- Alnus rubra is the largest alder in North America north of Mexico; it often forms extensive stands along streams and on low-lying flood plains in the Pacific Northwest. The strongly revolute margins of its leaf blades make it easily distinguished from all of the other alders in the flora . It is an important commercial tree ; the wood is used to make inexpensive furniture, small wooden items, and paper pulp. [source]
- Native Americans used various parts of plants of Alnus rubra medicinally as a purgative , an emetic, for aching bones, headaches, coughs, biliousness, stomach problems, scrofula sores, tuberculosis, asthma, and eczema, and as a general panacea (D. E. Moerman 1986). [source]
Common Names in English:
Oregon Alder, Red Alder
deciduous, monoecious. Stipules present, free
, often deciduous, rarely persistent
. Leaves alternate, simple
, usually doubly serrate, rarely simply serrate, lobulate
, or entire; veins pinnate. Flowers unisexual
. Male inflorescence precocious
, pendulous, with numerous
overlapping bracts; each bract usually subtending
a small dichasium with 1-3 male flowers; stamens as many as and opposite sepals or, if sepals obsolete
, then stamens of inflorescence to 20; filaments
very short, connate
or nearly so; anthers
2-loculed, thecae connate or separate, opening by longitudinal
slits. Female inflorescence pendulous or erect
, with numerous overlapping bracts; each bract subtending a small dichasium with 2 or 3 flowers; calyx with 1-6 scalelike lobes
, or obsolete; petals absent; ovary inferior, 2-loculed; styles 2, free; ovules 2, or 1 by abortion
, pendulous from near apex of each locule. Fruit a nut or nutlet
or not. Seed 1, with straight embryo and flat or thickened cotyledons, without endosperm.
Six genera and 150-200 species: mainly in Asia, Europe, and North and South America; six genera (one endemic) and 89 species (56 endemic) in China.
Because of evolutionary divergence within the Betulaceae, the family has often been divided into tribes (i.e. , Betuleae, Carpineae, and Coryleae) or more recently into subfamilies (J. J. Furlow, J. Arnold Arbor . 71: 1-67. 1990) .
, to 35 m
usually several, branching excurrent to deliquescent. Bark
of trunks and branches light gray to dark brown, thin, smooth
, close; lenticels
often present, pale
, sometimes horizontally expanded. Wood
nearly white, turning reddish upon exposure to air
, moderately light and soft, texture
fine. Branches, branchlets
, and twigs
nearly 2-ranked to diffuse
; young twigs uniform or ( Alnus subg. Alnobetula ) differentiated into long and short shoots
. Winter buds
(nearly sessile in Alnus subg. Alnobetula ), narrowly to broadly ovoid
, apex acute to rounded
, or ( Alnus subg. Alnobetula ) several, imbricate, smooth, or ( Alnus subg. Clethropsis ) sometimes none. Leaves borne on long or short shoots, 3-ranked to nearly 2-ranked. Leaf blade
, thin to leathery, base
variable, cuneate to rounded, margins
doubly serrate, serrate, serrulate
, or nearly entire, apex variable, acute to obtuse
or acuminate to rounded; surfaces glabrous
, abaxially sometimes resinous-glandular. Inflorescences: staminate
, in racemose clusters
or ( Alnus subg. Clethropsis ) solitary, formed ( Alnus subg. Alnus and Clethropsis ) during previous growing season
and exposed or enclosed in buds during winter, or ( Alnus subg. Clethropsis ) formed and expanding during same growing season, expanding before or with leaves; pistillate
to staminate catkins, solitary or in relatively small racemose clusters, erect
to nearly pendulous, ovoid to ellipsoid, firm; scales and flowers crowded, developing and maturing at same time as staminate catkins. Staminate flowers
in catkins, 3 per scale; stamens (3--) 4(--6) ; anthers
undivided. Pistillate flowers usually 2 per scale. Infructescences
erect or pendulous; scales persistent long after release
of fruits, with 5 lobes
, greatly thickened, woody. Fruits tiny samaras, lateral wings 2, leathery or membranaceous
, reduced or essentially absent in some species. x
Species ca. 25 (8 sp: forested temperate and boreal Northern Hemisphere; North America; Asia.
Alders resemble birches but are easily distinguished from them by the infructescences, which consist of persistent , 5-lobed, woody scales (versus deciduous, 3-lobed, thin scales). Except in members of Alnus subg. Alnobetula Petermann (which have nearly sessile buds with several imbricate scales), alders are also distinctive in their stipitate buds bearing two stipular scales. The fruits, borne two to a scale, are laterally winged , although the wings are sometimes reduced or absent.
