Overview
Cedrus (common name Cedar) is a genus of coniferous trees in the plant family Pinaceae. They are native to the mountains of the western Himalaya and the Mediterranean region, occurring at altitudes of 1,500?3,200 m in the Himalaya and 1,000?2,200 m in the Mediterranean.1]
Description
Cedars are trees up to 30?40 m (occasionally 60 m) tall with spicy-resinous scented wood, thick ridged or square-cracked bark, and broad, level branches. The shoots are dimorphic, with long shoots, which form the framewo rk of the branches, and short shoots, which carry most of the leaves. The leaves are evergreen and needle-like, 8?60 mm long, arranged in an open spiral phyllotaxis on long shoots, and in dense spiral clusters of 15?45 together on short shoots; they vary from bright grass-green to dark green to strongly glaucous pale blue-green, depending on the thickness of the white wax layer which protects the leaves from desiccation. The seed cones are barrel-shaped, 6?12 cm long and 3?8 cm broad, green maturing grey-brown, and, as in Abies, disintegrate at maturity to release the winged seeds. The seeds are 10?15 mm long, with a 20?30 mm wing; as in Abies, the seeds have 2?3 resin blisters, containing an unpleasant-tasting resin, thought to be a defence against squirrel predation. Cone maturation takes one year, with pollination in autumn and the seeds maturing the same time a year later. The pollen cones are slender ovoid, 3?8 cm long, produced in late summer and shedding pollen in autumn.[1][2]
Taxonomy
Cedars share a very similar cone structure with the firs (Abies) and were traditionally thought to be most closely related to them, but molecular evidence supports a basal position in the family.[3][4]
There are five taxa of Cedrus, assigned according to taxonomic opinion to between one and four different species.[1][5][6][7][8][9][10][11][12][13][14]:
- Deodar or Deodar Cedar, C. deodara (syn. C. libani subsp. deodara). Western Himalaya. Leaves bright green to pale glaucous green, 25?60 mm; cones with slightly ridged scales.
- Lebanon Cedar or Cedar of Lebanon C. libani. Cones with smooth scales; two (or up to four) subspecies:
- Lebanon Cedar C. libani subsp. libani. Mountains of Lebanon, western Syria and south-central Turkey. Leaves dark green to glaucous blue-green, 10?25 mm.
- Turkish Cedar C. libani subsp. stenocoma. Mountains of southwest Turkey. Leaves glaucous blue-green, 8?25 mm.
- Cyprus Cedar C. brevifolia (syn. C. libani subsp. brevifolia, C. libani var. brevifolia). Mountains of Cyprus. Leaves glaucous blue-green, 8?20 mm.
- Atlas Cedar C. atlantica (syn. C. libani subsp. atlantica). Atlas mountains in Morocco & Algeria. Leaves dark green to glaucous blue-green, 10?25 mm.
Ecology
Cedars are adapted to mountainous climates; in the Mediterranean they receive winter precipitation, mainly as snow, and summer drought, while in the western Himalaya, they receive primarily summer monsoon rainfall.[1]
Cedars are used as food plants by the larvae of some Lepidoptera species including Pine Processionary and Turnip Moth (recorded on Deodar Cedar).
Uses
Cedars are very popular ornamental trees, widely used in horticulture in temperate climates where winter temperatures do not fall below about -25 ?C. The Turkish Cedar is slightly hardier, to -30 ?C or just below. Extensive mortality of planted specimens can occur in severe winters where temperatures do drop lower.[15] Areas with successful long-term cultivation include the entire Mediterranean region, western Europe north to the British Isles, southern Australia and New Zealand, and southern and western North America.
Cedar wood and cedar oil are known to be a natural repellent to moths,[16] hence cedar is a popular lining for modern-day cedar chests and closets in which woolens are stored. This specific use of cedar is mentioned in The Iliad (Book 24), referring to the cedar-roofed or lined storage chamber where Priam goes to fetch treasures to be used as ransom. Cedar is also commonly used to make shoe trees as it can absorb moisture and de-odorise.
Timber of trees with similar names such as Western Red Cedar is frequently confused with genuine cedar.
The Cedar of Lebanon and to a lesser extent the Deodar have local cultural importance.
