Interesting Facts
Common Names
Common Names in English:
Trans Montane Abronia, Transmontane Sand Verbena, Transmontane Sandverbena
Description
Family Nyctaginaceae
Herbs, shrubs
, trees
, or sometimes spiny
vines
. Leaves opposite, alternate, or whorled
; stipules absent; petiole
usually present, well defined; leaf blade
simple
, herbaceous or slightly fleshy
, margin
entire. Inflorescences mostly terminal
, less often axillary
, of cymes, umbels, or verticils
, sometimes 1-flowered or fasciculate, often grouped into panicles; bracts often inconspicuous, sometimes forming calyxlike involucre, or large and brightly colored
. Flowers bisexual
, rarely unisexual
or polygamous, actinomorphic
. Perianth constricted
beyond the ovary, base
persistent
, closely enclosing ovary which appears inferior, limb petaloid
beyond constriction, tubular
, funnelform
, or campanulate
, apex 5-10-lobed, lobes
plicate
or valvate
in bud, persistent or caducous
. Disk absent. Stamens (1-) 3-5(-many), hypogynous, free
or connate
at base, involute
in bud; anthers
2-loculed, dehiscence longitudinal
. Ovary superior, 1-loculed; ovule 1. Style 1; stigma globose
. Fruit an achenelike anthocarp enclosed by persistent perianth, ribbed
or winged
, often glandular
. Seed 1; endosperm present; embryo straight or curved
.
About 30 genera and 300 species: tropics and subtropics, mainly in tropical
America; six genera (two introduced
) and 13 species (one endemic, three introduced) in China.[1]
Genus Abronia
Herbs, annual
or perennial
, sometimes cespitose or appearing acaulescent
, usually viscid-pubescent, from slender to stout taproot
(extensively rhizomatous
in Abronia bolackii). Stems prostrate
to erect
, unarmed
, without glutinous
bands
on internodes. Leaves usually basal and cauline (all basal in A. bigelovii and A. nana), unequal in each pair, petiolate
; blade
± thick and succulent, base
usually asymmetric
. Inflorescences axillary
(appearing scapose
in A. bigelovii and A. nana), pedunculate
, capitate clusters
, with peripheral flowers usually opening first; receptacle slightly rounded
to conic, without pedicel-like projections; bracts persistent
, not accrescent
, 5-10, distinct
, forming involucre, lanceolate to broadly ovate
, thinly papery
or scarious
, translucent
, occasionally thin and green. Flowers bisexual
, chasmogamous
; perianth radially symmetric
, funnelform
or salverform
, constricted
distal to ovary, abruptly expanded to 5-lobed limb; stamens 5-9, included
; styles included; stigmas linear
. Fruits winged
or not, usually fusiform
or turbinate
, in profile
rhombic
, cordate, or obdeltate, coriaceous
, glabrate
to viscid
puberulent
; wings
2-5, opaque
, subtly veined, not or only slightly extending beyond apex or base of body, distal margins sometimes dilated
and flattened, broader than lamina, interior hollow, forming cavity
, or filled with spongy
tissue
; sulci smooth
or slightly rugose
.
Species ca.
20: North America, Mexico.
Mature
to near-mature fruits are usually required for identification of Abronia species because of the variation
of vegetative
structures within each taxon
. Abronia appears to be in a state of active
evolution. Cross-pollination
readily occurs in the greenhouse, producing a variety of hybrids. Hybridization occasionally occurs in the field
.[2]
Physical Description
Species Abronia turbinata
Plants annual , infrequently perennial . Stems decumbent to ascending , much branched, elongate , reddish at least basally, glandular-pubescent , rarely glabrous or viscid-pubescent. Leaves: petiole 1-4.5 cm; blade broadly ovate to orbiculate, 1-5 × 0.5-3 cm, margins entire or ± repand and undulate , surfaces glabrous or sparsely glandular-pubescent. Inflorescences: peduncle longer than subtending petiole; bracts lanceolate to ovate, 3-10 × 1-5 mm, papery , puberulent to densely glandular-pubescent; flowers 15-35. Perianth: tube greenish to coral pink, 6-18 mm, limb white to pale pink, 5-8 mm diam. Fruits winged , turbinate , 3-8 × 3-6 mm, coriaceous , apex broadly tapered to prominent beak ; wings (2-) 5 (when 2, folded together) truncate distally with conspicuous dilations, cavities extending throughout. [source]
Habit: Forb/herb
Flowers: Bloom Period: February, March, April, May, June, July, August, September, October. • Flower Color: near white, pink, white
Size/Age/Growth
Size: under 6" tall.
Habitat
Sandy soils, desert scrub ; 900-2500 m [3].