The genus is diverse , including several very distinct lines of specialization. The shrubby or arborescent Alnus subg. Alnus is characterized by winter buds with long stalks and two valvate scales, inflorescences borne in racemose clusters, and development of both pistillate and staminate inflorescences during the growing season prior to anthesis , with these fully exposed during winter. It includes the common A. rubra, A. incana, A. oblongifolia, and A. serrulata. Alnus subg. Alnobetula (represented in North America by three subspecies of A. viridis ) consists of shrubby species of cold-climate regions. In this group, the buds are nearly sessile and covered by several imbricate scales. Both staminate and pistillate catkins are formed the season before anthesis, but only the staminate ones are exposed during winter. The predominantly Asian Alnus subg. Clethropsis (Spach) Regel is represented in America by a single species, A. maritima, a small tree or large shrub of stream banks, marshes, and the shores of shallow lakes . Members of this group are unique in that they bloom in autumn rather than spring . They also differ from other native species in Alnus in having essentially naked buds, leaves with semicraspedodromous venation (i.e. , with the secondary veins branching and anastomosing with each other near the margin before reaching the teeth), and solitary pistillate inflorescences borne in the axils of foliage leaves. All of the alders associate symbiotically with species of the actinomycete Frankia, leading to the formation of nodules on the roots of the plants and the fixation of atmospheric nitrogen.
Species Alnus rubra
Trees , to 28 m ; trunks often several, crowns narrow or pyramidal . Bark gray, smooth , darkening and breaking into shallow rectangular plates in age; lenticels inconspicuous. Winter buds stipitate , ellipsoid , 6--10 mm, apex rounded , long; stalks 2--8 mm; scales 2--3, outer 2 equal and valvate , usually heavily resin-coated. Leaf blade ovate to elliptic , 6--16 × 3--11 cm, leathery, base broadly cuneate to rounded, margins strongly revolute , deeply doubly serrate or crenate , with distinctly larger secondary teeth, apex acute to obtuse ; surfaces abaxially glabrous to sparsely pubescent . Inflorescences formed season before flowering and exposed during winter; staminate catkins in 1 or more clusters of 2--6, 3.5--14 cm; sistillate catkins in 1 or more clusters of 3--8. Flowering before new growth in spring . Infructescences ovoid to nearly globose , 1--3.5 × 0.6--1.5 cm; peduncles 1--10 mm. Samaras ovate or elliptic, wings much narrower than body, irregularly elliptic to obovate , leathery. 2 n = 28. Flowering early spring. [source]
Size: 30-40' tall.
Typically found at an altitude of 0 to 2,174 meters (0 to 7,133 feet).
Culture: Space 20-30' apart.
Soil: Minimum pH: 4.5 • Maximum pH: 7.5
Sunlight: Sun Exposure: Sun to Partial Shade.
Temperature: Cold Hardiness: 8b, 9a, 9b, 10a, 10b. (map)
- Whittaker & Margulis,1978
- Haeckel, 1866
- Cavalier-Smith, 1981
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Kenrick & Crane, 1997
- Brongniart, 1843
- Takhtajan, 1967
- Superorder: Faganae () - (Engler, 1892) Takhtajan, 1997
- Subclass: Rosidae () - Takhtajan, 1967
- Class: Spermatopsida () - Brongniart, 1843
- Infraphylum: Radiatopses () - Kenrick & Crane, 1997
- Subphylum: Euphyllophytina ()
- Phylum: Tracheophyta () - Sinnott, 1935 ex Cavalier-Smith, 1998 - Vascular Plants
- Subkingdom: Viridaeplantae () - Cavalier-Smith, 1981
- Kingdom: Plantae () - Haeckel, 1866 - Plants
A. rubra var. pinnatisecta Starker • Alnus glutinosa var. quercifolia Hartig • Alnus oregana Nutt. • Alnus oregona Nuttall • Alnus rubra f. pinnatisecta (Starker) Rehder • Alnus rubra var. pinnatisecta Starker • Alnus washinhtonia Wetzel
Publishing author : Desf. ex Steud. Publication : Nomencl. Bot. [Steudel] 29 1821
Members of the genus Alnus
ZipcodeZoo has pages for 28 species, subspecies, varieties, forms, and cultivars in this genus:
A. acuminata (Alder) · A. acuminata glabrata (Mexican Alder) · A. cordata (Italian Alder) · A. fallacina (Alder) · A. glutinosa (Black Alder) · A. glutinosa glutinosa (European Black Alder) · A. glutinosa 'Imperialis' (Black Alder) · A. glutinosa 'Pyramidalis' (Black Alder) · A. hirsuta (Manchurian Alder) · A. incana (Gray Alder) · A. incana incana (Speckled Alder) · A. incana rugosa (Gray Alder) · A. incana tenuifolia (Gray Alder) · A. japonica (Japanese Alder) · A. maritima (Seaside Alder) · A. nepalensis (Indian Alder) · A. nitida (West Himalayan Alder) · A. oblongifolia (Arizona Alder) · A. rhombifolia (Alder) · A. rubra (Oregon Alder) · A. rubra f. pinnatisecta (Oregon Alder) · A. serrulata (Alder) · A. viridis (Green Alder) · A. viridis crispa (Green Alder) · A. viridis fruticosa (Green Alder) · A. viridis sinuata (Green Alder) · A. viridis viridis (European Green Alder) · A. x fallacina (Alder)
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- A catalogue of the Cretaceous and Tertiary plants of North America by Frank Hall Knowlton. Washington: G.P.O., 1898. url .
