Overview
Taxonomy
The Subfamily Paronychioideae is a member of the Family Caryophyllaceae. Here is the complete "parentage" of Paronychioideae:
- Domain: Eukaryota
Whittaker & Margulis,1978 - eukaryotes
- Kingdom: Plantae
Haeckel, 1866 - Plants
- Subkingdom: Viridaeplantae
Cavalier-Smith, 1981 - Green Plants
- Phylum: Tracheophyta
Sinnott, 1935 ex Cavalier-Smith, 1998 - Vascular Plants
- Subphylum: Euphyllophytina
- Infraphylum: Radiatopses
Kenrick & Crane, 1997
- Class: Magnoliopsida
Brongniart, 1843 - Dicotyledons
- Subclass: Caryophyllidae
Takhtajan, 1967
- Superorder: Caryophyllanae
Takhtajan, 1967
- Order: Caryophyllales
Perleb, 1826
- Suborder: Caryophyllineae
- Family: Caryophyllaceae
(kar-ree-oh-fil-AY-see-ee)
Durande, 1782 ex A.L. de Jussieu, 1789, nom. cons.
- Subfamily: Paronychioideae
- Family: Caryophyllaceae
(kar-ree-oh-fil-AY-see-ee)
Durande, 1782 ex A.L. de Jussieu, 1789, nom. cons.
- Suborder: Caryophyllineae
- Order: Caryophyllales
Perleb, 1826
- Superorder: Caryophyllanae
Takhtajan, 1967
- Subclass: Caryophyllidae
Takhtajan, 1967
- Class: Magnoliopsida
Brongniart, 1843 - Dicotyledons
- Infraphylum: Radiatopses
Kenrick & Crane, 1997
- Subphylum: Euphyllophytina
- Phylum: Tracheophyta
Sinnott, 1935 ex Cavalier-Smith, 1998 - Vascular Plants
- Subkingdom: Viridaeplantae
Cavalier-Smith, 1981 - Green Plants
- Kingdom: Plantae
Haeckel, 1866 - Plants
The Subfamily Paronychioideae is further organized into finer groupings including:
- Tribe (3): Corrigioleae · Paronychieae · Polycarpeae
- Genus (20): Cardionema · Cometes · Corrigiola · Delia · Dicheranthus · Drymaria · Herniaria · Illecebrum · Loeflingia · Ortegia · Paronychia · Polycarpaea · Polycarpon · Pteranthus · Spergula · Spergularia · Sphaerocoma · Stipulicida · Telephium · Xerotia
- Species: ZipcodeZoo has pages for 1,138 species, subspecies, varieties, forms, and cultivars in the Subfamily Paronychioideae.
Genera
Cardionema
Herbs, perennial. Taproots often stout. Stems prostrate, much-branched proximally, terete to angular. Leaves opposite, not connate, connected by thickened margins from which stipules arise, sessile; stipules 2 per node, silvery, lanceolate to ovate, margins entire to irregularly fimbriate, apex 2-lobed; blade prominently 1-veined, needlelike, not succulent, apex spinose. Inflorescences axillary, 1-3-flowered clusters; bracts paired, resembling stipules, smaller. Flowers sessile; hypanthium cup-shaped, not abruptly expanded distally; sepals 5, distinct, olive-green, oblong to obovate, 1.2-2.8 mm (excluding awns), indurate, herbaceous, margins whitish, scarious, apex spinose, prominently hooded, awned, hoods projecting inward and enclosing developing fruit, awns arising from near apex, widely divergent, very stout, spinose, 3 awns and associated sepals well developed, alternating with 2 reduced sepals; nectaries not apparent; stamens 3-5; filaments distinct to base; staminodes 5, adnate basally with alternating filaments, ovate-triangular, apex acuminate; styles 2, distinct, subcapitate, 0.2 mm, glabrous proximally; stigmas 2, subterminal, obscurely papillate (50×). Utricles ellipsoid to cylindric, teeth absent, indehiscent. Seeds tan with red spot at hilum, narrowly ovate, not compressed, smooth, marginal wing absent, appendage absent; embryo peripheral, straight to slightly curved.[1] [more]
Cometes
Corrigiola
