or on rock, occasionally hemiepiphytic
or epiphytic. Stems creeping
, rarely arborescent
, sometimes climbing
, branched or unbranched, dictyostelic, bearing scales
. Leaves circinate
in bud, monomorphic
usually not articulate
to stem, scales usually persistent
, in cross
with 2--many roundish bundles, or bundles 2 and lunate
to commonly 1--5-pinnate or more divided
, leaf buds
absent or present. Veins pinnate or parallel in ultimate
segments, simple or forked
or anastomosing, areoles sometimes with included
free veinlets. Indument
on blade commonly of glands
, and/or scales, especially on rachis and costae abaxially. Sori borne abaxially on veins or at vein
(but usually not marginal
), or sporangia acrostichoid
and covering abaxial
surface, if in discrete sori then variously shaped (round
, or elongate
) ; receptacle not or only slightly elevated
, with or without indusium, indusium variously linear
, or reniform
, sometimes hoodlike, cuplike, or round. Sporangia with stalk
of 2--3 rows
; annulus vertical
by stalk. Spores all of 1 kind, usually not green (except Matteuccia, Onoclea ), oblong or reniform in outline, monolete, variously ornamented (often broadly winged
), 64 per sporangium (32 in apogamous spp.
) . Gametophytes green, aboveground, cordate, glabrous
or often bearing glands or hairs; archegonia and antheridia borne on lower surface, antheridia 3-celled.
Genera ca. 60, species perhaps exceeding 3000 (18 genera, 79 species in the flora ) : worldwide.
The family Dryopteridaceae has been variously circumscribed; it is here delimited in a manner similar to that of R. M. Tryon and A. F. Tryon (1982) but with the inclusion of Nephrolepis . In many works, the family has gone under the illegitimate name Aspidiaceae. Some authorities define Dryopteridaceae more narrowly, to exclude Athyrium, Deparia, Diplazium, Cystopteris, and Gymnocarpium (Athyriaceae or Woodsiaceae), Woodsia (Woodsiaceae), Lomariopsis (Lomariopsidaceae), Nephrolepis (Nephrolepidaceae or Davalliaceae), Onoclea and Matteuccia (Onocleaceae), and Ctenitis and Tectaria (Tectariaceae) . Characteristics holding Dryopteridaceae (as circumscribed here) together include the bilateral , monolete spores, often broadly winged perispore, absence of needlelike hairs, scaly stem and petiole bases, abaxial (nonmarginal) sori, base chromosome number of 40 or 41 (also 38 and 39 in Woodsia, 37 in Onoclea, 42 in Cystopteris ), and usually indusiate sori. Loss of indusium, dimorphism , areolate venation , and reduced blade dissection have occurred repeatedly along many evolutionary lines in Dryopteridaceae, and in general these characteristics are often not very useful in delimiting genera or assessing intergeneric relationships .
In some genera, especially Phanerophlebia and Polystichum, the blade bears very narrow scales (sometimes called microscales) that resemble uniseriate hairs. These scales may be only one or two cells wide. Every intergradation exists between these filiform microscales and more typical, wider scales, and the two types are the same color, generally tan to brownish. Microscales are probably not homologous with true hairs, which may be either unicellular or multicellular , uncolored or sometimes reddish (as in Tectaria and Ctenitis ), glandular (as in Woodsia ) or not. Hairs in Dryopteridaceae, if present at all, are generally readily distinguishable from the needlelike, transparent ones found in Thelypteridaceae.
List of Habitats
- 1 Forest
- 1.6 Forest - Subtropical/Tropical Moist Lowland [more info]
- Whittaker & Margulis,1978
- Haeckel, 1866
- Cavalier-Smith, 1981
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Infraphylum: Moniliformopses () - Kenrick & Crane, 1997, Nom. Nud.
- Subphylum: Euphyllophytina ()
- Phylum: Polypodiophyta () - Sinnott, 1935 ex Cavalier-Smith, 1998 - Ferns
- Subkingdom: Viridaeplantae () - Cavalier-Smith, 1981
- Kingdom: Plantae () - Haeckel, 1866 - Plants
Arachniodes hasseltii • Dryopteris hasseltii
: Polypodium hasseltii Blume
Basionym author: (Blume)
Members of the genus Acrorumohra
ZipcodeZoo has pages for 0 species, subspecies, varieties, forms, and cultivars in this genus:
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- IUCN. 2004. 2004 IUCN Red List of Threatened Species. www.iucnredlist.org. Downloaded on 23 November 2004.
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 16, 2012.
- China Plant Specialist Group 2004. Acrorumohra hasseltii. In: IUCN 2011. IUCN Red List of Threatened Species. Version 2011.2. <www.iucnredlist.org>. Downloadedon 29January2012.
- IUCN 2012. IUCN Red List of Threatened Species. Version 2011.2. . Downloaded on January 28, 2012.
Accessed through GBIF Data Portal February 29, 2008:
- Australian National Herbarium (CANB)
- Taiwan Biodiversity Information Facility: Magnoliophyta
- Biodiversity Heritage Library NamebankID: 5984189
- Globally Unique Identifier: urn:lsid:ipni.org:names:17425690-1
- International Plant Names Index (IPNI) ID: 17425690
- IUCN ID: 188785
- Zipcode Zoo Species Identifier: 1029499