- A class-book of botany, designed for colleges, academies, and other seminaries. Illustrated by a flora of northern, middle, and western states; particularly of the United States north of the Capitol, lat. 38 3/4. Claremont, N. H., Manufacturing Co., S. Ide, agent, 1850. url p. 499.
- A dictionary of English names of plants applied in England and among English-speaking people to cultivated and wild plants, trees, and shrubs, by William Miller; in two parts, English-Latin and Latin-English. London, J. Murray, 1884. url p. 2.
- A flora of California, by Willis Linn Jepson. San Francisco, Calif., Cunningham, Curtis & Welch, 1909- url p. 347, p. 347, p. 348, p. 423.
- A flora of northwest America containing brief descriptions of all the known indigenous and naturalized plants growing without cultivation north of California, west of Utah, and south of British Columbia / Portland, Or.: [s.n.], 1901. url .
- A flora of northwest America, containing brief descriptions of all the known indigenous and naturalized plants growing without cultivation north of California, west of Utah, and south of British Columbia. Portland, Or., 1903. url .
- A flora of northwest America: containing brief descriptions of all the known indigenous and naturalized plants growing without cultivation north of California, west of Utah, and south of British Columbia / by Thomas Howell. Vol. 1, Phanerogamae. Portland, Or.: [s.n.], 1903. url p. 614, p. 614.
- A manual of dangerous insects likely to be introduced in the United States through importations. Ed. by W. Dwight Pierce, entomologist, southern field crop insect investigations. Washington: Govt. print. off., 1918. url p. 11.
- A popular California flora, or Manual of botany for beginners. Containing descriptions of flowering plants growing in central California, and westward to the ocean. With illustrated introductory lessons, especially adapted to San Francisco, A.L. Bancroft, 1882. url p. 106.
- A popular California flora, or, Manual of botany for beginners. With illustrated introductory lessons, especially adapted to the Pacific Coast. To which is added an Analytical key to west coast botany. Containing brief descripti San Francisco, The Whitaker & Ray Co., 1896. url p. 106.
- A popular California flora: or, Manual of botany for beginners with illustrated introductory lessons, especially adapted to the Pacific Coast; to which is added an analytical key to west coast botany, containing brief descripti by Volney Rattan. San Francisco: Whitaker & Ray, 1896. url p. 106.
- A popular California flora: or, manual of botany for beginners. ..: to which is added an Analytical key to west coast botany. .. / San Francisco: Bancroft, 1892. url .
- A preliminary catalogue of the flora of Vancouver and Queen Charlotte Islands. Victoria, B.C., Printed by W. H. Cullin, 1921. url p. 32.
- A provisional host-index of the fungi of the United States, by W.G. Farlow and A.B. Seymour. Cambridge, 1888-91. url p. 106.
- A text-book of mycology and plant pathology / Philadelphia: P. Blakiston's Son & Co.c1917. url .
- American medicinal plants; an illustrated and descriptive guide to the American plants used as homopathic remedies: their history, preparation, chemistry and physiological effects. By Charles F. Millspaugh. 1887 New York, Boericke & Tafel, c1887. url page 143- 2.
- American plants. .. [Descriptions, bibliographical notes, synonymy, and other information, comp. from many sources] Charles Russell Orcutt, editor. San Diego, Calif., -1910. url p. 457.
- An illustrated dictionary of medicine, biology and allied sciences. .. by George M. Gould. 5th ed., with additions and corrections. PhiladelphiaP. Blakiston's Son1907 url p. 62.
- An illustrated flora of the Pacific States: Washington, Oregon, and California. Stanford University, Stanford University Press, 1923- url p. 513.
- An introduction to entomology. Ithaca, Comstock, 1888. url p. 166.
- Annotated list of the ferns and flowering plants of New York state, by Homer D. House. Albany, The University of the state of New York, 1924. url p. 272.
- Annual report / Albany: University of the State of New York, 1902-1918. url p. 56.
- Annual report of the Board of Regents of the Smithsonian Institution. Washington: Smithsonian Institution, -1965. url p. 454, p. 454.
- Annual report of the Bureau of American Ethnology to the Secretary of the Smithsonian Institution. Washington: U. S. Govt. Print. Off., 1897- url p. 145, p. 44, p. 483, p. 520.
- Annual report of the Regents / Albany: J.B. Lyon, State Printer, 1890-1903. url p. 408, p. 467.
- Arctic searching expedition: a journal of a boat-voyage through Rupert's Land and the Arctic Sea, in search of the discovery ships under command of Sir John Franklin. With an appendix on the physical geography of North America. By Sir John Richardson. .. London, Longman, Brown, Green, and Longmans, 1851. url p. 311.
- Bartonia;proceedings of the Philadelphia botanical club. .. 62 2004 Philadelphia, Philadelphia Botanical Club, Academy of Natural Sciences. url p. 64, p. 65, p. 68.