Herbs, annual or biennial [perennial]. Taproots slender. Stems decumbent to ascending, branched, terete. Leaves alternate or rarely subopposite, sometimes forming basal rosette, not connate, petiolate or sessile; stipules 2 per node, white, ovate, margins mostly entire, apex acuminate; petiole minute (subopposite leaves) ; blade 1-veined, narrowly oblanceolate to spatulate, not succulent, apex obtuse. Inflorescences terminal or axillary, dense cymes; bracts paired, smaller, scarious. Pedicels erect. Flowers: perianth and androecium slightly perigynous; hypanthium cup-shaped, not abruptly expanded distally; sepals 5, distinct, green or reddish brown, 1-1.5 mm, herbaceous, margins white-scarious, apex blunt, not hooded, not awned; nectaries not apparent; stamens 5; filaments distinct; staminodes 5, arising from hypanthium rim, ovate to oblong; styles 3, distinct, capitate, 0.1-0.2 mm, glabrous proximally; stigmas 3, capitate, obscurely papillate (at 50×). Utricles at least partly enclosed by persistent calyx/hypanthium, obscurely 3-gonous, indehiscent. Seeds white with tan band, reniform or subglobose, not compressed, finely papillose, marginal wing absent, appendage absent; embryo peripheral, curved. x = 8, 9.[2] [more]
Delia
Delia can refer to: [more]
Dicheranthus
Drymaria
Herbs, annual or perennial, caudices often branched. Taproots slender, elongate. Stems sprawling to erect, simple or branching proximally or throughout, terete. Leaves opposite or appearing whorled, connate by membranous to thickened line, petiolate or sessile, stipulate (D. pachyphylla not stipulate) ; stipules 2 per node, white to tan, simple or divided into segments, subulate to filiform, often minute, margins entire, apex acute to acuminate; blade 1-5-veined, linear to lanceolate, spatulate, ovate, reniform, or orbiculate, not succulent, apex rounded to acuminate. Inflorescences terminal or axillary, open to congested, bracteate cymes or umbelliform clusters or flowers solitary, axillary; bracts paired, scarious or central portion herbaceous. Pedicels erect to spreading or reflexed. Flowers: perianth and androecium hypogynous; sepals 5, distinct, white, lanceolate to oblong, ovate, or orbiculate, 1.5-4.8(-5) mm, herbaceous, margins white to purple, scarious, apex acuminate to rounded, hooded or not; petals (3-) 5, sometimes absent, white, claw narrow, tapering distally or with oblong or expanded, sessile or short-clawed trunk, auricles absent, blade apex divided into 2 or 4 lobes; nectaries at base of filaments opposite sepals; stamens 5; filaments distinct or briefly connate proximally; styles 3, occasionally 2, connate proximally for 2 of length, rarely to nearly distinct (D. cordata), filiform, 0.1-0.3 mm, glabrous proximally; stigmas 3, occasionally 2, linear along adaxial surfaces of styles (or branches), obscurely papillate (30×). Capsules ellipsoid to globose, opening by (2-) 3 spreading to recurved valves; carpophore absent. Seeds 3-25, tan, reddish brown, dark brown, black, or transparent (white embryo visible), horseshoe-, snail-shell- or teardrop-shaped, compressed laterally, at least somewhat, tuberculate, marginal wing absent, appendage absent. x = (11), 12.[3] [more]
Herniaria
Herbs, annual, biennial, or perennial. Taproots slender. Stems ascending or spreading to often prostrate, much-branched from base, mat-forming, terete. Leaves opposite, or distalmost alternate (from reduction of 1 member of a pair), connate by a line of tissue between adjacent stipules, sessile or virtually so; stipules 2 per node, inconspicuous, white, ovate to deltate, margins ciliate, apex acute; blade 1-veined, oblanceolate to elliptic or suborbiculate, not succulent, apex acute to rounded. Inflorescences densely clustered cymes usually on short lateral branches opposite a leaf; bracts paired, resembling stipules, smaller. Pedicels: flowers sessile. Flowers: hypanthium cup-shaped, not abruptly expanded distally; sepals 5, distinct, greenish to whitish green, lanceolate to oblong, 0.5-1.5 