Overview
The Magnoliaceae, or Magnolia Family, is a family in the order Magnoliales. It consists of two subfamilies:
- Magnolioideae, of which Magnolia is the most well-known genus.
- Liriodendroidae, a monogeneric subfamily, of which Liriodendron (Tulip trees) is the only genus.
Unlike most angiosperms, whose flower parts are in rings, the Magnoliaceae have their stamens and pistils in spirals on a conical receptacle.1] This arrangement is found in some fossil plants and is believed to be a basal or early condition for angiosperms. The flowers are also have parts not distinctly differentiated into sepals and petals, while angiosperms that evolved later tend to have distinctly differentiated sepals and petals. The poorly differentiated perianth parts that occupy both positions are known as tepals.
The family has approximately 225 species in 7 genera, although some classification systems include all of subfamily Magnoioideae in genus Magnolia. The family ranges across eastern North America, Mexico and Central America, the West Indies, tropical South America, southern and eastern India, Sri Lanka, Indochina, Malesia, China, Japan, and Korea.
Description
The monophyly of Magnoliaceae is supported by a number of shared morphological characters among the various genera included in the family. Most have flowers that are bisexual (with the exception of Kmeria and some species of Magnolia section Gynopodium), radial, and with an elongate receptacle. Leaves are alternate, simple, and sometimes lobed. The inflorescence is a solitary, showy flower with indistinguishable petals and sepals. Sepals range from six to many; stamens are numerous and feature short filaments which are poorly differentiated from the anthers. Carpels are usually numerous, distinct, and on an elongate receptacle.[1] The fruit is an aggregate of follicles which usually become closely appressed as they mature and open along the abaxial surface. Seeds have a fleshy coat and color that ranges from red to orange (except Liriodendron). Magnoliaceae flowers are beetle pollinated, except for Liriodendron, which is bee pollinated. The carpals of Magnolia flowers are especially thick to avoid damage by beetles who, land, crawl, and feast on them. The seeds of Magnolioideae are bird dispersed while the seeds of Liriodendron are wind dispersed.
Biogeography
Due to i ts great age, the geographical distribution of Magnoliaceae has become disjunct or fragmented as a result of major geologic events such as ice ages, continental drift, and mountain formation. This distribution pattern has isolated some species while keeping others in close contact. Extant species of Magnoliaceae are widely distributed in temperate and tropical Asia from the Himalayas to Japan and southwest through Malaysia and New Guinea. Asia is home to approximately two thirds of the species in Magnoliaceae, with the remainder of the family spread across the Americas with temperate species extending into the southern United States and tropical elements extending into Brazil and the West Indies.
Systematics
Due to this great deal of family wide morphological similarity, no consensus has yet emerged on the number of genera in the family. The development of DNA sequencing at the end of the twentieth century had a profound impact on the research of phylogenetic relationships within the family. The employment of ndhF and cpDNA sequences has refuted many of the traditionally accepted phylogenetic relationships within Magnoliaceae. For example, it has shown the genera Magnolia and Michelia to be paraphyletic when the remaining four genera of Magnolioideae are split out. In fact, even many of the subgenera (Magnolia subg. Magnolia, Magnolia subg. Talauma) have been found to be paraphyletic. Although no completely resolved phylogeny for the family has yet been determined, these technological advances have allowed systematists to broadly circumscribe major lineages.[2]
Economic Significance
As a whole, Magnoliaceae is not an economically significant family. With the exception of a great deal of ornamental cultivation, the economic significance of Magnolias is generally confined to the use of wood from certain timber species and the use of bark and flowers from several species believed to possess medicinal qualities. Magnolias have a rich cultural tradition in China where references to their healing qualities go back thousands of years. The Chinese have long used the bark of Magnolia officinalis, a magnolia native to the mountains of China with large leaves and fragrant white flowers, as a remedy for cramps, abdominal pain, nausea, diarrhea, and indigestion. Certain magnolia flowers, such as the buds of Magnolia liliflora, have been used to treat chronic respiratory infections, sinus infections, and lung congestion. Recently, magnolia bark has become incorporated into alternative medicine in the west where tablets made from the bark of M. officinalis have been marketed as an aid for anxiety, allergies, asthma, and weight-loss. Preliminary studies have suggested that compounds found in magnolia bark might have antibacterial and antifungal properties. To this day, however, there has never been a large scale study on the health effects of magnolia bark or flowers.
Photos
Taxonomy
The Subfamily Magnolioideae is a member of the Family Magnoliaceae. Here is the complete "parentage" of Magnolioideae:
- Domain: Eukaryota
Whittaker & Margulis,1978 - eukaryotes
- Kingdom: Plantae
Haeckel, 1866
- Subkingdom: Viridaeplantae
Cavalier-Smith, 1981 - Green Plants
- Phylum: Tracheophyta
Sinnott, 1935 Ex Cavalier-Smith, 1998 - Vascular Plants
- Subphylum: Euphyllophytina
- Infraphylum: Radiatopses
Kenrick & Crane, 1997
- Class: Magnoliopsida
Brongniart, 1843 - Dicotyledons
- Subclass: Magnoliidae
Novák Ex Takhtajan, 1967
- Superorder: Magnolianae
Takhtajan, 1967
- Order: Magnoliales
Bromhead, 1838
- Family: Magnoliaceae
(mag-NO-lee-uh)
A.l. De Jussieu, 1789, Nom. Cons.
- Subfamily: Magnolioideae
- Family: Magnoliaceae
(mag-NO-lee-uh)
A.l. De Jussieu, 1789, Nom. Cons.
- Order: Magnoliales
Bromhead, 1838
- Superorder: Magnolianae
Takhtajan, 1967
- Subclass: Magnoliidae
Novák Ex Takhtajan, 1967
- Class: Magnoliopsida
Brongniart, 1843 - Dicotyledons
- Infraphylum: Radiatopses
Kenrick & Crane, 1997
- Subphylum: Euphyllophytina
- Phylum: Tracheophyta
Sinnott, 1935 Ex Cavalier-Smith, 1998 - Vascular Plants
- Subkingdom: Viridaeplantae
Cavalier-Smith, 1981 - Green Plants
- Kingdom: Plantae
Haeckel, 1866
The Subfamily Magnolioideae is further organized into finer groupings including:
- Tribe (14): Arabideae · Bombycillini · Bougainvilleeae · Cisteae · Corydaleae · Gentianeae · Heliantheae · Lilieae · Magnolieae · Michelieae · Narcisseae · Oleeae · Tulipeae · Verbeneae
- Genus (105): Aethionema · Agapanthus · Allium · Angraecum · Arthropteris · Aubrieta · Baccaurea · Bergenia · Blandfordia · Blechnum · Bombycilla · Botrychium · Bougainvillea · Brachyotum · Campanula · Ceanothus · Ceiba · Centaurea · Centaurium · Cephalanthera · Cephalaria · Cephalotaxus · Chamaecyparis · Chlorogalum · Cistus · Citrus · Cleome · Cnidoscolus · Corydalis · Cosmos · Cotoneaster · Crataegosorbus · Cyclamen · Davallia · Deutzia · Diascia · Dichelostemma · Diospyros · Draba · Dryopteris · Epilobium · Eucomis · Euonymus · Festuca · Fremontodendron · Galpinia · Garrya · Gentiana · Geranium · Gladiolus · Gnolia · Heliotropium · Helleborus · Hemerocallis · Hippocrepis · Holboellia · Hypoxis · Juncus · Juniperus · Kniphofia · Lampranthus · Lavandula · Leucadendron · Leucosidea · Lilium · Limnanthes · Linum · Lupinus · Macrozamia · Magnolia · Mahonia · Malephora · Mandevilla · Manglietia · Marrubium · Michelia · Narcissus · Nuxia · Oplismenus · Oxalis · Paeonia · Pelargonium · Phacelia · Plantago · Polypodium · Polystichum · Pyrrhopappus · Rehmannia · Rhodocoma · Salix · Saxifraga · Silybum · Sorbus · Styrax · Syringa · Tecomanthe · Thamnochortus · Thlaspi · Tulipa · Ulmus · Vanilla · Verbena · Viburnum · Wattakaka · Zauschneria
- Species: ZipcodeZoo has pages for 1,730 species, subspecies, varieties, forms, and cultivars in the Subfamily Magnolioideae.
