p. 11.
- Report of the State Botanist. Albany, N.Y.: University of the State of New York, 1898-1925. url .
- Small, J. K. Flora of the southeastern United States;being descriptions of the seed-plants, ferns and fern-allies growing naturally in North Carolina, South Carolin, Georgia, Florida, Tennessee, Alabama, Mississippi, Arkansas, Louisiana and the Indian territory and in Oklahoma and Texas east of the one-hundredth meridian /by John Kunkel Small. .. 1903 New York: The author, 1903. url p. 391.
- The American journal of science and arts. New-Haven: S. Converse, url p. 303.
- The American journal of science. 5 1822 New Haven, Kline Geology Laboratory, Yale University. url p. 303.
- The flora of Long Island. Lancaster, Pa.New Era Print., 1899. url .
- The flora of North Carolina from Ranunculaceae to Salviniaceae / C.W. Hyams. Raleigh, N.C.: The North Carolina College of Agriculture and Mechanic Arts, Agricultural Experiment Station, 1899. url p. 335.
- The plants of Southern New Jersey with especial reference to the flora of the pine barrens and the geographic distribution of the species. Trenton, N. J., 1911. url , , , .
- The plants of Southern New Jersey; with especial reference to the flora of the Pine Barrens and the geographic distribution of the species. Trenton, 1911. url p. 386.
- The plants of southern New Jersey; with especial reference to the flora of the pine barrens and the geographic distribution of the species. Trenton, 1911. url p. 433.
- The vegetation of Shackleford Bank / by I.F. Lewis. Raleigh: Edwards & Broughton, 1917. url p. 25, p. 29.
- Kuan Ke-chien. 1979. Amaranthaceae. In: Kung Hsien-wu & Tsien Cho-po, eds., Fl. Reipubl. Popularis Sin. 25(2): 194241.
- Brenner, D. M. et al. 2000. Genetic resources and breeding of Amaranthus. Pl. Breed. Rev. 19: 227-285.
- Costea, M. and D. A. DeMason. 2001. Stem morphology and anatomy in Amaranthus L. (Amaranthaceae): Taxonomic significance. J. Torrey Bot. Soc. 128: 254-281.
- Costea, M., A. Sanders, and G. Waines. 2001. Preliminary results toward a revision of the Amaranthus hybridus species complex (Amaranthaceae). Sida 19: 931-974.
- Mosyakin, S. L. and K. R. Robertson. 1996. New infrageneric taxa and combinations in Amaranthus L. (Amaranthaceae). Ann. Bot. Fenn. 33: 275-281.
- Sauer, J. D. 1955. Revision of the dioecious amaranths. Madroño 13: 5-46.
- Sauer, J. D. 1967b. The grain amaranths and their relatives: A revised taxonomic and geographic survey. Ann. Missouri Bot. Gard. 54: 103-137.
- Sauer, J. D. 1972b. The dioecious amaranths: A new species name and major range extensions. Madroño 21: 426-434.
- Uline, E. B. and W. L. Bray. 1894. A preliminary synopsis of the North American species of Amaranthus. Bot. Gaz. 19: 267-273, 313-320.
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 10, 2012.
Accessed through GBIF Data Portal December 11, 2007:
- USDA PLANTS, USDA PLANTS Database
- Biodiversity Heritage Library NamebankID: 2646911
- Catalogue of Life Accepted Name Code: ITS-20744
- Global Biodiversity Information Facility Taxonkey: 13740642
- Globally Unique Identifier: urn:lsid:ipni.org:names:59642-1
- GRIN Nomen Number: 315346
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 20744
- International Plant Names Index (IPNI) ID: 327381-2
- Natural Heritage Network Species Identifier: PDAMA040Z0
- U.S. Fish and Wildlife Service Species Identifier: Q2MZ
- U.S.D.A. Plant Symbol: AMPU2
- Zipcode Zoo Species Identifier: 19692
- Bojian Bao, Thomas Borsch & Steven E. Clemants "Amaranthaceae". in Flora of China Vol. 5 Page 415. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Sergei L. Mosyakin & Kenneth R. Robertson "Amaranthus". in Flora of North America Vol. 4 Page 405, 406, 410. Oxford University Press. Online at EFloras.org. [back]
- "Amaranthus pumilus". in Flora of North America Vol. 4 Page 414, 428, 430. Oxford University Press. Online at EFloras.org. [back]
- Standard Deviation = 15.360 based on 32 observations. Terrestrial altitude and ocean depth information for each observation from British Oceanographic Data Centre. [back]