Overview
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Threatened |
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Interesting Facts
Common Names
Common Names in English:
Beach Sand Verbena
Description
Family Nyctaginaceae
Herbs, shrubs
, trees
, or sometimes spiny
vines
. Leaves opposite, alternate, or whorled
; stipules absent; petiole
usually present, well defined; leaf blade
simple
, herbaceous or slightly fleshy
, margin
entire. Inflorescences mostly terminal
, less often axillary
, of cymes, umbels, or verticils
, sometimes 1-flowered or fasciculate, often grouped into panicles; bracts often inconspicuous, sometimes forming calyxlike involucre, or large and brightly colored
. Flowers bisexual
, rarely unisexual
or polygamous, actinomorphic
. Perianth constricted
beyond the ovary, base
persistent
, closely enclosing ovary which appears inferior, limb petaloid
beyond constriction, tubular
, funnelform
, or campanulate
, apex 5-10-lobed, lobes
plicate
or valvate
in bud, persistent or caducous
. Disk absent. Stamens (1-) 3-5(-many), hypogynous, free
or connate
at base, involute
in bud; anthers
2-loculed, dehiscence longitudinal
. Ovary superior, 1-loculed; ovule 1. Style 1; stigma globose
. Fruit an achenelike anthocarp enclosed by persistent perianth, ribbed
or winged
, often glandular
. Seed 1; endosperm present; embryo straight or curved
.
About 30 genera and 300 species: tropics and subtropics, mainly in tropical
America; six genera (two introduced
) and 13 species (one endemic, three introduced) in China.[1]
Genus Abronia
Herbs, annual
or perennial
, sometimes cespitose or appearing acaulescent
, usually viscid-pubescent, from slender to stout taproot
(extensively rhizomatous
in Abronia bolackii). Stems prostrate
to erect
, unarmed
, without glutinous
bands
on internodes. Leaves usually basal and cauline (all basal in A. bigelovii and A. nana), unequal in each pair, petiolate
; blade
± thick and succulent, base
usually asymmetric
. Inflorescences axillary
(appearing scapose
in A. bigelovii and A. nana), pedunculate
, capitate clusters
, with peripheral flowers usually opening first; receptacle slightly rounded
to conic, without pedicel-like projections; bracts persistent
, not accrescent
, 5-10, distinct
, forming involucre, lanceolate to broadly ovate
, thinly papery
or scarious
, translucent
, occasionally thin and green. Flowers bisexual
, chasmogamous
; perianth radially symmetric
, funnelform
or salverform
, constricted
distal to ovary, abruptly expanded to 5-lobed limb; stamens 5-9, included
; styles included; stigmas linear
. Fruits winged
or not, usually fusiform
or turbinate
, in profile
rhombic
, cordate, or obdeltate, coriaceous
, glabrate
to viscid
puberulent
; wings
2-5, opaque
, subtly veined, not or only slightly extending beyond apex or base of body, distal margins sometimes dilated
and flattened, broader than lamina, interior hollow, forming cavity
, or filled with spongy
tissue
; sulci smooth
or slightly rugose
.
Species ca.
20: North America, Mexico.
Mature
to near-mature fruits are usually required for identification of Abronia species because of the variation
of vegetative
structures within each taxon
. Abronia appears to be in a state of active
evolution. Cross-pollination
readily occurs in the greenhouse, producing a variety of hybrids. Hybridization occasionally occurs in the field
.[2]
Physical Description
Biology
Reproduction
Duration: Perennial
Growth
Culture: Space 9-12" apart.
Soil: Minimum pH: 5.6 • Maximum pH: 6.5
Sunlight: Sun Exposure: Full Sun .
Moisture: Drought Tolerance: High
Temperature: Cold Hardiness: 7a, 7b, 8a, 8b, 9a, 9b. (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Caryophyllidae
(
)
- Takhtajan, 1967
- Superorder:
Caryophyllanae
(
)
- Takhtajan, 1967
- Order:
Caryophyllales
(
)
- Perleb, 1826
- Suborder:
Phytolaccineae
(
)
- Family:
Nyctaginaceae
(
)
- A.L. de Jussieu, 1789, nom. cons.
- four o'clocks, nyctaginacées
- Family:
Nyctaginaceae
(
- Suborder:
Phytolaccineae
(
- Order:
Caryophyllales
(
- Superorder:
Caryophyllanae
(
- Subclass:
Caryophyllidae
(
- Class:
Spermatopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Notes
Publishing author : Standl. Publication : Contr. U.S. Natl. Herb. xii. 314 (1909).