Genera
Aethionema
Perennial or annual herbs, often woody below, branched, erect or suberect, leafy, glabrous or rarely papillose. Leaves simple, usually sessile or subsessile, oblong or linear, glaucous. Racemes corymbose, usually many flowered, ebracteate. Flowers mediocre, rose, lilac or white, rarely yellowish; pedicls filiform, usually spreading in fruit. Sepals oblong, obtuse, rounded at apex; inner ±saccate at base; outer often somewhat hooded at apex. Petals obovate, cuneate or clawed, rarely oblong; claw 1-3-nerved. Stamens 6; filaments of longer stamens append-aged, dilated or linear; anthers often apiculate, ovate-orbicular. Lateral nectar glands in pairs, minute, semiglobose; middle usually absent. Ovary ± ellipsoid with narrowly flattened margin, 1-2-locular with 1-2 (rarely 3-4) ovules in each locule; stigma capitate, sub-sessile or on distinct short style. Siliculae ovate, elliptic or suborbicular, laterally flattened, usually winged, dehiscent, (rarely heterocarpic with dehiscent and indehiscent fruits), 1-4-seeded; apex generally deeply notched or emarginate; wing entire or variously dentate; seed ovate, brown, often minutely papillose; radicle incumbent, oblique or accumbent.[1] [more]
Agapanthus
Agapanthus , the "Lily of the Nile", is a genus of flower plants with six to ten species depending on how the different species are classified. They are all perennial plants native to South Africa. They have been placed either in the family Alliaceae, or separated into their own monogeneric family Agapanthaceae (e.g. Indices Nominum Supragenericorum Plantarum Vascularium). [more]
Allium
Herbs, perennial, scapose, from tunicate bulbs, with onion odor and taste. Bulbs solitary or clustered, dividing at base, or on rhizomes, reforming annually; outer coats generally brown or gray, smooth, fibrous, or with cellular reticulation (generally important in identification) ; inner coats membranous. Leaves generally withering from tip by anthesis, usually persistent, 1-12, basal; blade usually linear, terete, channeled, or flat (carinate in A. sativum, A. praecox, A. tuberosum, A. rotundum, A. neapolitanum, A. triquetrum, A. unifolium, and A. lacunosum), straight or ± falcate (coiled or circinate in A. nevadense and A. atrorubens), broader in A. victorialis and A. tricoccum, not petiolate (except in A. tricoccum and A. victorialis) . Scape usually persistent, terete or flattened. Inflorescences umbellate, flowering centripetally (centrifugally in A. schoenoprasum), sometimes replaced totally or partially by bulbils, subtended by spathe bracts; bracts conspicuous, ± fused, usually 3+-veined, equaling pedicel except in some introduced species, membranous. Flowers erect (pendent in A. triquetrum) ; tepals 6, in 2 similar whorls, ± distinct, petallike, usually becoming becoming dry and persisting; stamens 6, epipetalous; filaments in all but 1 native species broad at base, fused into ring (some introduced species and A. victorialis appendaged), linear, generally glabrous (A. rotundum and A. hoffmanii papillose to ciliate proximally) ; anthers and pollen variously colored; ovary superior, 3-lobed, sometimes crested with processes, 3-locular, usually 2 ovules per locule (6-8 in A. nigrum), crest processes 3 or 6, smooth except in A. haematochiton, A. sharsmithiae, and A. lacunosum; style 1; stigma capitate to ± 3-lobed; pedicel erect or spreading (lax in A. triquetrum) . Fruits capsular, dehiscence loculicidal. Seeds black, obovoid, finely cellular-reticulate, cells smooth or minutely roughened, with 1-8 papillae, without caruncle except in A. triquetrum. x = 7, 8, 9.[2] [more]
Angraecum
The Angraecum, abbreviated as Angcm in horticultural trade, common name Angrec or Comet Orchid, contains about 220 species, some of them among most magnificent of all orchids. They are quite varied vegetatively and florally and are adapted to dry tropical woodland habitat and have quite fleshy leaves as a consequence. Most are epiphytes, but a few are lithophytes. [more]
Arthropteris
Aubrieta
Aubrieta is a of about 12 species of flowering plants in the cabbage family Brassicaceae. The genus is named after Claude Aubriet, a French flower-painter. It originates from southern Europe east to central Asia but is now a common garden escape throughout Europe. It is a low, spreading plant, hardy, evergreen and perennial, with small violet, pink or white flowers, and inhabits rocks and banks. It prefers light, well-drained soil, is tolerant of a wide pH range, and can grow in partial shade or full sun. [more]
Baccaurea
Baccaurea is a of flowering plant belonging to the family Phyllanthaceae. The genus comprises about 80 species, distributed from Indomalesia to the West Pacific. [more]
Bergenia
Herbs perennial, forming large clumps. Rhizomes creeping, large, thick, scaly. Leaves all basal, ± persistent, simple, waxy, often leathery; petiole short, broad, sheathing at base; leaf blade thick, margin entire, crenate, or dentate. Infloresences cymose, bracteate. Flowers showy, large. Sepals 5. Petals 5, white, pink, red, or purple. Stamens 10. Carpels 2, basally connate; ovary 1/4 subsuperior, proximally 2-loculed with axile placentation and distally 1-loculed with marginal placentation; styles 2; ovules many. Fruit a capsule. Seeds numerous, dark brown, small.[3] [more]
Blandfordia
Blandfordia is a of flowering plants which are native to eastern Australia. Plants in this genus are commonly referred to as Christmas Bells due to the shape of their flowers and the timing of their flowering season in Australia. Blandfordia is the sole genus in the family Blandfordiaceae and was named by English botanist James Edward Smith in 1804 in honour of George Spencer Churchill, the Marquis of Blandford. [more]
Blechnum
Plants terrestrial or rarely on rock. Stems creeping to ascending or erect, slender to stout, sometimes climbing [rarely subarborescent]; scales brown or black. Leaves monomorphic or variously dimorphic, cespitose to scattered. Blades pinnatifid to 1-pinnate, rarely simple or 2-pinnate. Rachis and costae glabrous, scaly, or hairy abaxially. Veins free, often forked. Sori borne on vascular commissures parallel to costae, 1 per side, normally uninterrupted, linear, continuous along length of costa. Spores with perine smooth to variously winged or rugose. x = 28, 29, 31, 32, 34, 36.[4] [more]
Bombycilla
The waxwings form the genus Bombycilla of birds. According to most authorities, this is the only genus placed in the family Bombycillidae. [more]
Botrychium
Plants terrestrial. Roots occasionally branching laterally, yellowish to black, 0.5--2 mm diam., smooth or with corky ridges, not proliferous. Stems upright, forming caudex to 5 mm thick; gemmae absent or minute, spheric. Trophophores ascending to perpendicular to stem, sessile or stalked; blades linear, oblong, or deltate, simple to 5-pinnate, 4--25 × 1--35 cm. Pinnae (reduced to segments in many species) spreading to ascending, fan-shaped to lanceolate to linear; margins entire to dentate to lacerate, apex rounded or acute; veins free, arranged like ribs of fan or pinnate. Sporophores normally 1 per leaf, 1--3-pinnate, long-stalked, borne at ground level to high on common stalk. Sporangial clusters with sporangia sessile to short-stalked, almost completely exposed, borne in 2 rows on pinnate (except in very small plants) sporophore branches. Gametophytes broadly ovate, unbranched, 1--3 × 1--10 mm. x =44, 45, 92.[5] [more]
Bougainvillea
A genus in the Kingdom Animalia. [more]
Brachyotum
Brachyotum is a genus of in family Melastomataceae. [more]
Campanula
Plants perennial or annual, erect trailing or decumbent, glabrous, pubescent, or hirsute. Leaves simple, alternate or forming rosettes at the base. Inflorescence 1-many flowered, with racemes or spikes. Flowers blue to purple or white. Sepals 5, with or without reflexed appendages between lobes; calyx tube adnate to the ovary, segments 5-lobed. Corolla campanulate, funnel-shaped or tubular. Stamens 5, free, filaments dilated at the base. Ovary 3-locular; style cylindrical; stigmas 3. Fruit a capsule, elongated to ovoid, obovoid or round, with membran¬ous walls; dehiscence by irregular pores at the bases or the sides. Seeds minute, numerous.[6] [more]
Ceanothus
Ceanothus is a genus of about 50–60 species of shrubs or small trees in the buckthorn family Rhamnaceae. The genus is confined to North America, the center of its distribution in California, with some species (e.g. C. americanus) in the eastern United States and southeast Canada, and others (e.g. C. coeruleus) extending as far south as Guatemala. Most are shrubs 0.5–3 m tall, but C. arboreus and C. thyrsiflorus, both from California, can be small trees up to 6–7 m tall. [more]