mm, herbaceous, margins green, herbaceous, apex acute to subobtuse, not hooded, not awned; nectaries near inner surface of filament bases; stamens (2-) 4-5; filaments distinct; staminodes 5, arising from hypanthium rim, subulate-filiform, inconspicuous; styles 2, connate in proximal 3, filiform, 0.1-0.4 mm, glabrous proximally; stigmas 2, linear along adaxial surfaces of style branches, papillate (100×). Utricles at least partly enclosed by hypanthium, opening irregularly. Seeds dark brown or black, ovoid to lenticular, slightly laterally compressed, shiny, smooth, marginal wing absent, appendage absent; embryo peripheral, curved. x = 9.[4] [more]
Illecebrum
Loeflingia
Herbs, annual. Taproots slender. Stems prostrate to erect, usually dichotomously branched at or near base, terete. Leaves opposite, basal leaves absent, axillary leaf clusters often present, weakly connate, sessile; stipules 2 per node, ± white to silvery, bristlelike, margins entire, apex acuminate; blade 1-veined or obscurely so, subulate to oblong, not succulent, apex blunt to spine-tipped. Inflorescences axillary, 1(-2) -flowered; bracts absent. Pedicels: flowers sessile. Flowers: perianth and androecium hypogynous; sepals ± distinct, green, linear to lanceolate, 1.8-6.5 mm, herbaceous, margins silvery, often scarious, apex acute to spinose; petals absent or rudimentary; nectaries not apparent; stamens 3-5, arising from base of ovary; filaments distinct; styles 3, distinct, filiform, shorter than 0.1 mm, glabrous proximally; stigmas 3, linear along adaxial surface of styles, papillate (30×). Capsules lanceoloid to ovoid, opening by 3 slightly recurved valves; carpophore absent. Seeds 25-35, tan with reddish brown band on curved edge, tear-shaped, plump, minutely papillate on broadly grooved edge, marginal wing absent, appendage absent.[5] [more]
Ortegia
Paronychia
Herbs, annual, biennial, or perennial, sometimes with woody base. Taproots filiform to stout. Stems prostrate, ascending, or erect, simple or branched, terete to angular. Leaves opposite, connate by stipules from adjacent leaves, petiolate (basal) or sessile (cauline) ; stipules 2 per node, often conspicuous, white or silvery, subulate to lanceolate or ovate, margins entire or fimbriate, apex subobtuse or acute to acuminate, unlobed or sometimes deeply 2-fid; blade 1-veined, linear to elliptic, oblanceolate, or spatulate, sometimes thickened and succulent, apex obtuse or acute to acuminate or spinose. Inflorescences terminal or sometimes axillary, frequently much-branched or congested cymes, or flowers solitary; bracts paired, dimorphic (resembling leaf blades and stipules), often concealing flowers. Pedicels erect in fruit. Flowers bisexual or rarely unisexual, some plants also having staminate unisexual flowers, others also having pistillate unisexual flowers, not woolly, with hairs ± straight or tips coiled, 0.1-0.3 mm; hypanthium cup-shaped, tapering or expanded distally; sepals (3-) 5, connate proximally, white or yellowish to green or reddish or purplish brown, subulate to linear-oblong, lanceolate, spatulate, or ovate, 0.4-4.5 mm, margins translucent to white, scarious or papery, apex defined by a usually prominent adaxial hood, ascending to slightly descending, rounded to triangular, sometimes absent (P. americana, P. erecta), apex obtuse or rounded, usually with terminal or subterminal cusp, crest, mucro, or prominent awn (often thickened-conic proximally, spinose distally) ; nectar secreted from within hypanthium; stamens usually 5; filaments distinct or connate proximally with alternating staminodes; staminodes absent or 5, arising from hypanthium rim, subulate to narrowly triangular, filiform, or oblong; styles 1-2(-3), distinct or often connate proximally 10- 10 of length, subcapitate to filiform, 0.07-3.2 mm, glabrous proximally; stigmas 2(-3), subterminal or linear along adaxial surface of style branches, obscurely papillate (50×). Utricles ovoid to globose or rarely 4-angular, membranous, indehiscent. Seeds brown, subglobose to ellipsoid, laterally compressed, smooth, marginal wing absent, appendage absent; embryo peripheral, curved. x = 7, 8, 9.[6] [more]
