Interesting Facts
Common Names
Common Names in English:
Smooth Joyweed
Description
Family Amaranthaceae
Herbs, clambering
subshrubs
, shrubs
, or lianas. Leaves alternate or opposite, entire, exstipulate
. Flowers small, bisexual
or unisexual
, or sterile
and reduced, subtended by 1 membranous bract and 2 bracteoles, solitary or aggregated in cymes. Inflorescences elongated or condensed spikes (heads
), racemes
, or thyrsoid
structures of varying complexity. Bracteoles membranous or scarious
. Tepals 3-5, membranous, scarious or subleathery, 1-, 3-, 5-, or 7(-23) -veined. Stamens as many as tepals and opposite these, rarely fewer than tepals; filaments
free
, united
into a cup
at base
or ± entirely into a tube
, filament lobes present or absent, pseudostaminodes present or absent; anthers
(1- or) 2-loculed, dorsifixed
, introrsely dehiscent
. Ovary superior, 1-loculed; ovules 1 to many; style persistent
, short and indistinct or long and slender; stigma capitate, penicillate
, 2-lobed or forming 2 filiform
branches. Fruit a dry utricle or a fleshy
capsule, indehiscent, irregularly bursting, or circumscissile. Seeds lenticular
, reniform
, subglobose, or shortly cylindric
, smooth
or verruculose
.
About 70 genera and 900 species: worldwide; 15 genera (one introduced
) and 44 species (three endemic, 14 introduced) in China.
Morphology of the androecium, perianth (tepals), and the inflorescence has traditionally been used to circumscribe genera and tribes
. Pseudostaminodia are interstaminal appendages
with variously shaped apices. Filament appendages are the lateral
appendages of filaments (one on each side) . The basic structure of the inflorescence is the cyme (branchlets
arising from the bracteole axils, the bracteoles serving as bracts for upper flowers), which can be reduced to one flower with two bracteoles and a bract. Units
of dispersal
vary considerably (capsules opening with lower part persistent, flower and bracteoles falling together, or cymose
partial inflorescences breaking off above bract) and can be characteristic for genera. Several genera possess long trichomes
serving dispersal at the base of the tepals.[1]
Genus Alternanthera
Herbs or subshrubs
, annual
or perennial
. Stems prostrate
, decumbent
, ascending
, erect
, or floating, indumentum of simple
trichomes
. Leaves opposite, sessile or petiolate
; blade
lanceolate to ovate
, ovate-rhombic, or obovate-rhombic, margins
entire. Inflorescences axillary
or terminal
, sessile or pedunculate
, several-flowered cylindric
spikes or globose
heads
, without immediately subtending
leaves; bracts and bracteoles scarious
. Flowers bisexual
; tepals 5, distinct
; stamens 3-5; filaments
connate
basally into tube
or short cup
; pseudostaminodes 5, alternating with stamens; ovule 1; style 1, ca.
0.2 mm; stigma capitate or rarely 2-lobed. Utricles compressed
, ovoid
or obovoid
, indehiscence. Seeds 1, reddish brown, lenticular
or ovoid-oblong.
Species ca. 80: primarily American tropics and subtropics, some in Asia, Africa, and Australia.[2]
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Caryophyllidae
(
)
- Takhtajan, 1967
- Superorder:
Caryophyllanae
(
)
- Takhtajan, 1967
- Order:
Caryophyllales
(
)
- Perleb, 1826
- Suborder:
Chenopodiineae
(
)
- Family:
Amaranthaceae
(
)
- Adanson, 1763 ex A.L. de Jussieu, 1789, nom. cons.
- amaranthes, pigweed
- Subfamily:
Gomphrenoideae
(
)
- Tribe:
Gomphreneae
(
)
- Genus:
Alternanthera
(
)
- Forsskål, 1775
- [Latin alternans, alternating, and anthera, anther, referring to the alternation of pseudostaminodes and stamens]
- Specific epithet:
paronichyoides
- St.-Hil.
- Variety:
paronichyoides
- Botanical name: - Alternanthera paronichyoides var. paronichyoides St.-Hil.
- Variety:
paronichyoides
- Specific epithet:
paronichyoides
- St.-Hil.
