Overview
Tree from India to Indonesia and Australia with leathery, whorled leaves and small, greenish-white flowers. The bark yields an antimalarial drug.
Common Names
Click on the language to view common names.
Common Names in English:
Blackboard Tree, Blackboardtree, Devil Tree, Deviltree, Dita Bark, Ditabark, Milkwood-Pine, White Cheesewood
Common Names in Hindi:
चािटयान, Chatian
Common Names in Indonesia:
Pulai
Common Names in Sanskrit:
Saptaparna
Description
Family Apocynaceae
Trees
, shrubs
, or vines
, rarely subshrubs
or herbs, with latex or rarely watery juice. Leaves simple
, opposite, rarely whorled
or alternate, pinnately veined; stipules absent or rarely present. Inflorescences cymose
, terminal
or axillary
, with bracteoles. Flowers bisexual
, 5- [or 4]-merous, actinomorphic
. Calyx 5- or rarely 4-partite, quincuncial, basal glands
usually present. Corolla 5- or rarely 4-lobed, salverform
, funnelform
, urceolate
, or rarely rotate, lobes
overlapping to right
or left, rarely valvate
. Stamens 5 or rarely 4; filaments
short; anthers
mostly sagittate
, free
or connivent into a cone adherent
to pistil head
, dehiscing longitudinally, base
rounded
, cordate, sagittate, or prolonged into an empty spur; pollen granular
; disc ringlike or cup-shaped, 2-5-lobed, or absent. Ovaries superior, rarely half-inferior, connate
or distinct
, 1- or 2-locular; ovules (1 or) 2-numerous per locule. Style
1; pistil head capitate, conical
, or lampshade-shaped, base stigmatic, apex 2-cleft and not stigmatic
. Fruit a berry, drupe, capsule, or follicle. Seeds with or without coma; endosperm thick and often horny
, scanty, sometimes absent; embryo straight or nearly so, cotyledons often large, radicle terete
.
About 155 genera and 2000 species distributed primarily in the tropics and subtropics, poorly represented in the temperate regions
. Of the 44 genera and 145 species present in China, one genus and 38 species are endemic, and nearly 95% of the taxa grow in the southern and southwestern portions of the country.
Fruit type is highly diversified in the family
, and it is diagnostic of many genera. Genera 1-4 produce
1, 2-celled berries
from a flower; genus 5 produces 2, 1-celled berries from a flower; 6 and 7 produce mostly fleshy
follicles containing deeply indented
seeds with ruminate
endosperm; 8 has follicles and winged
seeds; 9 produces follicles and seeds with 2 comas; 10-12 have follicles with globose
seeds; 13-18 have drupes mostly with fleshy mesocarp; 19 has samaroid
fruit; 20 has spiny
capsules with seeds winged all around; and 21-44 have free or fused follicles and comose
seeds. Double
flowers are known only from cultivated forms of Nerium oleander, Tabernaemontana divaricata, and Wrightia religiosa.
Plants
of the Apocynaceae are often poisonous and are rich in alkaloids or glycosides, especially in the seeds and latex. Some species are valuable sources of medicine, insecticides
, fibers, and rubber.Bingtao Li, Antony J. M. Leeuwenberg & David J. Middleton "Apocynaceae". in Flora of China Vol. 16 Page 143. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org.
Genus Alstonia
Trees
or shrubs
, latex present. Branches whorled
, mostly 4 or 5 together. Leaves whorled, rarely opposite; lateral
veins numerous
, ending in a marginal
vein
. Cymes terminal
, usually 1-5 in thyrses
or compound
umbels, terminal. Flowers white, yellow, or pink. Calyx without glands
inside, lobes
connate
at base
. Corolla salverform
, tube
cylindric
, dilated
in distal half, pubescent
inside, lobes overlapping to right
or left. Stamens included
, inserted
near or above middle
of corolla tube; anthers
ovate
, free
from pistil head
, not caudate
; disc absent or of scales
. Ovaries 2, distinct
or connate, ovules numerous. Follicles 2, free or connate. Seeds oblong
or linear
, long bearded
at both ends; endosperm thin; cotyledons up to twice as long as radicle.
About 60 species: tropical
Asia, Africa, C America, N Australia, Pacific Islands; eight species in China."Alstonia". in Flora of China Vol. 16 Page 154. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org.
Physical Description
Flowers: Bloom Period: September. • Flower Color: pale green
Size/Age/Growth
Size: over 40' tall.
Biology
Growth
Soil: Minimum pH: 6.1 • Maximum pH: 8.5
Sunlight: Sun Exposure: Sun to Partial Shade.
Temperature: Cold Hardiness: 10a, 10b, 11. (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Magnoliophyta
(
)
- Sinnott, 1935 Ex Cavalier-Smith, 1998
- Flowering Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Magnoliopsida
(
)
- Brongniart, 1843
- Dicotyledons
- Subclass:
Lamiidae
(
)
- Takhtajan Ex Reveal, 1992
- Superorder:
Gentiananae
(
)
- Thorne Ex Reveal, 1992
- Order:
Apocynales
(
)
- Bromhead, 1838
- Family:
Apocynaceae
(
)
- A.l. De Jussieu, 1789
- Dogbane Family
- Subfamily:
Rauvolfioideae
(
)
- Tribe:
Alstonieae
(
)
- Genus:
Alstonia
(
)
- R. Brown, Mem. Wern. Nat. Hist. Soc. 1: 75. 1811.