Ceiba
Trees, deciduous; trunk swollen and/or buttressed, trunk and branches usually spiny. Petiole long; leaf blade palmately compound; leaflets 3-5(-9), shortly petiolulate, both surfaces glabrous, abaxially gray-white, base attenuate, margin serrate or entire. Flowers blooming before or simultaneously with leaf flush, solitary or 2-15-fascicled, pendulous, axillary or subterminal, actinomorphic, rarely zygomorphic. Calyx campanulate, truncate or irregularly 3-5(-12) -lobed, thickly fleshy, persistent. Petals 5, pink or yellow-white, connate at base and coherent to staminal tube, falling with stamens and style. Stamens (3-) 5-15; filament tube short, sometimes thickened or with sterile appendages, free parts of filament long to absent, each bearing 2(or 3) straight to strongly twisted thecae. Ovary syncarpous, 5-locular; ovules many per cell; style filiform; stigma capitate to lobulate. Capsule oblong or nearly obovate, pendulous, woody or leathery, loculicidally dehiscent into 5(or 6) valves, inner walls densely woolly. Seeds many, enclosed in wool.[7] [more]
Centaurea
Annuals, biennials, or perennials, 20-300 cm, glabrous or tomentose. Stems erect, ascending, or spreading, simple or branched. Leaves basal and cauline; petiolate or sessile; proximal blade margins often ± deeply lobed, (spiny in C. benedicta ), distal ± smaller, often entire, faces glabrous or ± tomentose, sometimes also villous, strigose, or puberulent, often glandular-punctate. Heads discoid, disciform, or radiant, borne singly or in corymbiform arrays. Involucres cylindric or ovoid to hemispheric . Phyllaries many in 6-many series, unequal, proximal part appressed, body margins entire. distal parts expanded into erect to spreading, usually ± dentate or fringed, linear to ovate appendages, spine. tipped or spineless. Receptacles flat, epaleate, bristly. Florets 10-many; outer usually sterile, corollas slender and inconspicuous to much expanded, ± bilateral; inner fertile, corollas white to blue, pink, purple, or yellow, bilateral or radial, often bent at junction of tubes and throats, lobes linear-oblong, acute; anther bases tailed, apical appendages oblong; style branches: fused portions with minutely hairy nodes, distinct portions minute. Cypselae ± barrel-shaped, ± compressed, smooth or ribbed, apices entire (denticulate in C. benedicta ), glabrous or with fine, 1-celled hairs, attachment scar. lateral (with or without elaiosomes) ; pappi 0 or ± persistent, of 1-3 series of smooth or minutely barbed, stiff bristles or narrow scales . x = 8, 9, 10, 11, 12, 13, 15.[8] [more]
Centaurium
Annuals. Stems erect, obscurely 4angled. Leaves opposite. Cymes pseudodichotomous, sometimes spicate. Flowers [4 or] 5merous. Calyx lobed nearly to base. Corolla salverform, lobes shorter than tube. Stamens inserted at throat of corolla tube; filaments filiform, erect at first, helically coiled later; anthers dehiscing longitudinally. Ovary partly 2-locular. Style linear; stigma lobes orbicular. Capsules 2valved, many seeded. Seed coat alveolate.[9] [more]
Cephalanthera
Herbs, perennial, achlorophyllous. Roots fleshy, slender, scattered along slender rhizome. Stems leafy. Leaves several, alternate; blade spreading, plicate, or reduced to appressed sheathing bracts. Inflorescences terminal, solitary spikes, lax; floral bracts often foliaceous, large to minute, scarious distally. Flowers few-many, resupinate, sessile, showy; lip white with yellow markings, divided by median constriction, saccate, distal end with fleshy parallel adaxial lamellae; pollinaria absent; pollinia 2; viscidia absent. Fruits capsules.[10] [more]
Cephalaria
Herbs, glabrous to pilose. Leaves sometimes divided. Involucral bracts coriaceous, the receptacular ones larger. Calyx cupular, limb many-toothed. Corolla 4-fid. Involucel 4-8-angled, limb toothed.[11] [more]
Cephalotaxus
Morphological characters and geographical distribution are the same as those for the family.[12] [more]
Chamaecyparis
Trees (rarely shrubs). Branchlets terete or rhombic in cross section, in fan-shaped or pinnately flattened sprays. Leaves opposite in 4 ranks. Adult leaves usually appressed, lateral and facial pairs similar, closely overlapping, scalelike, free portion of long-shoot leaves to ca. 7 mm; abaxial glands present or absent, circular to linear. Pollen cones with 2--3 pairs of sporophylls, each sporophyll with 2--4 pollen sacs. Seed cones maturing and opening in 1--2 years, nearly globose, glaucous, 4--12 mm; scales persistent, 2--5(--6) pairs, valvate, peltate or basifixed, thick and woody, terminal pair often fused. Seeds 1--4 per cone scale, lenticular, equally 2-winged; cotyledons 2--3. x = 11.[13] [more]
Chlorogalum
Herbs, perennial, from tunicate bulbs. Leaves basal; blade linear, reduced to scarious bracts in inflorescence. Inflorescences paniculate. Flowers scattered at intervals along branches, 1-several per node, each subtended by small bract; perianth white, purple, or pinkish; tepals 6, distinct, linear to oblong, with nectaries at base, persisting in fruit and twisting together above capsule; stamens 6, inserted on bases of tepals; anthers versatile; style slightly 3-lobed at apex; pedicel articulate. Fruits capsular, 3-valved, dehiscence loculicidal. Seeds 1 or 2 per locule.[14] [more]
Cistus
Citrus
Evergreen, small trees or shrubs, often spiny. Leaves simple, alternate, glandular punctate, petiole winged or margined. Flowers perfect or staminate, solitary or clustered in axillary racemes. Calyx 4-5-lobed, glabrous or pubescent. Petals (4-) 5(-8). Stamens 4-10 times the petals, polyadelphous. Ovary 10-14-locular, ovules biseriate or collateral. Fruit a fleshy hesperidium, globose to mamillate-oblong to oblate, rind tight or loose, with oil glands. Seeds embedded in pulpy vesicles.[15] [more]
Cleome
Cleome is a of flowering plants in the family Cleomaceae. Previously it had been placed in family Capparaceae, until DNA studies found the Cleomaceae genera to be more closely related to Brassicaceae than Capparaceae. The APG II system allows for Cleome and the other members of Cleomaceae to be included in Brassicaceae. [more]
Cnidoscolus
Cnidoscolus is a genus of the family Euphorbiaceae. [more]
Corydalis
A genus in the Kingdom Animalia. [more]
Cosmos
Annuals [perennials or subshrubs], 30-250 cm. Stems usually 1, erect or ascending, branched distally or ± throughout. Leaves mostly cauline; opposite; petiolate or sessile; blades usually 1-3-pinnately lobed [undivided], ultimate margins usually entire, faces usually glabrous, sometimes glabrate, hispid, puberulent, or scabridulous. Heads radiate, borne singly or in corymbiform arrays. Calyculi of [5-]8 basally connate, ± linear to subulate, herbaceous (striate) bractlets. Involucres hemispheric or subhemispheric [cylindric], 3-15 mm diam. Phyllaries persistent, [5-]8 in ± 2 series, distinct, lanceolate, lance-oblong, lance-ovate, or oblong, ± equal, membranous or herbaceous, margins ± scarious. Receptacles flat, paleate; paleae falling, linear, flat or slightly concave-convex, scarious (entire). Ray florets [0, 5] 8 (more in "double" cultivars), neuter; corollas white to pink or purple, or yellow to red-orange. Disc florets 10-20[-80+], bisexual, fertile; corollas yellow [orange] (at least distally), tubes shorter than funnelform throats, lobes 5, ± deltate (staminal filaments hairy near anthers; style branches linear, flattened, thicker distally, hirtellous, appendages relatively slender). Cypselae (dark brown or black) relatively slender, quadrangular-cylindric or -fusiform [outer somewhat obcompressed], sometimes slightly arcuate, attenuate-beaked, not winged [winged], faces glabrous or hispid to scabridulous or ± setose, sometimes papillate, usually with 1 groove; pappi persistent [falling], of 2-4[-8] retrorsely [antrorsely] barbed awns, sometimes 0. x = 12.[16] [more]
Cotoneaster
Shrubs, rarely small trees, erect, decumbent, or prostrate, deciduous, semievergreen, or evergreen. Branchlets mostly terete, rarely slightly angulate, unarmed. Winter buds small; scales several, imbricate, exposed. Leaves alternate, simple, shortly petiolate; stipules caducous, usually subulate, small; margin of leaf blade entire, venation camptodromous. Inflorescences terminal or axillary, cymose or corymbose, sometimes flowers several fascicled or solitary. Hypanthium turbinate or campanulate, rarely cylindric, adnate to ovary. Sepals 5, persistent, short. Petals 5, erect or spreading, imbricate in bud, white, pink, or red. Stamens 10-20(-22), inserted in mouth of hypanthium. Ovary inferior or semi-inferior, 2-5-loculed; carpels 2-5, connate abaxially, free adaxially; ovules 2 per carpel, erect; styles 2-5, free; stigmas dilated. Fruit a drupe-like pome, red, brownish red, or orange to black, with persistent, incurved, fleshy sepals, containing pyrenes; pyrenes (1 or) 2-5, bony, 1-seeded; seeds compressed; cotyledons plano-convex.[17] [more]