Polycarpaea
Polycarpaea is a genus of plant in family Caryophyllaceae. It contains the following species (but this list may be incomplete): [more]
Polycarpon
Herbs, annual. Taproots slender. Stems prostrate to erect, branched, terete to finely ridged. Leaves opposite or in whorls of 4, not connate, petiolate; stipules 2 per node, silvery, lanceolate to triangular-ovate, margins entire or irregularly cut, apex acuminate to aristate; blade 1-veined, spatulate or oblanceolate to ovate or elliptic, not succulent, apex obtuse, sometimes mucronate. Inflorescences terminal or axillary, dense or lax cymes; bracts paired or absent. Pedicels erect. Flowers: perianth and androecium perigynous; hypanthium minute, cup-shaped, not abruptly expanded distally; sepals distinct, green, lanceolate to elliptic or ovate, often keeled, 1-2.5 mm, herbaceous, margins white, scarious, apex acute, ± hooded, ± awned; petals often fugacious, 5, white, blade apex emarginate; nectaries between filament bases; stamens 3-5; filaments shortly connate distally around ovary; style 1, obscurely 3-branched, filiform, 0.1-0.3 mm, glabrous proximally; stigmas 3, linear along adaxial surface of style branches, papillate (30×). Capsules ovoid to spherical, opening by 3 incurved or twisting valves; carpophore present. Seeds ca. 8-15, whitish, ovoid to lenticular or triangular, laterally compressed to angular, papillate or granular, marginal wing absent, appendage absent. x = [7], 8, 9.[7] [more]
Pteranthus
Spergula
Spergula is a genus of about five plants belonging to the family Caryophyllaceae. Their usual English name is spurrey. Commonly found in grassland, the genus originated in the Northern Hemisphere, but is now found worldwide. [more]
Spergularia
Spergularia is a genus of the Caryophyllaceae family, and consists of sea-spurreys. [more]
Sphaerocoma
Stipulicida
Herbs, annual (or short-lived perennial?). Taproots filiform to stout. Stems diffuse to erect, repeatedly dichotomous, terete. Leaves opposite (cauline) or rosulate (basal), connate (distally) or not (proximally), petiolate (basal) or sessile (cauline) ; stipules 2 per node (cauline leaves) or forming tuft of to 14+ per node (basal leaves), white to tan, filiform, forming incised or notched nodal fringe; blade 1-veined, spatulate to suborbiculate (basal) or scalelike, subulate to triangular (cauline), not succulent, apex obtuse. Inflorescences terminal, compact, few-flowered cymes; bracts paired, scalelike. Pedicels erect. Flowers: perianth and androecium hypogynous; sepals distinct, reddish brown, elliptic to obovate, 0.8-2 mm, scarious, margins scarious, apex acute to obtuse or mucronate; petals 5, white, blade apex entire to erose; nectaries as minute, rounded lobes flanking filament bases; stamens 3-5; filaments distinct; styles 3, distinct or nearly so, capitate, ca. 0.2 mm, glabrous proximally; stigmas 3, terminal, obscurely papillate (30×). Capsules ellipsoid to globose, opening by 3 recurved valves; carpophore absent. Seeds ca. 20, golden chestnut to reddish brown, ± triangular, laterally compressed, lustrous, reticulate, marginal wing absent, appendage absent.[8] [more]
Telephium
Xerotia
More info about the Genus Xerotia may be found here.