- Alstonia
- Specific epithet:
scholaris
- (L.) R.Br.
- Botanical name: - Alstonia scholaris (L.) R. Br.
- Specific epithet:
scholaris
- (L.) R.Br.
- Genus:
Alstonia
(
- Tribe:
Alstonieae
(
- Subfamily:
Rauvolfioideae
(
- Family:
Apocynaceae
(
- Order:
Apocynales
(
- Superorder:
Gentiananae
(
- Subclass:
Lamiidae
(
- Class:
Magnoliopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Magnoliophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Unambiguous Synonyms
- Alstonia scholaris (L.) R. Br.
- Alstonia scholaris R. Br.
- Echites scholaris L.
- Echites scholaris Linnaeus
- Pala scholaris (L.) Roberty
Notes
Name
Status: Accepted Name
. Latest taxonomic
scrutiny: 05-Apr-2001
Place of publication
: Asclepiadeae 64. 1810 (Mem. Wern. Nat. Hist. Soc. 1:75. 1811)
Name verified on 27-Apr-1995 by ARS Systematic Botanists. Last updated: 26-May-1998
Similar Species
Members of the genus Alstonia
There are approximately 132 species in this genus. Here are just 100 of them:
A. actinophylla · A. acuminata · A. angustifolia · A. angustiloba (Pulai Tree) · A. annamensis · A. balansae · A. batino · A. beatricis · A. beccarii · A. boonei · A. boulindaensis · A. brassii · A. breviloba · A. calophylla · A. ciliata · A. cochinchinensis · A. comptonii · A. congensis · A. constricta · A. constricta var. constricta · A. constricta var. mollis · A. constricta var. montmariensis · A. coriacea · A. costata · A. costata var. fragrans · A. costulata · A. cuneata · A. curtisii · A. deplanchei · A. deplanchei var. ndokoaensis · A. duerckheimiana · A. edulis · A. elliptica · A. esquirolii · A. eximia · A. ficifolia · A. filipes · A. fragrans · A. gilletii · A. glabriflora · A. glaucescens · A. godeffroyi · A. grandifolia · A. guangxiensis · A. henryi · A. iwahigensis · A. kurzii · A. lanceolata · A. lanceolifera · A. lanceolifera var. oleoidea · A. latifolia · A. legouixiae · A. legouixiae var. obtusalabastra · A. legouxiae var. linearis · A. lenormandii · A. lenormandii var. comptonii · A. lenormandii var. minutifolia · A. linearifolia · A. linearis · A. longifolia · A. longissima · A. lucida · A. macrantha · A. macrophylla (Devil Tree) · A. mairei · A. marquisensis · A. micrahtha · A. mollis · A. montana · A. muelleriana · A. muelleriana var. muelleriana · A. muelleriana var. parvifolia · A. neriifolia · A. oblongifolia · A. odontophora · A. oleandraefolia · A. ophioxyloides · A. pachycarpa · A. pacifica · A. pangkorensis · A. parvifolia · A. paucinervia · A. paupera · A. pedicellata · A. penangiana · A. pittieri · A. pittierii · A. plumosa · A. plumosa f. glabra · A. plumosa var. communis · A. pneumatophora · A. polyphylla · A. quaternata · A. reineckeana · A. retusa · A. roeperi · A. rostrata · A. rubiginosa · A. rupestris · A. saligna
Bibliography
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Notes
Contributors
- Bisby, F.A., Y.R. Roskov, M.A. Ruggiero, T.M. Orrell, L.E. Paglinawan, P.W. Brewer, N. Bailly, J. van Hertum, eds (2007). Species 2000 & ITIS Catalogue of Life: 2007 Annual Checklist. Species 2000: Reading, U.K.
- Global Biodiversity Information Facility. Accessed November 15, 2007. http://www.gbif.org Mediated distribution data from 8 providers.
- MBLWHOI Library: Universal Biological Index and Organizer. uBio.org accessed July 20, 2008.
- USDA, ARS, National Genetic Resources Program. Germplasm Resources Information Network - (GRIN) [Online Database]. National Germplasm Resources Laboratory, Beltsville, Maryland. URL (May 05, 2008)
- World Conservation Monitoring Centre 1998. In IUCN 2008. 2008 IUCN Red List of Threatened Species. IUCNRedList.org. Downloaded July 19, 2008.
Data Sources
Accessed through GBIF Data Portal November 15, 2007:
- Australian National Herbarium
- , Australian National Herbarium
- Herbarium of the University of Aarhus, The AAU Herbarium Database
- Missouri Botanical Garden, Missouri Botanical Garden
- National Herbarium of New South Wales, NSW herbarium collection
- National Herbarium of New South Wales, Plants of Papua New Guinea
Identifiers
- Biodiversity Heritage Library NamebankID: 2664137
- Catalogue of Life Accepted Name Code: ITS-184803
- Global Biodiversity Information Facility Taxonkey: 13729507
- Globally Unique Identifier: urn:lsid:ipni.org:names:105973-3
- GRIN Nomen Number: 2688
- International Plant Names Index (IPNI) ID: 76587-1
- IUCN ID: 32295
- Zipcode Zoo Species Identifier: 407769