Crataegosorbus
Cyclamen
Cyclamen is a of 23 species of flowering plants, traditionally classified in the family Primulaceae, but in recent years reclassified in the family Myrsinaceae (Kallersjo et al. 2000). The genus is most widely known by its scientific name Cyclamen being taken into common usage; other names occasionally used include sowbread and sometimes, confusingly, Persian violet (it is not related to the violets), or primrose (neither is it a primrose). [more]
Davallia
Davallia (deersfoot fern, hare's foot fern) is a genus of about 40 species of in the family Davalliaceae. They are epiphytic ferns, with fronds arising from long aerial rhizomes which grow on and over thick bark on trees or on rock crevices. [more]
Deutzia
Shrubs stellate hairy. Branchlets opposite; buds enclosed by imbricate scales. Leaves opposite, exstipulate, subdeciduous. Inflorescences racemose, paniculate, corymbose, or cymose, rarely a solitary flower. Calyx tube adnate to ovary, campanulate, 5-lobed. Petals 5, induplicate, valvate, or imbricate. Stamens 10(-15), 2-seriate; filaments subulate, flat, or dilated and apex 2-dentate; anthers shortly stalked, subglobose. Ovary inferior, rarely subinferior, 3-5-loculed; ovules numerous, in many series on fleshy placenta. Styles 3(-5), free; stigma terminal or decurrent. Fruit a capsule, subglobose, 3(-5) -valved, dehiscing loculicidally or between styles. Seeds numerous, oblong, compressed; testa membranous, reticulate, apex winged; embryo borne in middle of fleshy endosperm.[18] [more]
Diascia
Dichelostemma
Herbs, perennial, scapose, from fibrous-coated corms. Leaves 1-5, basal; blade narrowly lanceolate, usually keeled and channeled, margins entire. Scape solitary, usually weak, curved to twining, cylindrical, smooth to scabrous. Inflorescences umbellate or racemose, usually dense, 2-20-flowered, bracteate; bracts 2-4, ± papery, not enclosing flower buds. Flowers: perianth 6-tepaled, distinctly connate proximally into tube, tube cylindrical, ovoid, or campanulate, occasionally globose or urceolate, soft, limb lobes similar; perianth appendages arising from intersection of perianth tube and limb lobes, leaning toward or away from anthers, forming corona; anthers basifixed, held close to style; stamens 3 (6 in Dichelostemma capitatum), epitepalous; filaments entirely adnate to perianth tube; staminodia absent (except in D. volubile) ; pistil 3-carpellate; ovary superior, sessile or stipitate, 3-locular, ovules several; style 1; stigma weakly 3-lobed; pedicel erect or flexuous, articulate beneath perianth, usually shorter than flowers. Fruits capsular, 3-angled, usually ovoid, firm, dehiscence loculicidal. Seeds black, sharply angled, coat with crust. x = 9 (8 in D. ida-maia).[19] [more]
Diospyros
Trees or shrubs, deciduous or evergreen. Terminal buds absent. Branchlet tips sometimes forming a spine. Leaves alternate, occasionally minutely translucent dotted or with gland pits. Flowers dioecious or polygamous. Male flowers in axillary cymes, usually on basal part of current year's branchlets, deciduous soon after anthesis; stamens 4 to numerous, often paired and forming 2 whorls; ovary rudimentary. Female flowers usually solitary, axillary; staminodes 1--16 or absent; stigma often 2-cleft. Calyx usually 3--5(--7) -lobed, sometimes truncate. Corolla urn-shaped, campanulate, or tubular, 3--5(--7) -lobed, deciduous. Berries fleshy to somewhat leathery, usually with an enlarged persistent calyx. Seeds 1--10(or more), often laterally compressed.[20] [more]
Draba
Herbs perennial, rarely annual, biennial (or subshrubs with woody stems). Trichomes simple, forked, stellate, malpighiaceous, or dendritic, stalked or sessile, often more than 1 kind present. Stems erect or ascending, sometimes prostrate, leafy or leafless and plants scapose. Basal leaves petiolate, often rosulate, simple, entire or toothed, rarely lobed. Cauline leaves petiolate or sessile, cuneate or auriculate at base, entire or dentate, sometimes absent. Racemes bracteate or ebracteate, elongated or not in fruit. Fruiting pedicels slender, erect, ascending, or divaricate. Sepals ovate, oblong, or elliptic, base of lateral pair not saccate or subsaccate, margin usually membranous. Petals yellow, white, pink, purple, orange (or rarely red) ; blade obovate, spatulate, oblong, oblanceolate, orbicular, or linear, apex obtuse, rounded, or rarely emarginate; claw obscurely to strongly differentiated from blade. Stamens 6, tetradynamous; filaments dilated or not at base; anthers ovate or oblong, obtuse at apex. Nectar glands 1, 2, or 4, distinct or confluent and subtending bases of all stamens; median glands present or absent; lateral glands toothlike, semiannular, or annular. Ovules 4 to numerous per ovary. Fruit dehiscent, silicles or rarely siliques, ovate, elliptic, oblong, orbicular, ovoid, globose, lanceolate, or linear, latiseptate or terete, sometimes spirally twisted; valves distinctly or obscurely veined, glabrous or pubescent; replum rounded; septum complete, membranous, translucent; style distinct or obsolete, glabrous; stigma capitate, entire or slightly 2-lobed. Seeds biseriate, wingless (or rarely winged), oblong, ovate, or orbicular, flattened; seed coat minutely reticulate, not mucilaginous when wetted; cotyledons accumbent.[21] [more]
Dryopteris
Plants terrestrial, rarely on rock. Stems short-creeping to erect, stolons absent. Leaves monomorphic, green through winter or dying back in winter. Petiole ca. 1/4--2/3 blade length, bases swollen or not; vascular bundles more than 3, arranged in an arc, ± round in cross section. Blade deltate-ovate to lanceolate, 1--3-pinnate-pinnatifid, gradually reduced distally to pinnatifid apex, herbaceous to somewhat leathery. Pinnae not articulate to rachis, segment margins entire, crenate, or serrate, spinulose or not; proximal pinnae reduced (several pairs), same size as or enlarged relative to more distal pinnae, sessile to petiolulate, equilateral or often inequilateral with pinnules on basiscopic side longer than those on acroscopic side; costae adaxially grooved, grooves continuous from rachis to costae to costules; indument of linear to ovate scales abaxially, also sometimes with glands, blades ± glabrous adaxially. Veins free, forked. Sori in 1 row between margin and midrib, round; indusia round-reniform, attached at narrow sinus, persistent or caducous. Spores brownish, coarsely rugose or with folded wings. x = 41.[22] [more]
Epilobium
Herbs perennial [or annual, sometimes suffrutescent], with leafy rosettes, stolons, soboles (shoots), or turions (subterranean globose buds with fleshy scales) . Stems glabrous to pubescent, often with lines of hairs decurrent from margins of petioles. Leaves opposite, becoming alternate and bractlike in inflorescence; petiolate or sessile; stipules absent; bracteoles absent. Inflorescences simple or branched racemes, panicles, spikes, or corymbs. Flowers 4-merous, often protandrous, with floral tube, producing nectar at base of style. Petals pink to rose-purple or white [or rarely cream-colored or orange-red], obcordate or obtrullate, notched at apex. Stamens 8, in two unequal whorls; pollen yellow, shed in tetrads. Style erect; stigma entire or 4-lobed. Fruit an elongate, slender capsule, 4-loculed, loculidical. Seeds many [or rarely only 4], generally with terminal coma of silky hairs [or coma rarely lacking]. 2n = [18, 20, 24, 26, 28, 30, 32,] 36, [38, 60].[23] [more]
Eucomis
Euonymus
Trees or shrubs, the latter sometimes scandent. Leaves oppsite; stipules caducous. Flowers bisexual, 4-5-merous. Calyx flat or recurved. Disc broad, fleshy, 4-5-lobed. Petals 4-5, rounded, spreading, often with coloured veins. Stamens 4-5, inserted on the disc. Ovary sunken in the disc; style short. Fruit a capsule, 3-5-lobed, angled or winged, rarely echinate, dehiscence loculicidal. Seeds 1-3 in each cell, enclosed in a fleshy aril.[24] [more]
Festuca
Perennials, tufted, shoots extra- or intra-vaginal. Leaf sheath margins usually free, rarely connate, sometimes with auricles; leaf blades folded to conduplicate and filiform, sometimes flat; ligule membranous. Inflorescence an open, contracted or spikelike panicle. Spikelets with 2 to several florets, uppermost floret usually reduced; rachilla usually scabrid, rarely smooth or pubescent; disarticulating above glumes and between florets; glumes usually unequal, herbaceous to scarious, rarely subleathery, lower glume often small, 1-veined, upper glume usually shorter than lowest lemma, 3(-5) -veined; lemmas usually similar in texture to glumes, often subleathery at least with age, usually ± laterally compressed but not keeled, rounded on back at least toward base, usually 5-veined, veins sometimes prominent, apex acuminate, entire or notched, awned or awnless; palea subequal to lemma, keels scabrid, rarely smooth. Stamens 3. Ovary glabrous or hairy on top. Caryopsis oblong or linear, usually ventrally sulcate, usually free from lemma and palea, hilum long-linear. x = 7.[25] [more]