- Genus:
Alternanthera
(
- Tribe:
Gomphreneae
(
- Subfamily:
Gomphrenoideae
(
- Family:
Amaranthaceae
(
- Suborder:
Chenopodiineae
(
- Order:
Caryophyllales
(
- Superorder:
Caryophyllanae
(
- Subclass:
Caryophyllidae
(
- Class:
Spermatopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Synonyms
Achyranthes paronichyoides auct. non (L.) Lam. • Alternanthera polygonoides auct. non (L.) R. Br. Ex Sweet
Notes
Name Status: Accepted Name . Latest taxonomic scrutiny: 15-Mar-2000
Similar Species
Members of the genus Alternanthera
ZipcodeZoo has pages for 33 species, subspecies, varieties, forms, and cultivars in this genus:
A. bettzichiana (Calicoplant) · A. bettzickiana (Baptist Plant) · A. bettzickiana 'Brilliantissima' (Joyweed) · A. brasiliana (Brazil Chaff-Flower) · A. caracasana (Mat Chaff-Flower) · A. crucis (West Indian Joyweed) · A. dentata 'Purple Knight' (Calico Plant) · A. dentata 'Red Marble' (Calico Plant) · A. dentata 'Rubiginosa' (Calico Plant) · A. echinocephala (Seaurchin Joyweed) · A. ficoidea 'Aurea' (Chartreuse Josephs Coat) · A. ficoidea 'Chartreuse' (Chartreuse Josephs Coat) · A. ficoidea 'Grenadine' (Josephs Coat) · A. ficoidea 'New Burgundy' (Baptist Plant) · A. ficoidea 'Purple Threadleaf' (Baptist Plant) · A. ficoidea 'Rosea Nana' (Dwarf Rose Josephs Coat) · A. ficoidea 'Snow Carpet' (Brazilian Snow Flower) · A. flavescens (Yellow Chaff-Flower) · A. flavogrisea (Alternanthera) · A. hassleriana (Hassler's Alternanthera) · A. lanuginosa (Woolly Tidestromia) · A. maritima (Beach Alternanthera) · A. palmeri (Palmer's Joyweed) · A. paronichyoides (Smooth Chaff-Flower) · A. paronichyoides var. amazonica (Smooth Joyweed) · A. paronichyoides var. paronichyoides (Smooth Joyweed) · A. philoxeroides (Alligator Weed) · A. pungens (Creeping Chaffweed) · A. reineckii (Red Hygrophila) · A. sessilis (Sessile Chaff-Flower) · A. tenella (Alternanthera Snow Ball) · A. tenella tenella (Sanguinaria) · A. trigyna (Island Joyweed)
More Info
- Search for Pictures: images.google.com
- Search for Scholarly Articles: Google Scholar
- Search using Scientific Name and Vernacular Names: All the Web | AltaVista Canada | AltaVista | Excite | Google | HotBot | Lycos
- Search using Specialized Databases: GenBank | Medline | Scirus | CISTI/CAL | Agricola Periodicals | Agricola Books
Further Reading
- Kuan Ke-chien. 1979. Amaranthaceae. In: Kung Hsien-wu & Tsien Cho-po, eds., Fl. Reipubl. Popularis Sin. 25(2): 194241.
- Mears, J. A. 1978. The nomenclature and type collections of the widespread taxa of Alternanthera (Amaranthaceae). Proc. Acad. Nat. Sci. Philadelphia 129: 1-21.
Notes
Contributors
- Bisby, F.A., Y.R. Roskov, M.A. Ruggiero, T.M. Orrell, L.E. Paglinawan, P.W. Brewer, N. Bailly, J. van Hertum, eds (2007). Species 2000 & ITIS Catalogue of Life: 2007 Annual Checklist. Species 2000: Reading, U.K.
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 10, 2012.
Identifiers
- Catalogue of Life Accepted Name Code: ITS-181940
- Global Biodiversity Information Facility Taxonkey: 13890303
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 181940
- Natural Heritage Network Species Identifier: PDAMA03092
- Zipcode Zoo Species Identifier: 1012260
Footnotes
- Bojian Bao, Thomas Borsch & Steven E. Clemants "Amaranthaceae". in Flora of China Vol. 5 Page 415. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Steven E. Clemants "Alternanthera". in Flora of North America Vol. 4 Page 406, 435, 447. Oxford University Press. Online at EFloras.org. [back]