- Biosphere Reserves, Compilation 4, October 1986: programme on man and the biosphere (MAB) IUCN Conservation Monitoring Centre url p. 462.
- Botany Cambridge, Mass., John Wilson and Son, 1880 url p. 483, p. 80.
- Botany of Yakutat Bay, Alaska / by Frederick Vernon Coville; with a field report by Frederick Funston. Washington: GPO, 1895 [i.e. 1896] url , , p. 332.
- Botany. By W. H. Brewer, Sereno Watson, and Asa Gray. Boston, Little, Brown, 1880. url p. 483, p. 80.
- Botany. Cambridge, Mass.Welch, Bigelow, University Press, 1876-80. url p. 80.
- Britton, N. L. (ed.). North American flora. [New York]New York Botanical Garden. url p. 680.
- Bulletin - United States National Museum. Washington: Smithsonian Institution Press, [etc.];1877-1971. url p. 152, p. 240, p. 345, p. 346.
- Bulletin of the California Academy of Sciences. [S.l.: The Academy], 1884- url p. 256, p. 260, p. 261, p. 266, p. 351, p. 444.
- Bulletin of the Illinois State Laboratory of Natural History. Bloomington, Ill.: The Laboratory, 1876-1918. url , p. 561.
- Bulletin of the Torrey Botanical Club. 21 1894 New York: Torrey Botanical Club, 1870-1996 url p. 160, p. 580, p. 641, p. 76.
- Bulletin of the U.S. Department of Agriculture. [Washington, D.C.?]: Supt. of Docs., G.P.O., 1913-1923. url p. 33.
- Bulletin. California Academy of Sciences. San Francisco. San Francisco url p. 256, p. 260, p. 265, p. 266, p. 351, p. 444.
- Butterflies and moths of Newfoundland and Labrador: the macrolepidoptera / [Ottawa]: Agriculture Canada, 1980. url p. 123.
- C. E. Hobbs Botanical hand-book of common local, English, botanical and pharmacopial names arranged in alphabetical order, of most of the crude vegetable drugs, etc., in common use. Especially designed as a reference book for druggists and apothecaries. Comp. and pub. by Charles E. Hobbs. Boston, Printed by C.C. Roberts, 1876. url p. 117, p. 142, p. 235.
- C.E. Hobbs' Botanical hand-book: of common local, English, botanical and pharmacopœial names arranged in alphabetical order, of most of the crude vegetable drugs, etc., in common use: their properties, productions and uses in an abb Boston: Printed by C.C. Roberts, 1876. url , , , .
- California fish and game. [San Francisco, etc.]: State of California, Resources Agency, Dept. of Fish and Game. url p. 207, p. 248, p. 266.
- Canadian forests, forest trees, timber and forest products Montreal: Dawson, 1884. url .
- Canadian forests: forest trees, timber and forest products / by H. B. Small. Montreal: Dawson, 1884. url p. 37.
- Catalog of hymenoptera in America north of Mexico / prepared cooperatively by specialists on the various groups of Hymenoptera under the direction of Karl V. Krombein. .. [et al.]. Washington: Smithsonian Institution Press, 1979- url p. 99.
- Catalogue of Canadian plants. .. Montreal [etc.]1883-1902. url , , , , , , , , p. 102, p. 106, p. 122, p. 122, p. 151, p. 151, p. 162, p. 162, p. 167, p. 167, p. 169, p. 355, p. 43, p. 43, p. 437, p. 437.
- Catalogue of the collections in the museum of the Pharmaceutical Society of Great Britain compiled by E.M. Holmes. London, The Society, 1878. url p. 272.
- Contributions from the United States National Herbarium 11 1906 Washington, D.C.: Smithsonian Institution Press, 1890- url p. 219, p. 328, p. 332, p. 345, p. 350, p. 352.
- Contributions from the University of Michigan Herbarium. 5 1941 Ann Arbor: University Herbarium, University of Michigan, 1939- url p. 41, p. v.
- Contributions to the natural history of Alaska results of investigations made chiefly in the Yukon district and the Aleutian Islands; conducted under the auspices of the Signal Service, United States Army, extending from May, 1874, to August, 1881 prepared under the direction of W.B. Hazen, Chief Signal Officer of the Army, by L. M. Turner. Washington: G.P.O., 1886. url .
- Cyclopedia of American horticulture, comprising suggestions for cultivation of horticultural plants, descriptions of the species of fruits, vegetables, flowers and ornamental plants sold in the United by L. H. Bailey. .. assisted by William Miller. .. and many expert cultivators and botanists. London: The Macmillan company, 1909. url p. 48.
- Directory of Wetlands of International Importance. IUCN url p. 757.
- Disturbance and forest health in Oregon and Washington / technical coordinators: Sally Campbell, Leon Liegel; editor: Martha H. Brookes. Portland, Or.: USDA Forest Service, Pacific Northwest Research Station,  url p. 88.