Fremontodendron
The flannelbush or flannel bush (Fremontodendron, syn. Fremontia) is a of two species of shrubs from the southwestern United States and northwest Mexico. They are treated within the Sterculiaceae by the most of the authors, in the tribe Fremontodendreae together with the genus Chiranthodendron, but also included in the family Malvaceae (fide APG). [more]
Galpinia
Garrya
Garrya is a genus of about 18 species of in the family Garryaceae, native to North and Central America and the Caribbean. They are commonly referred to as silktassel. [more]
Gentiana
Herbs annual, biennial, or perennial. Rootstock with a fibrous primary root and secondary rootlets, with a stout ± fleshy or woody taproot, or with several linear-cylindric roots from a collar. Stems ascending to erect, striate or angled, in perennial species sometimes both flowering and vegetative. Leaves opposite, rarely whorled, sometimes forming a basal rosette. Inflorescences axillary or terminal, 1 to few-flowered cymes, sometimes in terminal clusters and/or axillary whorls. Flowers (4 or) 5- (or 6-8) -merous. Calyx lobes filiform to ovate, with a prominent midvein. Corolla tubular, salverform, funnelform, obconic, or urceolate, very rarely rotate; tube usually much longer than lobes; plicae between lobes. Stamens inserted on corolla tube; filaments basally ± winged; anthers free or rarely contiguous. Glands 5-10 at ovary base. Pistil sessile or on a long gynophore. Style usually short, linear, less often long and filiform; stigma lobes free or connate, recurved, usually oblong to linear, rarely expanded and rounded. Capsule cylindric to ellipsoid and wingless or narrowly obovoid to obovoid (narrowly ellipsoid in G. winchuanensis) and winged, many seeded. Seeds wingless or winged; seed coat minutely reticulate, rugose, simply areolate, or with complex spongy areolation.[26] [more]
Geranium
Annual or perennial herbs. Leaves usually alternate, stipulate, variously divided. Peduncles (1-) 2-flowered. Flowers often showy, regular, usually 5-merous. Petals alternating with 5 nectiferous glands. Stamens (5-) 10, staminodes occasional. Carpels usually 5, adnate, separating septifragally from the central axis at maturity. Ovary 5-lobed. Fruit schizocarpic, of 5 mericarps which remain attached to an elastically coiling stylar axis upwards; mericarps without apical pits.[27] [more]
Gladiolus
Herbs, perennial, from corms. Stems simple or branched. Leaves 1-9; blade lanceolate to linear, plane or margins and/or midribs variously raised and thickened (then H- or X-shaped in cross section), or evidently terete, midribs and margins much thickened, grooved; grooves 4, narrow, longitudinal. Inflorescences spicate, partly to fully secund or with flowers weakly distichous; bracts green, sometimes flushed grayish purple, unequal, outer usually exceeding inner, acute or inner forked or notched apically. Flowers somewhat fragrant, zygomorphic [actinomorphic]; tepals basally connate into tube, variously colored, usually with contrasting markings comprising nectar guide on outer tepals, usually unequal, dorsal tepal largest, arched to hooded over stamens, outer 3 tepals narrower; perianth tube obliquely funnel-shaped to cylindric; stamens usually unilateral; anthers usually parallel; style usually arching over stamens, dividing into 3 filiform branches, these distally expanded. Capsules usually slightly inflated, oblong to ellipsoid or globose [rarely nearly cylindric], softly cartilaginous. Seeds usually many, broadly winged; rarely few, wingless, globose or angular; seed coat light to dark brown. x = 15.[28] [more]
Gnolia
Heliotropium
Herbs annual or perennial, rarely subshrubs, pubescent or strigose, rarely scabrous. Leaves alternate, less often opposite, sessile or petiolate. Cymes terminal, rarely axillary, unilateral, scorpioid, bracteate or not. Calyx 5-parted. Corolla white or light bluish purple, less often yellow, cylindric or funnelform, strigose outside, glabrous and rarely appressed pubescent inside; throat frequently pubescent; limb 5-parted; lobes orbicular, sometimes linear, margin with folds or undulate. Filaments extremely short; anthers included. Ovary completely or incompletely divided into 4 lobes; ovules 4. Style terminal; stigma conical or ringlike. Fruit nutletlike dry drupes, without evident mesocarp at maturity, endocarp bony, dividing into 4 1-seeded or 2 2-seeded mericarps. Seeds straight or curved, usually with a thin endosperm.[29] [more]
Helleborus
Herbs [subshrubs], perennial, from tough, short rhizomes [rhizomes absent]. Leaves basal and cauline, basal leaf much larger [all leaves cauline], petiolate; cauline leaves alternate. Leaf blade pedately or palmately compound or deeply parted [undivided], lobes narrowly elliptic to oblanceolate or lanceolate, margins sharply toothed [entire]. Inflorescences terminal, 3-4-flowered cymes, to 25 cm or flowers solitary or paired; bracts ±leaflike, divided, not forming involucre. Flowers bisexual, radially symmetric; sepals persistent in fruit [not persistent], 5, yellowish green [white, pink, or purple], plane, ovate to elliptic, 19-30(-50) mm; petals 5-15, distinct, green or brown, funnel-shaped, ± 2-lipped, clawed, 4-8 mm; nectary in center of "funnel"; stamens 30-60; filaments filiform; staminodes absent between stamens and pistils; pistils [2-]3-6[-10], simple, proximally connate [distinct or completely connate]; ovules several per pistil; style present. Fruits follicles [capsules], aggregate, sessile, oblong, sides with prominent transverse veins; beak terminal, straight, 5-15 mm. Seeds usually ± carinate. x = 8.[30] [more]
Hemerocallis
Herbs, perennial, scapose, clump-forming, rhizomatous, from fibrous or fleshy contractile roots often enlarged at ends; rhizomes spreading. Leaves many, basal, sessile, 2-ranked, bases sheathing; blade long-linear, keeled, apex acuminate. Inflorescences 2, in terminal helicoid cyme, or solitary. Flowers mostly diurnal and ephemeral, slightly irregular, showy; tepals 6, connate basally into short, funnelform to campanulate tube, distinct parts imbricate, spreading, inner broader than outer; stamens 6, adnate to throat of perianth tube; filaments curved upward, distinct, unequal; anthers dorsifixed, 2-locular, linear-oblong, dehiscence introrse; ovary superior, green, 3-locular, conic, septal nectaries present; style curved upwards; stigma indistinctly 3-lobed or capitate. Fruits capsular, leathery, dehiscence loculicidal. Seeds rarely produced (sterile) or many. x = 11.[31] [more]
Hippocrepis
Hippocrepis is a of ornamental plants in the family Fabaceae. [more]
Holboellia
Woody climbers, evergreen or deciduous. Monoecious. Stems twining. Winter bud outer scales numerous, seriate. Leaves alternate, usually long petiolate; leaf blade palmately 3--9-foliolate or pinnately 3-foliolate; petiolules usually unequal with terminal one longest; leaflet margins entire. Inflorescences axillary, clustered, corymbiform racemes, rarely elongated racemes; several to many flowered. Sepals 6, greenish white or purple, oblong, slightly fleshy, outer 3 valvate, inner 3 usually smaller. Petals 6, minute, opposite stamens, rudimentarily nectariferous. Male flowers: stamens 6, free; connective apically appendaged or unappendaged. Pistillodes 3, minute. Female flowers: carpels 3, straight. Stigma terminal or oblique, often cleft. Fruit oblong to ellipsoid, fleshy, indehiscent. Seeds embedded in pulp.[32] [more]
Hypoxis
Herbs, perennial, scapose, glabrous or sparsely to densely pubescent, often pilose, pubescence including at least some irregularly stellate trichomes, rhizomatous or cormose. Stems subterranean, usually vertical, fleshy. Leaves grasslike; blade linear to setaceous. Scape usually shorter than leaves. Inflorescences depauperate racemes or umbels, borne singly in leaf axils, bracteate. Flowers: tepals 6, spreading, distinct, often greenish abaxially, yellow adaxially, outer usually ± pilose abaxially; anthers 6, spreading, shortly connate at bases; ovary inferior, usually densely pubescent to pilose, sometimes glabrate; style erect. Fruits capsular, crowned by persistent flower parts throughout maturation. Seeds (5-) 10-50 per capsule, ± globose, hilum and micropyle prominent, surfaces sharply to bluntly muricate or with rounded pebbling, sometimes with iridescent, membranous coat.[33] [more]