Biology
Reproduction
Duration: Annual , Perennial
Growth
Culture: Space 6-9" apart.
Soil: Minimum pH: 6.1 • Maximum pH: 7.5
Sunlight: Sun Exposure: Full Sun .
Moisture: Drought Tolerance: High
Temperature: Cold Hardiness: 8a, 8b, 9a, 9b, 10a, 10b. (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Caryophyllidae
(
)
- Takhtajan, 1967
- Superorder:
Caryophyllanae
(
)
- Takhtajan, 1967
- Order:
Caryophyllales
(
)
- Perleb, 1826
- Suborder:
Phytolaccineae
(
)
- Family:
Nyctaginaceae
(
)
- A.L. de Jussieu, 1789, nom. cons.
- four o'clocks, nyctaginacées
- Family:
Nyctaginaceae
(
- Suborder:
Phytolaccineae
(
- Order:
Caryophyllales
(
- Superorder:
Caryophyllanae
(
- Subclass:
Caryophyllidae
(
- Class:
Spermatopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Synonyms
Abronia exalata Standl. • Abronia orbiculata Standl.
Notes
Publishing author : Torr. ex S.Watson Publication : Botany [Fortieth Parallel] 285 (t. 31) 1871 [Sep-Dec 1871]
Similar Species
Members of the genus Abronia
ZipcodeZoo has pages for 40 species, subspecies, varieties, forms, and cultivars in this genus:
A. alba (Whiteflower Sand Verbena) · A. alpina (Ramshaw Meadow Abronia) · A. ameliae (Amelia Sand Verbena) · A. ammophila (Tweedy's Sand Verbena) · A. angustifolia (Narrow-Leaf Sandverbena) · A. argillosa (Clay Sand Verbena) · A. aurita (Cope's Arboreal Alligator Lizard) · A. bigelovii (Galisteo Sand Verbena) · A. bolackii (Bolacks Sand Verbena) · A. carletoni (Carleton's Sand Verbena) · A. carletonii (Carleton's Sand Verbena) · A. elliptica (Dwarf Sandverbena) · A. fragrans (Four Oclock) · A. gracilis (Narrow Sandverbena) · A. insularis (Island Sand Verbena) · A. latifolia (Coastal Sand Verbena) · A. macrocarpa (Large-Fruited Sand Verbena) · A. maritima (Red Sand Verbena) · A. mellifera (Honey-Scent Sand Verbena) · A. minor (Little Sand Verbena) · A. nana (Dwarf Sand Verbena) · A. nana covillei (Coville's Dwarf Abronia) · A. nana S.Watson var. harrisii Welsh (Harris' Dwarf Sand Verbena) · A. nana S.Watson var. nana S.Watson (Dwarf Sand Verbena) · A. nana var. covillei (Coville's Dwarf Sand Verbena) · A. nana var. harrisii (Harris' Dwarf Sand Verbena) · A. nana var. nana (Dwarf Sand Verbena) · A. neurophylla (Beach Sand Verbena) · A. platyphylla (Broadleaf Sand Verbena) · A. pogonantha (Mojave Sand Verbena) · A. turbinata (Trans Montane Abronia) · A. umbellata (Pink Sand Verbena) · A. umbellata breviflora (Pink Sand Verbena) · A. umbellata variabilis (Beach Sandverbena) · A. umbellata var. umbellata (Sand-Verbena) · A. umbellata subsp. breviflora (Pink Sand Verbena) · A. umbellata subsp. variabilis (Variable Pink Sand Verbena) · A. villosa (Desert Sand Verbena) · A. villosa var. aurita (Sand Verbena) · A. villosa var. villosa (Wooly Desert Sand Verbena)
More Info
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Further Reading
- A flora of California, by Willis Linn Jepson. San Francisco, Calif., Cunningham, Curtis & Welch, 1909- url p. 456.
- American plants. .. [Descriptions, bibliographical notes, synonymy, and other information, comp. from many sources] Charles Russell Orcutt, editor. San Diego, Calif., [1907]-1910. url p. 279.
- An illustrated flora of the Pacific States: Washington, Oregon, and California. Stanford University, Stanford University Press, 1923-[60] url p. 108, p. 111, p. 112.
- Biologia centrali-americana; or, Contributions to the knowledge of the fauna and flora of Mexico and Central America. London, Pub. for the editors by R. H. Porter and Dulau & co., 1879-88. url .
- Botany. Washington, Govt. Print. Off., 1871. url , .
- Britton, N. L. (ed.). North American flora. 21 1918 [New York]New York Botanical Garden. url p. 253, p. 254.
- Bulletin of the Southern California Academy of Sciences Los Angeles, Calif.: The Academy, 1971- url p. 77.