- Entomological news, and proceedings of the Entomological Section of the Academy of Natural Sciences of Philadelphia. Philadelphia[Entomological Rooms of the Academy of Natural Sciences] url p. 357.
- Erigenia: journal of the Illinois Native Plant Society. Carbondale, Ill.: The Society, 1982- url p. 69.
- FWS/0BS. [Washington]Fish and Wildlife Service, U.S. Dept. of the Interior. url p. 61.
- Fifty years of botany; golden jubilee volume of the Botanical Society of America, edited by William Campbell Steere. New York, McGraw-Hill, 1958. url p. 518.
- Flora of the Queen Charlotte Islands. [Ottawa: Queen's Printer], 1968-. url p. 144, p. 154, p. 306, p. 307, p. 53, p. 82, p. 84, p. 85, p. 87.
- Flora of the state of Washington / by Charles V. Piper. Washington: G.P.O., 1906 url p. 219.
- Flora of the state of Washington. By Charles V. Piper. Washington, Govt. Print. Off., 1906. url p. 219.
- Forestry research report. Urbana, Ill., Agricultural Experiment Station, Dept. of Forestry, University of Illinois at Urbana-Champaign. url , .
- Garden and forest; a journal of horticulture, landscape art and forestry. New York: The Garden and forest publishing co., 1888-97. url p. 343, p. 47, p. 484, p. 632.
- Great Basin naturalist memoirs. 1992 [Provo, Utah]Brigham Young University, 1976-1992. url p. 1247.
- Identification of the economic woods of the United States, including a discussion of the structural and physical properties of wood, New York, John Wiley & sons, inc.; [etc., etc.]1919. url , p. 107.
- Inland wetlands of the United States: evaluated as potential registered natural landmarks / Richard H. Goodwin, William A. Niering. [Washington]: National Park Service: may be purchased from the Supt. of Docs., Govt. Print. Off., 1975. url p. 18.
- Journal of ethnobiology. 8-9 1988-1989 Flagstaff, Ariz.: Center for Western Studies, 1981- url p. 103, p. 178, p. 183, p. 250, p. 7.
- Journal of the proceedings of the Linnean Society. Botany. 7 1864 London: Longman, Brown, Green, Longmans, & Roberts: ||Williams and Norgate, 1857-1864. url p. 134.
- Key to important woody plants of eastern Oregon and Washington, by Doris W. Hayes and George A. Garrison. Washington, U.S. Govt. Print. Off., 1960. url p. 120, p. 214, p. 22.
- Leaf-mining insects / by James G. Needham, Stuart W. Frost, Beatrice H. Tothill. Baltimore: The Williams & Wilkins company, [c1928] url p. 302.
- Leaflets of western botany. San Fransisco:[J. T. Howell], 1932-1966. url .
- Manual of the trees of North America (exclusive of Mexico) by Charles Sprague Sargent, with seven hundred and eighty-three illustrations from drawings by Charles Edward Faxon and Mary W. Gill. Boston and New York, Houghton Mifflin company, 1922. url p. 222, p. 899, p. 899.
- Memoirs of the Torrey Botanical Club. 9 1900 Durham, N.C.: Published for the Club by the Seeman Printery, 1889- url p. 133, p. 228, p. 264.
- Montana wetlands management study / prepared for Montana Department of Natural Resources and Conservation. Denver, Colo.: Bureau of Reclamation,  url p. 10, p. 44, p. 47.
- Muhlenbergia. Chico, Calif. [etc] url , p. 148, p. 153.
- Museum bulletin / Albany, N.Y.: University of the State of New York, 1908-1916. url p. 54, p. 56.
- National list of scientific plant names. [Washington, D.C.]: U.S. Dept. of Agriculture, Soil Conservation Service, 1982- url p. 17.
- Native woody plants of the United States, their erosion-control and wildlife values. Washington, U. S. Govt. print. off., 1938. url , , , , p. 46.
- Natural history of New York. Albany: D. Appleton: 1842-1894. url p. 202, p. 202, p. 203.
- New illustrated encyclopedia of gardening, unabridged. Edited by T.H. Everett, with contributions from twenty horticulturists and authorities in the United States and Canada. New York, Greystone Press[1964-] url p. 81.
- North American trees (exclusive of Mexico and tropical United States) A handbook designed for field use, with plates and distribution maps. Ames, Iowa State University Press url p. 156, p. 157, p. 159.
- North American trees: being descriptions and illustrations of the trees growing independently of cultivation in North America, north of Mexico and the West Indies / New York: H. Holt and Co., 1908. url .
- North American trees: guide to Charles F. Millspaugh Hall, by Samuel J. Record. Chicago, 1934. url p. 118, p. 62.
- North American trees; being descriptions and illustrations of the trees growing independently of cultivation in North America, north of Mexico and the West Indies; with the assistance of John Adolph Shafer. New YorkHolt1908 url p. 260, p. 262.
- Note on the distribution of some of the more important trees of British Columbia by George M. Dawson. S.l.: s.n., 1880? url .
