Abronia villosa

• Overview
• Interesting Facts
• Common Names
• Description
• Biology
• Taxonomy
• Similar Species
• More Info
• Further Reading
• Notes

Overview

Interesting Facts

Common Names

Description

Family Nyctaginaceae

Herbs, shrubs , trees , or sometimes spiny vines . Leaves opposite, alternate, or whorled ; stipules absent; petiole usually present, well defined; leaf blade simple , herbaceous or slightly fleshy , margin entire. Inflorescences mostly terminal , less often axillary , of cymes, umbels, or verticils , sometimes 1-flowered or fasciculate, often grouped into panicles; bracts often inconspicuous, sometimes forming calyxlike involucre, or large and brightly colored . Flowers bisexual , rarely unisexual or polygamous, actinomorphic . Perianth constricted beyond the ovary, base persistent , closely enclosing ovary which appears inferior, limb petaloid beyond constriction, tubular , funnelform , or campanulate , apex 5-10-lobed, lobes plicate or valvate in bud, persistent or caducous . Disk absent. Stamens (1-) 3-5(-many), hypogynous, free or connate at base, involute in bud; anthers 2-loculed, dehiscence longitudinal . Ovary superior, 1-loculed; ovule 1. Style 1; stigma globose . Fruit an achenelike anthocarp enclosed by persistent perianth, ribbed or winged , often glandular . Seed 1; endosperm present; embryo straight or curved .

About 30 genera and 300 species: tropics and subtropics, mainly in tropical America; six genera (two introduced ) and 13 species (one endemic, three introduced) in China.[1]

Genus Abronia

Herbs, annual or perennial , sometimes cespitose or appearing acaulescent , usually viscid-pubescent, from slender to stout taproot (extensively rhizomatous in Abronia bolackii). Stems prostrate to erect , unarmed , without glutinous bands on internodes. Leaves usually basal and cauline (all basal in A. bigelovii and A. nana), unequal in each pair, petiolate ; blade ± thick and succulent, base usually asymmetric . Inflorescences axillary (appearing scapose in A. bigelovii and A. nana), pedunculate , capitate clusters , with peripheral flowers usually opening first; receptacle slightly rounded to conic, without pedicel-like projections; bracts persistent , not accrescent , 5-10, distinct , forming involucre, lanceolate to broadly ovate , thinly papery or scarious , translucent , occasionally thin and green. Flowers bisexual , chasmogamous ; perianth radially symmetric , funnelform or salverform , constricted distal to ovary, abruptly expanded to 5-lobed limb; stamens 5-9, included ; styles included; stigmas linear . Fruits winged or not, usually fusiform or turbinate , in profile rhombic , cordate, or obdeltate, coriaceous , glabrate to viscid puberulent ; wings 2-5, opaque , subtly veined, not or only slightly extending beyond apex or base of body, distal margins sometimes dilated and flattened, broader than lamina, interior hollow, forming cavity , or filled with spongy tissue ; sulci smooth or slightly rugose .

Species ca. 20: North America, Mexico.

Mature to near-mature fruits are usually required for identification of Abronia species because of the variation of vegetative structures within each taxon . Abronia appears to be in a state of active evolution. Cross-pollination readily occurs in the greenhouse, producing a variety of hybrids. Hybridization occasionally occurs in the field .[2]

Physical Description

Biology

Reproduction

Duration: Annual

Growth

Culture: Space 18-24" apart.

Soil: Minimum pH: 6.1 • Maximum pH: 7.8

Sunlight: Sun Exposure: Full Sun .

Temperature: Cold Hardiness: 8a, 8b, 9a, 9b, 10a, 10b. (map)

Taxonomy

Synonyms

Abronia umbellata Lam. f. villosa (S.Watson) Voss • Abronia Umbellata Villosa

Notes

Publishing author : S.Watson Publication : Amer. Naturalist 7: 302 1873

Similar Species

Members of the genus Abronia

ZipcodeZoo has pages for 40 species, subspecies, varieties, forms, and cultivars in this genus:

A. alba (Whiteflower Sand Verbena) · A. alpina (Ramshaw Meadow Abronia) · A. ameliae (Amelia Sand Verbena) · A. ammophila (Tweedy's Sand Verbena) · A. angustifolia (Narrow-Leaf Sandverbena) · A. argillosa (Clay Sand Verbena) · A. aurita (Cope's Arboreal Alligator Lizard) · A. bigelovii (Galisteo Sand Verbena) · A. bolackii (Bolacks Sand Verbena) · A. carletoni (Carleton's Sand Verbena) · A. carletonii (Carleton's Sand Verbena) · A. elliptica (Dwarf Sandverbena) · A. fragrans (Four Oclock) · A. gracilis (Narrow Sandverbena) · A. insularis (Island Sand Verbena) · A. latifolia (Coastal Sand Verbena) · A. macrocarpa (Large-Fruited Sand Verbena) · A. maritima (Red Sand Verbena) · A. mellifera (Honey-Scent Sand Verbena) · A. minor (Little Sand Verbena) · A. nana (Dwarf Sand Verbena) · A. nana covillei (Coville's Dwarf Abronia) · A. nana S.Watson var. harrisii Welsh (Harris' Dwarf Sand Verbena) · A. nana S.Watson var. nana S.Watson (Dwarf Sand Verbena) · A. nana var. covillei (Coville's Dwarf Sand Verbena) · A. nana var. harrisii (Harris' Dwarf Sand Verbena) · A. nana var. nana (Dwarf Sand Verbena) · A. neurophylla (Beach Sand Verbena) · A. platyphylla (Broadleaf Sand Verbena) · A. pogonantha (Mojave Sand Verbena) · A. turbinata (Trans Montane Abronia) · A. umbellata (Pink Sand Verbena) · A. umbellata breviflora (Pink Sand Verbena) · A. umbellata variabilis (Beach Sandverbena) · A. umbellata var. umbellata (Sand-Verbena) · A. umbellata subsp. breviflora (Pink Sand Verbena) · A. umbellata subsp. variabilis (Variable Pink Sand Verbena) · A. villosa (Desert Sand Verbena) · A. villosa var. aurita (Sand Verbena) · A. villosa var. villosa (Wooly Desert Sand Verbena)

More Info

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Further Reading

• A flora of California, by Willis Linn Jepson. San Francisco, Calif., Cunningham, Curtis & Welch, 1909- url p. 455, p. 455.
• Abstract of the proceedings of the Linnaean Society of New York. [New York]: The Society, [1889-1932]. url p. 62.
• An illustrated flora of the Pacific States: Washington, Oregon, and California. Stanford University, Stanford University Press, 1923-[60] url p. 108, p. 109, p. 110.
• Annual record of science and industry. New York: Harper & Brothers, 1872-[1879] url p. 377.
• Botany. Cambridge, Mass.Welch, Bigelow, University Press, 1876-80. url p. 4.
• Britton, N. L. (ed.). North American flora. 21 1918 [New York]New York Botanical Garden. url p. 243, p. 246.
• Bulletin of the Torrey Botanical Club. 6 1875-1879 New York: Torrey Botanical Club, 1870-1996 url p. 313, p. 682, p. 686.
• Carnegie Institution of Washington publication. Washington, Carnegie Institution of Washington, 1902- url p. 108.
• Catalog of hymenoptera in America north of Mexico / prepared cooperatively by specialists on the various groups of Hymenoptera under the direction of Karl V. Krombein. .. [et al.]. Washington: Smithsonian Institution Press, 1979- url , , p. 1934, p. 1986.
• Contributions from the New York Botanical Garden. New York: The Garden, 1899- url p. 52, p. 682, p. 686.
• Contributions from the United States National Herbarium 4 1893 Washington, D.C.: Smithsonian Institution Press, 1890- url p. 178, p. 183, p. 315, p. 316, p. 32, p. 395, p. 396, p. 40, p. 45, p. V.
• Field book of western wild flowers, by Margaret Armstrong in collaboration with J.J. Thornber. .. with five hundred illustrations in black and white, and forty-eight plates in color drawn from nature by the author. 1915 London, C. [sic] P. Putnam's Sons, 1915. url index, page 581, p. 581.
• Flora of the Rocky Mountains and adjacent plains, Colorado, Utah, Wyoming, Idaho, Montana, Saskatchewan, Alberta, and neighboring parts of Nebraska, South Dakota, North Dakota, and British Columbia / by P.A. Rydberg. New York: The author, 1922. url p. 256.
• Flora of the Yellowstone National Park / by Frank Tweedy. Washington, D.C.: Published by the author, 1886. url p. 59, p. 59.
• Journal of the New York Botanical Garden. 33 1932 Lancaster, Pa.: Published for the Garden by the New Era Printing Co., 1900- url p. 249, p. 283, p. 94.
• Key to the Rocky Mountain flora. Colorado, Utah, Wyoming, Idaho, Montana, Saskatchewan, Alberta, and parts of Nebraska, South Dakota, North Dakota, and British Columbia. New York, [c1919] url .
• Key to the Rocky Mountain flora; Colorado, Utah, Wyoming, Idaho, Montana, Saskatchewan, Alberta, and parts of Nebraska, South Dakota, North Dakota, and British Columbia. New York, 1919. url p. 79.
• Muhlenbergia. Chico, Calif. [etc] url p. 139.
• Occasional papers - San Diego Society of Natural History. San Diego, The Society. url p. 47.
• Phytologia memoirs. Plainfield, N.J.: H.N. Moldenke and A.L. Moldenke, 1980- url p. 201.
• Pittonia: a series of papers relating to botany and botanists /by Edward L. Greene. 1 1887-1889 Berkeley, Calif. [etc.]: Doxey & Co. [etc.], 1887-1905. url p. 81.
• Report upon United States Geographical surveys west of the one hundredth meridian, Washington, Govt. print. off., 1875-89. url , , .
• Standardized plant names; a catalogue of approved scientific and common names of plants in American commerce. Salem, Mass., 1923. url p. 460.
• Studies on Rocky Mountain flora. New York, New York Botanical Garden, 1899-1913. url p. 686.
• The American botanist: a monthly journal for the plant lover. Binghamton, N.Y.: Willard N. Clute & Co., 1901-1948. url p. 49.
• The English flower garden and home grounds; design and arrangement shown by existing examples of gardens in Great Britain and Ireland, followed by a description of the plants, shrubs and trees for the open-air garden and their culture, by W. Robinson. .. Illustrated with many engravings on wood. London: J. Murray, 1900. url p. 407.
• The Garden: an illustrated weekly journal of gardening in all its branches. London: [s.n., url p. 116.
• The Gardeners' chronicle: a weekly illustrated journal of horticulture and allied subjects. London: [Gardeners Chronicle], 1874-1955. url p. 239, p. 267, p. 300.
• The Great Basin naturalist. 35 1975 Provo, Utah: M.L. Bean Life Science Museum, Brigham Young University, 1939-1999. url p. 116, p. 3.
• The IUCN Mammal Red Data Book. Part 1: threatened mammalian taxa of the Americas and the Australasian zoogeographic region (excluding Cetacea) IUCN url p. 501, p. 501.
• The Salton Sea; a study of the geography, the geology, the floristics, and the ecology of a desert basin, Washington, D.C.Carnegie Institution of Washington1914 url p. 108.
• Torreya. Burlington, Vt., Torrey Botanical Club, 1901-1945. url p. 161, p. 58.
• University of California publications in zoology. Berkeley: University Press, 1906- url p. 78.
• Western flower guide, wild flowers of the Rockies and west to the Pacific, Garden City, Doubleday, Page & company, 1917. url p. 271, p. 46.
• Zoe:a biological journal. 4 1893 San Diego, Calif. [etc.]Zoe Publishing Co. url p. 158, p. 158, p. 68, p. 68.
• Lu Dequan. 1996. Nyctaginaceae. In: Tang Changlin, ed., Fl. Reipubl. Popularis Sin. 26: 114.
• Galloway, L. A. 1975. Systematics of North American desert species of Abronia and Tripterocalyx. Brittonia 27: 328-347.
• Tillett, S. S. 1967. The maritime species of Abronia (Nyctaginaceae). Brittonia 19: 299-327.