Juncus
Herbs, perennial or rarely annual, rhizomatous or cespitose. Culms round or flattened in cross section. Cataphylls often present at culm base. Leaves: sheaths open; blade flat, channeled, ensiform or terete, sometimes septate, margins involute. Inflorescences terminal or pseudoaxillary, monochasia or dichasia, usually with monochasial branches, cymes or 1--many heads in racemes or panicles; bracteoles 2 or absent. Flowers: tepals (4--) 6 in 2 whorls; stamens (2--) 3--6. Capsules 1-locular or 3-locular, septicidal. Seeds many, ellipsoid to ovoid, sometimes tailed.[34] [more]
Juniperus
Shrubs or trees evergreen. Branchlets terete, 3--6 angled, variously oriented, but not in flattened sprays. Leaves opposite in 4 ranks or in whorls of 3. Adult leaves closely appressed to divergent, scalelike to subulate, free portion to ca. 10 mm (to ca. 15 mm in Juniperus communis ) ; abaxial gland visible or not, elongate to hemispheric ( J. ashei ), sometimes exuding white crystalline deposit. Pollen cones with 3--7 pairs or trios of sporophylls, each sporophyll with 2--8 pollen sacs. Seed cones maturing in 1 or 2 years, globose to ovoid and berrylike, 3--20 mm, remaining closed, usually glaucous; scales persistent, 1--3 pairs, peltate, tightly coalesced, thick and fleshy or fibrous to obscurely woody. Seeds 1--3 per scale, round to faceted, wingless; cotyledons 2--6. x = 11.[35] [more]
Kniphofia
Kniphofia (Tritoma, Red hot poker, Torch lily, Poker plant) is a genus of plants in the family that includes 70 or more species native to Africa. Some species have been commercially used horticulturally and are commonly known for their bright, rocket-shaped flowers. [more]
Lampranthus
Lampranthus is a genus of plants in the family . One of the species in this genus is L. roseus, the mini ice plant. Other species in this genus include L, haworthii and L. aberdeen. All Lampranthus species flower between June and August with flower colors including red, orange, peach, yellow and light pink through to magenta and purple. [more]
Lavandula
Plants small shrubs, rarely herbs. Verticillasters 2-10-flowered, in crowded terminal spikes; bracteoles small or absent. Flowers short pedicellate or subsessile. Calyx ovoid-tubular to tubular, slightly dilated in fruit, straight, 13-15-veined, 2-lipped; upper lip entire, protracted into an appendage; lower lip equally (2-) 4-toothed, teeth narrower than those of upper lip. Corolla blue or purple; tube exserted, throat ± dilated; limb 2-lipped, upper lip 2-lobed, lower 3-lobed. Stamens 4, included, anterior 2 longer; anther cells apically confluent. Style inserted at ovary base, apex 2-cleft, lobes flattened, ovate, connate. Nutlets smooth, shiny, each with a basal-dorsal areole.[36] [more]
Leucadendron
Leucadendron is a genus of about 80 species of in the family Proteaceae, endemic to South Africa, where they are a prominent part of the fynbos vegetation. [more]
Leucosidea
Leucosidea sericea, commonly known as Oldwood, is a species of evergreen tree and large shrub native to the regions of Southern Africa. It is the sole species of genus Leucosidea. [more]
Lilium
Herbs, perennial, bulbose. Bulbs whitish, rarely yellowish or purplish, often stained brown, erect and ovoid (hereafter ovoid), irregular and chunky (chunky), slanted in ground and ± elongate (subrhizomatous), or horizontally elongate (rhizomatous), sometimes branching if rhizomatous, rarely if not, 1.411.7 × 1.319 cm, 0.13 times taller than long, annual growth usually obscure; scales (modified leaves) numerous, fleshy and starchy, usually densely covering rhizomes, rarely bearing leaf blades known as basal leaves or their abscission scars, often notched or segmented, longest 0.811.9 cm; roots on each bulb either contractile and concentrically wrinkled and thick (to 5 mm), or for nutrition and thinner, fibrous. Stems erect, green, sometimes purple, rarely glaucous, to 3.1 m, ± glabrous, often with adventitious stem roots above bulb. Buds usually rounded in cross section, sometimes ± triangular. Leaves numerous, usually ± evenly distributed along stem, rarely concentrated proximally, scattered or more commonly in 112(24) whorls with some scattered at stem base and apex, 320(40) leaves per whorl, sessile, drooping at tips to ascending, 1.729 × 0.25.6 cm, 1.634 times longer than wide; blade green and somewhat lighter abaxially, rarely paler, linear, lanceolate, elliptic, or obovate, sometimes oblanceolate, especially in proximal leaves, often somewhat lanceolate in distal leaves, margins entire, undulate or not, usually glabrous and smooth or occasionally slightly papillose, sometimes roughened abaxially by ± deltoid epidermal spicules, apex acute to obtuse or rarely acuminate; principal veins usually 3, usually glabrous and smooth abaxially, sometimes with ± deltoid epidermal spicules, rarely impressed adaxially. Inflorescences maturing acropetally, terminal, racemose or umbellate (in small plants), usually open, bracteate, 125 (45) -flowered; bracts usually 12 per flower, often with one lanceolate and very wide and the other linear or filiferous. Flowers pendent, nodding, horizontal, ascending, or erect, radially or slightly bilaterally symmetric, fragrant or not; perianth campanulate, funnelform, or with sepals and petals strongly reflexed in form of a Turks-cap; sepals and petals usually differentiated, sometimes indistinctly so, recurved or reflexed, distinct, orange, red, yellow, pink, or white, usually with adaxial magenta or maroon spots concentrated in proximal 1/22/3, ± lanceolate and narrowed or rarely clawed, glabrous (pubescent strip at base in L. lancifolium), nectaries present on each but often more developed on sepals, basal, green, usually hidden but occasionally exposed and forming visible green star at adaxial base of perianth; sepals 3, occasionally ridged abaxially, 3.112 × 0.62.6 cm, apex usually acute; petals 3, ridged abaxially, with 2 adaxial longitudinal median rounded ridges, 311.2 × 0.63.4 cm, apex usually acute, often more widely than sepal apex; stamens 6, opposite sepals and petals, distinct, included to strongly exserted; filaments ± parallel to style or spreading, diverging to 31° from flower axis, color variable but usually pale green or nearly translucent; anthers versatile, color variable, usually purplish, becoming darker, oblong, 0.32.6 cm; pollen cream, yellow, peach, tan, orange, rust, or brown, usually becoming lighter; pistil compound, 3-lobed, 3-locular, oblong, 2.110.5 cm; ovary superior, 0.83.5 cm, axile placentas 6, ovules as many as seeds, a few developing without embryos; style initially parallel to flower axis, usually elongating and curving toward periphery, usually pale green, round in cross section; stigma 3-lobed, hollow in older flowers; pedicel not articulate, 0.832 cm. Fruits erect, green maturing to brown, capsular, 3-valved, not strongly winged, ± oblong-obovate, 1.57.7 × 0.83.3 cm, 1.14.8 times longer than wide, base constricted, dehiscence loculicidal. Seeds 67330, light brown with darker ovate embryo in center, 6-ranked, flattened into 60° wedge, verrucose. x = 12.[37] [more]
Limnanthes
Limnanthes is a of annual herbaceous plants commonly known as the meadowfoams. The nine species are all native to the coastal regions of western North America, where they typically grown in marshy habitats, such as the margins of vernal pools. [more]
Linum
Erect herbs, rarely shrubby below. Leaves alternate, linear to lanceolate, sessile. Flowers ebracteate, 5-merous, in cymes or racemes. Sepals entire. Petals linear to lanceolate or obovate. Glands 5, adnate to the staminal tube, opposite the petals. Stamens 5, connate at the base; staminodes present or absent. Carpels 5; ovary 10-locular due to secondary septa; styles 5, free or connate at the base; stigma linear to capitate. Fruit a septicidal capsule.[38] [more]
Lupinus
Herbs or shrubs. Leaf generally palmately multifoliolate; stipules adnate.. Inflorescence a terminal raceme. Bract caducous, bracteoles mostly persistent. Calyx 2-lipped. Vexillum orbicular or broadly ovate. Wing petals connate at the apex. Keel beaked. Stamens monadelphous, anthers alternately long and short. Ovary sessile, 2-many ovuled, style incurved, glabrous or with a ring of hairs beneath the terminal stigma. Fruit compressed, 2-valved, often constricted between the seeds. Seed estrophiolate.[39] [more]
Macrozamia