- Pesticides monitoring journal. [Washington, D.C.: For sale by the Supt. of Docs., U.S. G.P.O., 1967-1981. url p. 213.
- Phytologia. Bronx Park, New York, H.A. Gleason and H.N. Moldenke, url p. 15.
- Pittonia: a series of papers relating to botany and botanists /by Edward L. Greene. 3 1896 Berkeley, Calif. [etc.]: Doxey & Co. [etc.], 1887-1905. url p. 70.
- Plant species of concern / Helena, Mont.: Montana Natural Heritage Program, c2003. url .
- Potter's cyclopaedia of botanical drugs and preparations / by R.C. Wren. London: Potter & Clarke,  url p. 277, p. viii.
- Potter's cyclopedia of botanical drugs and preparations / by R. C. Wren. London: Potter & Clarke, [1900-1950?]. url p. 152, p. viii.
- Practical forestry: a treatise on the propagation, planting, and cultivation, with a description, and the botanical and popular names of all the indigenous trees of the United States, both evergreen and deciduous, togeth New York: Orange Judd Co., 1914, c1884. url .
- Proceedings of the Academy of Natural Sciences of Philadelphia. Philadelphia, Academy of Natural Sciences of Philadelphia url p. 311, p. 366, p. 41, p. 91, p. 91.
- Proceedings of the American Academy of Arts and Sciences. Boston: Metcalf and Co., 1846-1958 url p. 402, p. 662.
- Proceedings of the Entomological Society of Washington. Washington, etc.: Entomological Society of Washington url p. 1014, p. 191, p. 222, p. 247, p. 269, p. 281, p. 414, p. 463, p. 659.
- Proceedings of the United States National Museum. 17 1894 Washington: Smithsonian Institution Press, [etc.] url p. 220, p. 442, p. 649.
- Proceedings. Halifax url p. 497.
- Range plant handbook. Washington, U. S. Govt. print. off., 1937. url p. vi.
- Report of the Director to the Board of Trustees for the year. .. / Chicago Natural History Museum. [Chicago]: The Museum, -1963. url p. 50.
- Report of the State Botanist. Albany, N.Y.: University of the State of New York, 1898-1925. url , p. 56.
- Report of the State Entomologist on injurious and other insects of the state of New York. Albany: Weed, Parsons and Co.; University of the State of New York, 1882- url p. 408, p. 467.
- Report on an exploration in the Yukon district, N.W.T., and adjacent northern portion of British Columbia, 1887 by George M. Dawson. Montreal: Dawson, 1888. url , , .
- Report on the forest wealth of Canada Ottawa: S.E. Dawson, 1895. url .
- Report on the forests of North America (exclusive of Mexico) / by Charles S. Sargent. Washington [D.C.]: G.P.O., 1884. url p. 163, p. 164, p. 221, p. 582.
- Report on the injurious and other insects of the State of New York. Albany: State Entomologist, 1882-1898. url p. 467.
- Report upon forestry. From the committee appointed to memorialize Congress and the state legislatures, regarding the cultivation of timber and the preservation of forests, prepared under the direction of the Commissioner of Agriculture in pursuance of an Act of Congress approved August l5, 1876, by Franklin B. Hough. Washington, Govt. Print. Off., 1878. url p. 591.
- Select extra-tropical plants readily eligible for industrial culture or naturalisation with indications of their native countries and some of their uses / by Baron Ferd. von Mueller. Melbourne: R. S. Brain, govt. printer, 1895. url p. 33.
- Selected vertebrate endangered species of the seacoast of the United States / prepared by National Fish and Wildlife Laboratory, U.S. Fish and Wildlife Service. [Washington]: The Service: [for sale by the Supt. of Docs., U.S. Govt. Print. Off.], 1980. url p. 2.
- Special scientific report--wildlife / U.S. Department of the Interior, Fish and Wildlife Service. Washington, D.C.: The Service, 1949- url p. 642.
- Technical series. Washington, Govt. print. off., 1895-1915. url p. 6, p. 7.
- The American journal of science and arts. New-Haven: S. Converse, url p. 33.
- The American journal of science. 45 1843 New Haven, Kline Geology Laboratory, Yale University. url p. 33.
- The Bradley bibliography; a guide to the literature of the woody plants of the world published before the beginning of the twentieth century; Cambridge, Riverside Press, 1911-18. url p. 117.
- The Canadian field-naturalist. Ottawa, Ottawa Field-Naturalists' Club. url , p. 121, p. 151, p. 188, p. 213, p. 223, p. 236, p. 307, p. 308, p. 316, p. 348, p. 356, p. 37, p. 401, p. 423, p. 431, p. 455, p. 456, p. 466, p. 49, p. 55, p. 626, p. 647, p. 679, p. 758, p. 772.
- The Canadian naturalist and quarterly journal of science: with proceedings of the Natural History Society of Montreal. Montreal: Dawson, 1869-1883. url p. 331.
- The Gardeners' chronicle and agricultural gazette. London: published for the proprietors, 1844-1873. url p. 1150, p. 1432.