Similar Species
Members of the genus Abronia
ZipcodeZoo has pages for 40 species, subspecies, varieties, forms, and cultivars in this genus:
A. alba (Whiteflower Sand Verbena) · A. alpina (Ramshaw Meadow Abronia) · A. ameliae (Amelia Sand Verbena) · A. ammophila (Tweedy's Sand Verbena) · A. angustifolia (Narrow-Leaf Sandverbena) · A. argillosa (Clay Sand Verbena) · A. aurita (Cope's Arboreal Alligator Lizard) · A. bigelovii (Galisteo Sand Verbena) · A. bolackii (Bolacks Sand Verbena) · A. carletoni (Carleton's Sand Verbena) · A. carletonii (Carleton's Sand Verbena) · A. elliptica (Dwarf Sandverbena) · A. fragrans (Four Oclock) · A. gracilis (Narrow Sandverbena) · A. insularis (Island Sand Verbena) · A. latifolia (Coastal Sand Verbena) · A. macrocarpa (Large-Fruited Sand Verbena) · A. maritima (Red Sand Verbena) · A. mellifera (Honey-Scent Sand Verbena) · A. minor (Little Sand Verbena) · A. nana (Dwarf Sand Verbena) · A. nana covillei (Coville's Dwarf Abronia) · A. nana S.Watson var. harrisii Welsh (Harris' Dwarf Sand Verbena) · A. nana S.Watson var. nana S.Watson (Dwarf Sand Verbena) · A. nana var. covillei (Coville's Dwarf Sand Verbena) · A. nana var. harrisii (Harris' Dwarf Sand Verbena) · A. nana var. nana (Dwarf Sand Verbena) · A. neurophylla (Beach Sand Verbena) · A. platyphylla (Broadleaf Sand Verbena) · A. pogonantha (Mojave Sand Verbena) · A. turbinata (Trans Montane Abronia) · A. umbellata (Pink Sand Verbena) · A. umbellata breviflora (Pink Sand Verbena) · A. umbellata variabilis (Beach Sandverbena) · A. umbellata var. umbellata (Sand-Verbena) · A. umbellata subsp. breviflora (Pink Sand Verbena) · A. umbellata subsp. variabilis (Variable Pink Sand Verbena) · A. villosa (Desert Sand Verbena) · A. villosa var. aurita (Sand Verbena) · A. villosa var. villosa (Wooly Desert Sand Verbena)
More Info
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- Search for Scholarly Articles: Google Scholar
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- Search using Specialized Databases: GenBank | Medline | Scirus | CISTI/CAL | Agricola Periodicals | Agricola Books
Further Reading
- 1997 IUCN red list of threatened plants Cambridge: IUCN, World Conservation Union, 1998 url p. 435.
- An illustrated flora of the Pacific States: Washington, Oregon, and California. Stanford University, Stanford University Press, 1923-[60] url p. 110.
- Britton, N. L. (ed.). North American flora. 21 1918 [New York]New York Botanical Garden. url p. 247.
- Leaflets of western botany. San Fransisco:[J. T. Howell], 1932-1966. url p. 60.
- Lu Dequan. 1996. Nyctaginaceae. In: Tang Changlin, ed., Fl. Reipubl. Popularis Sin. 26: 114.
- Galloway, L. A. 1975. Systematics of North American desert species of Abronia and Tripterocalyx. Brittonia 27: 328-347.
- Tillett, S. S. 1967. The maritime species of Abronia (Nyctaginaceae). Brittonia 19: 299-327.
Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 9, 2012.
Identifiers
- Biodiversity Heritage Library NamebankID: 2646073
- Catalogue of Life Accepted Name Code: ITS-19568
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 19568
- International Plant Names Index (IPNI) ID: 95-2
- Natural Heritage Network Species Identifier: PDNYC010J0
- U.S.D.A. Plant Symbol: ABNE
- Zipcode Zoo Species Identifier: 68269
Footnotes
- Dequan Lu & Michael G. Gilbert "Nyctaginaceae". in Flora of China Vol. 5 Page 430. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Leo A. Galloway "Abronia". in Flora of North America Vol. 4 Page 16, 61. Oxford University Press. Online at EFloras.org. [back]