Macrozamia is a genus of 38-40 species of , in the family Zamiaceae, endemic to Australia. The majority of the species occur in eastern Australia in southeast Queensland and New South Wales, with one species in the Macdonnell Ranges of Northern Territory and three in southern Western Australia. The common name Burrawang, originally referring to M. communis in the Daruk Australian Aboriginal language, is often used for all the species in the genus. [more]
Magnolia
Trees or shrubs, deciduous or evergreen. Pith homogeneous or diaphragmed. Leaves distinctly alternate or sometimes crowded in terminal whorl-like clusters; stipules early deciduous, free or adnate to and proximal on petiole. Leaf blade: base deeply cordate or auriculate or cuneate to abruptly narrowed or rounded, margins entire, apex obtuse or acute to acuminate; surfaces abaxially chalky white or green to glaucous, pubescent or glabrous. Flowers protogynous, appearing with or before leaves; tepals 9-15, petaloid, usually spreading, creamy white, rarely greenish, yellow, or orange-yellow, outermost tepals sepaloid, sometimes strongly reflexed, greenish; stamens on elongate torus, early deciduous; filaments white or purple, very short; anthers introrse or latrorse. Follicles persistent, coalescent, forming conelike aggregate, abaxially dehiscent. Seeds with red, pink, or orange oily aril, extruded from follicles and suspended by funiculi. x =19.[40] [more]
Mahonia
Evergreen, unarmed shrubs, similar to the genus Berberis L., but leaves pinnately compound and inflorescence mostly a fascicle of dense spike-like racemes.[41] [more]
Malephora
Shrubs or subshrubs, perennial, succulent. Roots fibrous. Stems erect to prostrate, rooting at older nodes. Leaves cauline, opposite, sessile; stipules absent; blade slightly connate basally, triangular to cylindric, smooth, usually glaucous. Inflorescences axillary or terminal, flowers solitary or in cymes, pedicellate; bracts absent. Flowers showy, tubular, to 5 cm diam.; calyx lobes 4-6, tapered, unequal; petals (including petaloid staminodia) 40-65, distinct, yellow, pink, orange, or reddish to purple; stamens 150, erect, bases often hairy; nectary often present; pistil 8-12-carpellate; ovary mostly inferior, flat apically, 8-12-loculed, each locule with basal pocket containing 1-2 or no seeds; placentation parietal; style absent; stigmas 8-12, plumose. Fruits capsules, persistent, dehiscence loculicidal, not separating into segments; locule lids present, winged. Seeds 75, lenticular, rough; tubercles in rows.[42] [more]
Mandevilla
Lianas usually glabrous, with latex. Leaves opposite; stipules interpetiolar, reduced to many linear segments. Racemes axillary, few flowered. Flowers large. Calyx deeply divided, with many basal glands inside. Corolla funnelform; tube narrow, more than 2 cm, limb campanulate; faucal scales absent; lobes overlapping to right. Stamens inserted and included in widened part of corolla tube; filaments short; anthers oblong, adherent at middle to pistil head, cells obtusely caudate; disc 5-cleft. Ovaries free, glabrous; ovules numerous. Style glabrous; pistil head with a 2-cleft mucro. Follicles long, slender. Seeds narrowly oblong, not beaked, comose.[43] [more]
Manglietia
Manglietia is a genus of in family Magnoliaceae. It contains the following species (but this list may be incomplete): [more]
Marrubium
Marrubium (horehound or hoarhound) is a genus of about 40 species of in the family Lamiaceae, native to temperate regions of Europe and Asia. [more]
Michelia
Michelia is a of flowering plants belonging to the Magnolia family (Magnoliaceae). The genus includes about 50 species of evergreen trees and shrubs, native to tropical and subtropical south and southeast Asia (Indomalaya), including southern China. [more]
Narcissus
Herbs perennial, scapose, from ovoid, tunicate bulbs. Leaves (1-) several; blade linear to ligulate, flat to semiterete, fleshy. Inflorescences umbellate in clusters of 2-20, or solitary, spathaceous; spathe 1-valved, enclosing buds, membranous or papery. Flowers pedicellate or sessile, erect or declinate, often fragrant; tepals 6, connate proximally, distinct and reflexed to ascending distally, yellow and/or white; perianth tube surmounted by a cupular to trumpetlike corona with margins often frilled; stamens 6, epitepalous, often of 2 lengths; filaments separate from corona; anthers basifixed; ovary inferior, 3-locular; style often exserted; stigma minutely 3-lobed. Fruits capsular, 3-locular, papery to leathery, dehiscence loculidical. Seeds numerous, subglobose, often with elaiosomes; testa black. x = 7, 11.[44] [more]
Nuxia
Nuxia is a genus of in family Scrophulariaceae. It contains the following species (but this list may be incomplete): [more]
Oplismenus
Perennials or annuals. Culms trailing, ascending from a decumbent base, the internodes mostly with a villous line adaxial to the subtending leaf. Leaf blades ovate, lanceolate or linear-lanceolate, often with cross veins; ligule a ciliate membrane. Inflorescence composed of several unilateral racemes spaced along a central axis; racemes elongate or reduced to fascicles of a few spikelets, spikelets usually paired on short pedicels, the lowermost often reduced. Spikelets lanceolate to oblong, weakly dorsally or laterally compressed, florets 2; glumes subequal, 1/2-3/4 spikelet length, often pilose, the lower or both tipped by a viscid awn; lower lemma equaling the spikelet, acute to shortly awned, its palea absent or much reduced; upper lemma papery or subcoriaceous, smooth and shiny, acute, indistinctly crested. x = 9.[45] [more]
Oxalis
Mostly bulbous herbs with acidic juice. Aerial stem reduced, creeping or rhizomatous. Leaves digitately compound, showing sleep movements. Flowers regular. Stamens monadelphous at the base; filaments 5 long and 5 short, alternating with one another. Ovary 5 locular with one or more ovules in each locule. Often heterostylous (in some European species) . Fruit capsular. Seeds with an elastic testa.[46] [more]
Paeonia
Morphological characters and geographical distribution are the same as those of the family.[47] [more]
Pelargonium
Perennial with rarely entire leaves. Flowers showy, umbellate, irregular. Posticous sepal prolonged into a nectiferous spur. Fruit beaked.[48] [more]
Phacelia
Phacelia, the phacelias or scorpionweeds, is a large genus. It contains about 200 species of herbs, native of Western North America (the most), Eastern USA and South America. [more]
Plantago
Annual or perennial, or mostly acaulescent herbs, sometimes an undershrub. Leaves usually all radical, sometimes cauline. Flowers hermaphrodite, dimorphic, polygamous, usually in many-flowered cylindrical spikes, rarely capitate. Calyx lobes subequal or 2 large and 2 small. Corolla tube cylindrical, sometimes contracted at the throat, usually equalling the calyx, sometimes exserted, lobes 4, equal, patent. Stamens 4, inserted at or above the middle of the corolla tube, often exserted. Ovary 2-locular or 3-4-locular with false free septa; locules with 1 to many ovules. Capsule membranous, circumscissile. Seeds usually peltate, testa thin, mucilaginous, albumen fleshy.[49] [more]
Polypodium
A genus in the Kingdom Animalia. [more]
Polystichum
Plants terrestrial. Stems decumbent to erect, stolons absent. Leaves monomorphic (dimorphic in P. acrostichoides ), evergreen. Petiole 1/9--1 times length of blade, bases swollen or not; vascular bundles more than 3, arranged in an arc, ± round in cross section. Blade linear-lanceolate to broadly lanceolate, 1--3-pinnate, gradually reduced distally to pinnatifid apex, somewhat leathery to leathery. Pinnae not articulate to rachis, segment or pinna margins spinulose-toothed (except P. lemmonii ) ; proximal pinnae (several pairs) usually gradually reduced, sessile to short-petiolulate, bases usually inequilateral with acroscopic lobe; costae adaxially grooved, grooves continuous from rachis to costae; indument of linear to lanceolate scales on costae and sometimes between veins abaxially (microscales), ± glabrous or similarly scaly adaxially (scales forming loosely tangled network over blade and sori in P. dudleyi ). Veins free, forked, rarely ( P. imbricans ) anastomosing. Sori in 1 row (to several) between midrib and margins, round (confluent, covering abaxial surface in P. acrostichoides ) ; indusia peltate, persistent or caducous [absent]. Spores yellow or brownish to black, with inflated folds. x = 41.[50] [more]
Pyrrhopappus