Bibliography
- Akeroyd, J. R. 1993. Herniaria. In: T. G. Tutin et al., eds. 1993+. Flora Europaea, ed. 2. 1+ vol. Cambridge and New York. Vol. 1, pp. 182-184.
- Barneby, R. C. and E. C. Twisselmann. 1970. Notes on Loeflingia (Caryophyllaceae). Madrono 20: 398-408.
- Coker, P. D. 1962. Corrigiola litoralis L. J. Ecol. 50: 833-840.
- Core, E. L. 1939. A taxonomic revision of the genus Siphonychia. J. Elisha Mitchell Sci. Soc. 55: 339-345.
- Core, E. L. 1940. Notes on the mid-Appalachian species of Paronychia. Virginia J. Sci. 1: 110-116.
- Core, E. L. 1941. The North American species of Paronychia. Amer. Midl. Naturalist 26: 369-397.
- Duke, J. A. 1961. Preliminary revision of the genus Drymaria. Ann. Missouri Bot. Gard. 48: 173-268.
- Fernald, M. L. 1936c. Notes on Paronychia, Anychia. Rhodora 38: 416-421.
- Gilbert, M. G. 1987. The taxonomic position of the genera Telephium and Corrigiola. Taxon 36: 47-49.
- Hartman, R. L. 1974. Rocky Mountain species of Paronychia (Caryophyllaceae): A morphological, cytological, and chemical study. Brittonia 26: 256-263.
- Hermann, F. 1937. Übersicht über die Herniaria-Arten des Berliner Herbars. Repert. Spec. Nov. Regni Veg. 42: 203-224.
- Howell, J. T. 1941. Notes on Polycarpon. Leafl. W. Bot. 3: 80.
- Shinners, L. H. 1962c. Siphonychia transferred to Paronychia (Caryophyllaceae). Sida 1: 101-103.
- Turner, B. L. 1983b. The Texas species of Paronychia (Caryophyllaceae). Phytologia 54: 9-23.
- Vogel, A. 1997. Die Verbreitung, Vergesellschaftung und Populationsökologie von Corrigiola litoralis, Illecebrum verticillatum und Herniaria glabra (Illecebraceae). Berlin.
- Ward, D. B. 1977c. Keys to the flora of Florida. 2. Paronychia (Caryophyllaceae). Phytologia 35: 414-41
- Williams, F. N. 1896. A revision of the genus Herniaria. Bull. Herb. Boissier 4: 556-570.
Footnotes
- Ronald L. Hartman "Cardionema". in Flora of North America Vol. 5. Oxford University Press. Online at EFloras.org.
- John W. Thieret, Richard K. Rabeler "Corrigiola". in Flora of North America Vol. 5. Oxford University Press. Online at EFloras.org.
- Ronald L. Hartman "Drymaria". in Flora of North America Vol. 5. Oxford University Press. Online at EFloras.org.
- John W. Thieret, Ronald L. Hartman, Richard K. Rabeler "Herniaria". in Flora of North America Vol. 5. Oxford University Press. Online at EFloras.org.
- Ronald L. Hartman, Richard K. Rabeler "Loeflingia". in Flora of North America Vol. 5. Oxford University Press. Online at EFloras.org.
- Ronald L. Hartman, John W. Thieret, Richard K. Rabeler "Paronychia". in Flora of North America Vol. 5. Oxford University Press. Online at EFloras.org.
- John W. Thieret, Richard K. Rabeler "Polycarpon". in Flora of North America Vol. 5. Oxford University Press. Online at EFloras.org.
- Ann Swanson, Richard K. Rabeler "Stipulicida". in Flora of North America Vol. 5. Oxford University Press. Online at EFloras.org.
Sources
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