- The Gardeners' chronicle: a weekly illustrated journal of horticulture and allied subjects. London: [Gardeners Chronicle], 1874-1955. url p. 684.
- The Great Basin naturalist. Provo, Utah: M.L. Bean Life Science Museum, Brigham Young University, 1939-1999. url p. 228, p. 34.
- The IUCN Mammal Red Data Book. Part 1: threatened mammalian taxa of the Americas and the Australasian zoogeographic region (excluding Cetacea) IUCN url p. 470, p. 470.
- The Journal of geology. Chicago: University of Chicago Press, 1893- url p. 532.
- The Proceedings and transactions of the Nova Scotian Institute of Science. Halifax, Nova Scotian Institute of Science. url p. 154, p. 497.
- The University of Kansas science bulletin. 24 1936 [Lawrence]: University of Kansas, 1902-1996. url p. 135, p. 231, p. 231, p. 265, p. 265, p. 271, p. 271, p. 78, p. 82, p. 86, fig. 21, page 87, p. 88, p. 91.
- The century supplement to the dictionary of gardening, a practical and scientific encyclopaedia of horticulture for gardeners and botanists / by Geo. Nicholson. .. [et al.]. Hyde Park, Mass.: Geo. T. King; 1901. url p. 34.
- The complete herbalist, or The people their own physicians by the use of nature's remedies: describing the great curative properties found in the herbal kingdom. .. / by Dr. O. Phelps Brown. Jersey City, N. J.: Published by the author, 1897. url p. 36.
- The farmer's dictionary: a vocabulary of the technical terms recently introduced into agriculture and horticulture from various sciences, and also a compendium of practical farming, the latter chiefly from the works of the Re edited by D. P. Gardner. New York: Harper, 1846. url p. 585.
- The naturalist in Vancouver Island and British Columbia London: R. Bentley, 1866. url .
- The reflector: a brief resume of facts condensed for ready reference embracing cookery, household economy, poultry, stock and farm; together with diseases of man, beast and fowl and history cause, symptoms and ratio by W.S. Hames. Oklahoma City: Printers Publishing Co., c1912 url p. 103.
- The silva of North America a description of the tree which grow naturally in North America exclusive of Mexico / by Charles Sprague Sargent; illustrated with figures and analyses drawn from nature by Charles Edward Faxon. Boston;Houghton, Mifflin, 1894 url , .
- The silva of North America: a description of the trees which grow naturally in North America exclusive of Mexico /by Charles Sprague Sargent. .. illustrated with figures and analyses drawn from nature by Charles Edward Faxon. .. 14 1902 Boston, New York: Houghton, Mifflin and company, 1891-1902. url p. 111, p. 185, p. 61, p. 69, p. 73.
- The standard cyclopedia of horticulture; a discussion, for the amateur, and the professional and commercial grower, of the kinds, characteristics and methods of cultivation of the species of plants grown in the regions of the United States a Illustrated with colored plates, four thousand engravings in the text, and ninety-six full-page cuts. New York, Macmillan, 1919 [c1914] url p. 251.
- The timbers of commerce and their identification, London, W. Rider & son, limited, 1918. url .
- The trees of California. San FranciscoCunningham, Curtis & Welch url p. 148, p. 149, p. 15, p. 150, p. 151.
- The trees of Great Britain & Ireland / by Henry John Elwes and Augustine Henry. Edinburgh: Priv. print., 1906-13. url , p. 1949, p. 936, p. 956.
- The woods of the United States. With an account of their structure, qualities, and uses. With geographical and other notes upon the trees which produce them. By C.S. Sargent. New York: D. Appleton and Co., 1885. url p. 156, p. 161, p. 163, p. 167, p. 171, p. 175, p. 178, p. 98.
- The woods of the United States: with an account of their structure, qualities and uses; with geographical and other notes upon the trees which produce them / by C.S. Sargent. New York: Appleton, 1885. url p. 156, p. 161, p. 163, p. 167, p. 171, p. 175, p. 178, p. 98.
- Timber management: a programmatic environmental analysis record for BLM-administered lands in western Oregon / Office, 1974 url p. 46.
- Transactions of the Canadian Institute. Toronto: Canadian Institute. url p. 116.
- Transactions of the Massachusetts Horticultural Society. [S.l.: s.n.], 1843-1920. url p. 178.
- Transactions of the Scottish Arboricultural Society. Edinburgh: The Society, -1887. url p. 451.
- Tree ancestors; a glimpse into the past, by Edward Wilber Berry. Baltimore, Williams & Wilkins, 1923. url p. 118.
- Trees: the yearbook of agriculture 1949. Washington, D.C.: U.S. Dept. of Agriculture: 1949. url p. 767, p. 811, p. 917.
- Trees: the yearbook of agriculture, 1949 / the United States Department of Agriculture. Washington, D.C.: United States Government Printing Office, 1949 url p. 767, p. 811, p. 917.