Annuals or perennials, 5-100+ cm; taprooted or rhizomatous (roots producing tuberiform swellings in P. grandiflorus). Stems usually 1, sometimes 2-5+, erect, unbranched or branched proximally and/or distally, glabrous or pilosulous. Leaves basal or basal and cauline; basal ± petiolate, distal usually sessile; blades oblong, elliptic, or ovate to lanceolate or linear, margins entire or dentate to pinnately lobed (faces usually glabrous, sometimes pilosulous near margins). Heads borne singly or in loose, corymbiform arrays. Peduncles not inflated distally, sometimes bracteate. Calyculi of 3-13+, deltate to subulate or filiform bractlets. Involucres cylindric, 4-5[-8+] mm diam. Phyllaries 8-21+ in ± 2 series (reflexed in fruit), linear, equal, margins often scarious, apices acute (often thickened or bearing keel-like flaps near tips). Receptacles ± convex, pitted, glabrous, epaleate. Florets (20-) 30-150+; corollas yellow to whitish. Cypselae reddish brown to stramineous, bodies ± fusiform, beaks (± concolorous with bodies) ± filiform, fragile, grooves (or broad ribs) 5, faces transversely rugulose, glabrous; pappi (borne on discs at tips of beaks) persistent, double: outer coroniform (of whitish, relatively short, spreading, sometimes curly, hairs), inner of 80-120+, rufous to stramineous, subequal, barbellulate bristles in 2-3+ series. x = 6.[51] [more]
Rehmannia
Herbs, perennial. Rhizomes present. Stems erect, simple or branched from base. Basal leaves rosulate, stem leaves alternate and petiolate; leaf blade varying in shape, often hairy, margin toothed or lobed. Inflorescences sometimes scapose. Flowers axillary and solitary or in terminal racemes, pedicellate. Bracteoles present or absent. Calyx 5(-7) -lobed. Corolla purple-red or yellow, tubular; tube slightly curved or straight, dorsiventrally compressed, with 2 plaits from tube base to throat; limb 2-lipped, 5-lobed. Stamens 4, didynamous, rarely 5 and 1 smaller than other 4, included; anthers coherent in pairs, locules fertile. Ovary base with a disc, 2-loculed, rarely 1-loculed; ovules numerous. Stigma 2-lamellate. Capsule with persistent calyx, loculicidal. Seeds minute; seed coat reticulate.[52] [more]
Rhodocoma
Salix
Trees or shrubs deciduous, rarely evergreen (if shrubs, then erect, ascending procumbent, creeping, or cushion-shaped) ; pith terete. Branches terete. Terminal bud usually absent; buds with single scale. Leaves alternate, rarely subopposite or opposite; stipules small, free, deciduous or persistent, developed mainly on vigorous branchlets; petiole short; leaf blade variously shaped, often long and narrow. Flowering precocious, coetaneous, or serotinous; catkins upright or spreading, rarely pendulous; bracts entire, persistent or caducous. Flowers entomophilous or anemophilous, each with 1 or 2 glands: 1 abaxial (dorsal) or absent and 1 adaxial (ventral), i.e., abaxial gland between bract and stipe, adaxial gland between stipe and rachis. Male flower: stamens 2-many; filaments free or partly to completely connate, usually exceeding bracts; anthers 2-loculed (rarely 4-loculed if filaments connate), opening lengthwise. Female flower: ovary 2-loculed, sessile or stipitate; style 1, short, slender, or absent, entire or 2-cleft; stigmas 1 or 2, lobed or entire. Capsule 2-valved. Seeds mostly green or gray-green, small, surrounded by fine hairs.[53] [more]
Saxifraga
Herbs perennial, rarely annual or biennial. Stem cespitose or simple. Leaves both basal and cauline, petiolate or not; leaf blade simple, entire, margin dentate or lobate; cauline leaves usually alternate, rarely opposite. Inflorescence a solitary flower or few- to many-flowered cyme, bracteate. Flowers usually bisexual, sometimes unisexual, actinomorphic, rarely zygomorphic; receptacle cyathiform or saucer-shaped. Sepals (4 or) 5(or 7 or 8) . Petals (4 or) 5, yellow, orange, white, or red to purple, callose or not, distinctly veined, margin usually entire. Stamens (8 or) 10; filaments subulate or clavate. Carpels 2, usually connate at least in placental region; ovary superior to inferior, usually 2-loculed; placentation usually axile; ovules many; integuments 1 or 2; nectary disc sometimes well developed, annular or semiannular. Fruit a 2-valved capsule. Seeds many.[54] [more]
Silybum
Annuals or biennials, taprooted, 15-300 cm, herbage glabrous, puberulent, or slightly tomentose, spiny. Stems erect, usually simple. Leaves basal and cauline; petiolate (basal and proximal cauline) or sessile (distal cauline) ; blades adaxially variegated, margins dentate and often coarsely pinnately lobed, teeth and lobes spine-tipped, glabrous or puberulent. Heads discoid, borne singly, terminal and in distal axils. ( Peduncles with reduced leaflike bracts.) Involucres ovoid to spheric, 15-60 mm diam. Phyllaries many in 4-6 series. unequal, outer and mid with appressed bases and spreading, lanceolate to ovate, spiny-fringed, terminal appendages, at least mid spine-tipped, innermost with erect, flat, entire, spineless apices. Receptacles flat, epaleate, covered with whitish bristles. Florets 25-100+; corollas pink to purple, tubes slender, distally bent, abruptly expanded into short throats, lobes linear; stamen filaments connate. anther bases sharply short-tailed, anther appendages oblong; style branches: fused portions with slightly swollen subterminal nodes, distally cylindric, distinct portions minute. Cypselae ovoid, slightly compressed, not ribbed, apices with smooth, entire rims, glabrous, basal attachment scars slightly angled; pappi falling in rings, outer of many minutely barbed, basally connate, subulate scales, inner of minute smooth bristles. x = 17.[55] [more]
Sorbus
Trees or shrubs, usually deciduous. Winter buds usually rather large, ovoid, conical, or spindle-shaped, sometimes viscid; scales imbricate, several, glabrous or pubescent. Leaves alternate, membranous or herbaceous; stipules caducous, simple or pinnately compound, plicate or rarely convolute in bud; leaf blade usually serrate, sometimes nearly entire, venation craspedodromous or camptodromous, glabrous or pubescent. Inflorescences compound, rarely simple corymbs or panicles. Hypanthium campanulate, rarely obconical or urceolate. Sepals 5, ovate or triangular, glabrous, pubescent, or tomentose, sometimes glandular along margin. Petals 5, glabrous or pubescent, base clawed or not. Stamens 15-25(-44) in 2 or 3 whorls, unequal in length; anthers ovoid or subglobose. Carpels 2-5, partly or wholly adnate to hypanthium; ovary semi-inferior to inferior, 2-5-(-7) loculed, with 2 or 3(or 4) ovules per locule, one usually abortive; styles 2-5, free or partially connate, glabrous or pubescent. Fruit a pome, white, yellow, pink, or brown to orange or red, ovoid or globose to ellipsoid or oblong, usually small, glabrous or pubescent, laevigate or with small lenticels, apically with sepals persistent or caducous leaving an annular scar, with 2-5(-7) locules, each with 1 or 2 exendospermous seeds; seeds several, with thin perisperm and endosperm enclosing embryo with compressed cotyledons.Trees or shrubs, usually deciduous. Winter buds usually rather large, ovoid, conical, or spindle-shaped, sometimes viscid; scales imbricate, several, glabrous or pubescent. Leaves alternate, membranous or herbaceous; stipules caducous, simple or pinnately compound, plicate or rarely convolute in bud; leaf blade usually serrate, sometimes nearly entire, venation craspedodromous or camptodromous, glabrous or pubescent. Inflorescences compound, rarely simple corymbs or panicles. Hypanthium campanulate, rarely obconical or urceolate. Sepals 5, ovate or triangular, glabrous, pubescent, or tomentose, sometimes glandular along margin. Petals 5, glabrous or pubescent, base clawed or not. Stamens 15-25(-44) in 2 or 3 whorls, unequal in length; anthers ovoid or subglobose. Carpels 2-5, partly or wholly adnate to hypanthium; ovary semi-inferior to inferior, 2-5-(-7) loculed, with 2 or 3(or 4) ovules per locule, one usually abortive; styles 2-5, free or partially connate, glabrous or pubescent. Fruit a pome, white, yellow, pink, or brown to orange or red, ovoid or globose to ellipsoid or oblong, usually small, glabrous or pubescent, laevigate or with small lenticels, apically with sepals persistent or caducous leaving an annular scar, with 2-5(-7) locules, each with 1 or 2 exendospermous seeds; seeds several, with thin perisperm and endosperm enclosing embryo with compressed cotyledons.[56] [more]
Styrax
Trees or shrubs, stellate pubescent or scaly, rarely glabrous. Leaves usually alternate. Inflorescences axillary or terminal, racemes, panicles, or cymes, sometimes 1-flowered or several-flowered in a fascicle; bracteoles small, early deciduous. Flowers bisexual. Calyx cup-shaped, 5-toothed, rarely 2--6-lobed or truncate. Corolla campanulate; lobes 5(--7), imbricate or valvate. Stamens (8--) 10(--13), equal or rarely unequal in length; filaments flattened, free, sometimes basally adnate to corolla; anthers oblong. Ovary superior, 3-locular when young, becoming 1-locular; ovules 1--4 per locule, placentation parietal. Style subulate or filiform; stigma capitate or 3-lobed. Drupe