Notes

Contributors

• "Abronia villosa". in Flora of North America Vol. 4 Page 62, 66, 67, 6. Published by Oxford University Press. Online at EFloras.org.
• Bisby, F.A., Y.R. Roskov, M.A. Ruggiero, T.M. Orrell, L.E. Paglinawan, P.W. Brewer, N. Bailly, J. van Hertum, eds (2007). Species 2000 and ITIS Catalogue of Life: 2007 Annual Checklist. Species 2000: Reading, U.K.
• Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 9, 2012.
• Global Biodiversity Information Facility. Accessed February 28, 2008. http://www.gbif.org Mediated distribution data from 5 providers.
• Ruggiero M., Gordon D., Bailly N., Kirk P., Nicolson D. (2011). The Catalogue of Life Taxonomic Classification, Edition 2, Part A. In: Species 2000 and ITIS Catalogue of Life: 2011 Annual Checklist (Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D., eds). DVD; Species 2000: Reading, UK.
• The International Plant Names Index. Accessed Dec 27, 2011.
• USDA, ARS, National Genetic Resources Program. Germplasm Resources Information Network - (GRIN) [Online Database]. National Germplasm Resources Laboratory, Beltsville, Maryland. URL (May 01, 2008)
• USDA, NRCS. 2005. The PLANTS Database, Version 3.5 (http://plants.usda.gov). National Plant Data Center, Baton Rouge, LA 70874-4490 USA.

Data Sources

Accessed through GBIF Data Portal February 28, 2008:

• Arizona State University, International Institute for Species Exploration: Arizona State University Vascular Plant Herbarium
• Arizona State University, International Institute for Species Exploration: The Deaver Herbarium, Northern Arizona University
• Berkeley Natural History Museums: University and Jepson Herbaria DiGIR provider
• Missouri Botanical Garden
• USDA PLANTS: USDA PLANTS Database
• Utah State University: USU-UTC Specimen Database

Identifiers

• Biodiversity Heritage Library NamebankID: 2646078
• Catalogue of Life Accepted Name Code: ITS-19577
• Globally Unique Identifier: urn:lsid:ipni.org:names:604025-1
• GRIN Nomen Number: 316869
• Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 19577
• International Plant Names Index (IPNI) ID: 122-2
• Natural Heritage Network Species Identifier: PDNYC010P0
• U.S.D.A. Plant Symbol: ABVI
• Zipcode Zoo Species Identifier: 69181

Footnotes

1. Dequan Lu & Michael G. Gilbert "Nyctaginaceae". in Flora of China Vol. 5 Page 430. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
2. Leo A. Galloway "Abronia". in Flora of North America Vol. 4 Page 16, 61. Oxford University Press. Online at EFloras.org. [back]

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