- Bulletin of the Torrey Botanical Club. 16 1889 New York: Torrey Botanical Club, 1870-1996 url p. 174, p. 684.
- Contributions from the United States National Herbarium 4 1893 Washington, D.C.: Smithsonian Institution Press, 1890- url p. 178, p. 183, p. 318, p. 327, p. 4, p. 40, p. V, p. XIII.
- Entomological news, and proceedings of the Entomological Section of the Academy of Natural Sciences of Philadelphia. Philadelphia[Entomological Rooms of the Academy of Natural Sciences] url p. 39.
- Muhlenbergia. Chico, Calif. [etc] url p. 194, p. 91.
- National list of scientific plant names. [Washington, D.C.]: U.S. Dept. of Agriculture, Soil Conservation Service, 1982- url p. 5.
- Phytologia. Bronx Park, New York, H.A. Gleason and H.N. Moldenke, url p. 42, p. 43.
- Proceedings of the Academy of Natural Sciences of Philadelphia. Philadelphia, Academy of Natural Sciences of Philadelphia url p. 167.
- Proceedings of the California Academy of Sciences. San Francisco, The Academy. url p. 313.
- Report of the geological exploration of the fortieth parallel / Washington, D.C.: G.P.O., 1870-80 url , p. 285.
- Science. New York, N.Y.: [s.n.]1880- url p. 626.
- The Great Basin naturalist. 43 1983 Provo, Utah: M.L. Bean Life Science Museum, Brigham Young University, 1939-1999. url p. 117.
- Watson, S. Botany /by Sereno Watson, aided by Daniel C. Eaton, and others. 5 1871 Washington, D.C.: G.P.O., 1871. url p. 285.
- Wild flowers of Canada published exclusively with the Montreal Star, Montreal; by special artists and botanists. Montreal: Montreal Star, [1895?] url .
- Zoe:a biological journal. 4 1893 San Diego, Calif. [etc.]Zoe Publishing Co. url p. 10, p. 10, p. 125, p. 233.
- Lu Dequan. 1996. Nyctaginaceae. In: Tang Changlin, ed., Fl. Reipubl. Popularis Sin. 26: 114.
- Galloway, L. A. 1975. Systematics of North American desert species of Abronia and Tripterocalyx. Brittonia 27: 328-347.
- Tillett, S. S. 1967. The maritime species of Abronia (Nyctaginaceae). Brittonia 19: 299-327.
Notes
Contributors
- "Abronia turbinata". in Flora of North America Vol. 4 Page 62, 64, 66. Published by Oxford University Press. Online at EFloras.org.
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 9, 2012.
- Global Biodiversity Information Facility. Accessed February 28, 2008. http://www.gbif.org Mediated distribution data from 5 providers.
- Ruggiero M., Gordon D., Bailly N., Kirk P., Nicolson D. (2011). The Catalogue of Life Taxonomic Classification, Edition 2, Part A. In: Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist (Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D., eds). DVD; Species 2000: Reading, UK.
- The International Plant Names Index. Accessed Dec 27, 2011.
- USDA, ARS, National Genetic Resources Program. Germplasm Resources Information Network - (GRIN) [Online Database]. National Germplasm Resources Laboratory, Beltsville, Maryland. URL (May 01, 2008)
- USDA, NRCS. 2005. The PLANTS Database, Version 3.5 (http://plants.usda.gov). National Plant Data Center, Baton Rouge, LA 70874-4490 USA.
Data Sources
Accessed through GBIF Data Portal February 28, 2008:
- Berkeley Natural History Museums: University and Jepson Herbaria DiGIR provider
- Burke Museum: Vascular Plant Collection - University of Washington Herbarium (WTU)
- Oregon State University: Vascular Plant Collection
- USDA PLANTS: USDA PLANTS Database
- Utah State University: USU-UTC Specimen Database
Identifiers
- Biodiversity Heritage Library NamebankID: 2646076
- Catalogue of Life Accepted Name Code: ITS-19571
- Globally Unique Identifier: urn:lsid:ipni.org:names:604021-1
- GRIN Nomen Number: 448065
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 19571
- International Plant Names Index (IPNI) ID: 326892-2
- Natural Heritage Network Species Identifier: PDNYC010M0
- U.S.D.A. Plant Symbol: ABTU
- Zipcode Zoo Species Identifier: 69702
Footnotes
- Dequan Lu & Michael G. Gilbert "Nyctaginaceae". in Flora of China Vol. 5 Page 430. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Leo A. Galloway "Abronia". in Flora of North America Vol. 4 Page 16, 61. Oxford University Press. Online at EFloras.org. [back]
- "Abronia turbinata". in Flora of North America Vol. 4 Page 62, 64, 66. Oxford University Press. Online at EFloras.org. [back]