- Utilization of hardwoods growing on southern pine sites / Peter Koch. Washington, D.C.: For sale by the Supt. of Docs., U.S. G.P.O.,  url p. 2691, p. 3100.
- Washington State Coastal Zone Management Program amendment no. 3: approval of the Grays Harbor estuary management plan: program draft environmental impact statement. [Olympia, Wash.?]: Grays Harbor Regional Planning Commission; 1983. url p. 10, p. 9.
- West Coast botany: an analytical key to the flora of the Pacific Coast in which are described over eighteen hundred species of flowering plants growing west of the Sierra Nevada and Cascade crests, from San Diego to Pug by Volney Rattan. San Francisco: Whitaker & Ray, 1898. url p. 71.
- Wood; a manual of the natural history and industrial applications of the timbers of commerce. LondonE. Arnold1908 url p. 125.
- Woody plants of the western national parks; containing keys for the identification of trees and shrubs, by Virginia Long Bailey and Harold Edwards Bailey. Notre Dame, Ind., University Press, 1949. url p. 259, p. 3, p. 4.
- Year book / Carnegie Institution of Washington. Washington, D.C.: Carnegie Institution of Washington, 1903- url p. 350.
- Zoe:a biological journal. 2 1891 San Diego, Calif. [etc.]Zoe Publishing Co. url p. 257, p. 274.
- Li Pei-chun & Cheng Sze-hsu. 1979. Betulaceae. In: Kuang Ko-zen & Li Pei-chun, eds., Fl. Reipubl. Popularis Sin. 21: 44-137.
- Furlow, J. J. 1979. The systematics of the American species of Alnus (Betulaceae). Rhodora 81: 1--121, 151--248.
- Hylander, N. 1957. On cut-leaved and small-leaved forms of Alnus glutinosa and A. incana. Svensk Bot. Tidskr. 51: 437--453.
- Murai, S. 1964. Phytotaxonomical and geobotanical studies on gen. Alnus in Japan (III). Taxonomy of whole world species and distribution of each sect. Bull. Gov. Forest Exp. Sta. 171: 1--107.
- Trappe, J. M., J. F. Franklin, R. F. Tarrant, and G. M. Hansen, eds. 1968. Biology of Alder.... Portland.
- "Alnus rubra". in Flora of North America Vol. 3. Published by Oxford University Press. Online at EFloras.org.
- Brands, S.J. (comp.) 1989-2007. Systema Naturae 2000. The Taxonomicon. Universal Taxonomic Services, Amsterdam, The Netherlands. Accessed March 28, 2007.
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 10, 2012.
- Global Biodiversity Information Facility. Accessed November 11, 2007. http://www.gbif.org Mediated distribution data from 11 providers.
- Ruggiero M., Gordon D., Bailly N., Kirk P., Nicolson D. (2011). The Catalogue of Life Taxonomic Classification, Edition 2, Part A. In: Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist (Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D., eds). DVD; Species 2000: Reading, UK.
- The International Plant Names Index. Accessed Dec 27, 2011.
- USDA, ARS, National Genetic Resources Program. Germplasm Resources Information Network - (GRIN) [Online Database]. National Germplasm Resources Laboratory, Beltsville, Maryland. URL (April 25, 2008)
- World Checklist of Selected Plant Families. Release date: November 27, 2009
Accessed through GBIF Data Portal November 11, 2007:
- Berkeley Natural History Museums, University and Jepson Herbaria DiGIR provider
- Canadian Museum of Nature, Canadian Museum of Nature Herbarium
- Forest Research Institute, Department of Natural Forests, Herbarium
- Missouri Botanical Garden, Missouri Botanical Garden
- Oregon State University, Vascular Plant Collection
- UK National Biodiversity Network, Botanical Society of the British Isles - Vascular Plants Database
- USDA PLANTS, USDA PLANTS Database
- University of Alaska Museum of the North, University of Alaska Museum of the North Herbarium
- University of Washington Burke Museum, Vascular Plant Collection - University of Washington Herbarium
- Utah State University, USU-UTC Specimen Database
- Biodiversity Heritage Library NamebankID: 2646011
- Catalogue of Life Accepted Name Code: Kew-6584
- Global Biodiversity Information Facility Taxonkey: 4490920
- Globally Unique Identifier: urn:lsid:ipni.org:names:294987-1
- GRIN Nomen Number: 105118
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 19474
- International Plant Names Index (IPNI) ID: 294986-1
- Natural Heritage Network Species Identifier: PDBET01060
- U.S.D.A. Plant Symbol: ALOR
- Zipcode Zoo Species Identifier: 19685
- Pei-chun Li & Alexei K. Skvortsov "Betulaceae". in Flora of China Vol. 4 Page 286. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- "Alnus". in Flora of North America Vol. 3. Oxford University Press. Online at EFloras.org. [back]
- "Alnus rubra". in Flora of North America Vol. 3. Oxford University Press. Online at EFloras.org. [back]
- Mean = 363.430 meters (1,192.356 feet), Standard Deviation = 446.350 based on 225 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]