Etymology
Both the Latin words cedrus and the generic name cedrus are derived from the Greek 'kedros'. Ancient Greek and Latin used the same word, kedros and cedrus respectively, for different species of plants now classified in the genera Cedrus and Juniperus (juniper). Species of both genera are native to the area where Greek language and culture originated, though as the word "kedros" does not seem to be derived from any of the languages of the Middle East, it has been suggested the word may originally have applied to Greek species of juniper and was later adopted for species now classified in the genus Cedrus because of the similarity of their aromatic woods.[17] The name was similarly applied to citron and the word citrus is derived from the same root.[18] However, as a loan word in English, cedar had become fixed to its biblical sense of Cedrus by the time of its first recorded usage in AD 1000.[19]
The name "cedar" has more recently (since about 1700[19]) been applied to many other trees (such as Western redcedar; in some cases the botanical name alludes to this usage, such as the genus Calocedrus (meaning "beautiful cedar"), also known as Incense-cedar). Such usage is regarded by some authorities [20] as a misapplication of the name to be discouraged.
See also
- Lebanon Cedar
- Cedars of God
Ecology
Cedars are adapted to mountainous climates; in the Mediterranean they receive winter precipitation, mainly as snow, and summer drought, while in the western Himalaya, they receive primarily summer monsoon rainfall.[1]
Cedars are used as food plants by the larvae of some Lepidoptera species including Pine Processionary and Turnip Moth (recorded on Deodar Cedar).
Uses
Cedars are very popular ornamental trees, widely used in horticulture in temperate climates where winter temperatures do not fall below about -25 ?C. The Turkish Cedar is slightly hardier, to -30 ?C or just below. Extensive mortality of planted specimens can occur in severe winters where temperatures do drop lower.[15] Areas with successful long-term cultivation include the entire Mediterranean region, western Europe north to the British Isles, southern Australia and New Zealand, and southern and western North America.
Cedar wood and cedar oil are known to be a natural repellent to moths,[16] hence cedar is a popular lining for modern-day cedar chests and closets in which woolens are stored. This specific use of cedar is mentioned in The Iliad (Book 24), referring to the cedar-roofed or lined storage chamber where Priam goes to fetch treasures to be used as ransom. Cedar is also commonly used to make shoe trees as it can absorb moisture and de-odorise.
Timber of trees with similar names such as Western Red Cedar is frequently confused with genuine cedar.
The Cedar of Lebanon and to a lesser extent the Deodar have local cultural importance.
Etymology
Both the Latin words cedrus and the generic name cedrus are derived from the Greek 'kedros'. Ancient Greek and Latin used the same word, kedros and cedrus respectively, for different species of plants now classified in the genera Cedrus and Juniperus (juniper). Species of both genera are native to the area where Greek language and culture originated, though as the word "kedros" does not seem to be derived from any of the languages of the Middle East, it has been suggested the word may originally have applied to Greek species of juniper and was later adopted for species now classified in the genus Cedrus because of the similarity of their aromatic woods.[17] The name was similarly applied to citron and the word citrus is derived from the same root.[18] However, as a loan word in English, cedar had become fixed to its biblical sense of Cedrus by the time of its first recorded usage in AD 1000.[19]
The name "cedar" has more recently (since about 1700[19]) been applied to many other trees (such as Western redcedar; in some cases the botanical name alludes to this usage, such as the genus Calocedrus (meaning "beautiful cedar"), also known as Incense-cedar). Such usage is regarded by some authorities [20] as a misapplication of the name to be discouraged.
See also
- Lebanon Cedar
- Cedars of God
References
- ^ a b c d Farjon, A. (1990). Pinaceae. Drawings and Descriptions of the Genera. Koeltz Scientific Books ISBN 3-87429-298-3.
- ^ Frankis, M. & Lauria, F. (1994). The maturation and dispersal of cedar cones and seeds. International Dendrology Society Yearbook 1993: 43?46.
- ^ Liston A., D.S. Gernandt, T.F. Vining, C.S. Campbell, D. Pi?ero. 2003. Molecular Phylogeny of Pinaceae and Pinus. In Mill, R.R. (ed.): Proceedings of the 4th Conifer Congress. Acta Hort 615: Pp. 107-114.
- ^ Wang, X.-Q., Tank, D. C. and Sang, T. (2000): Phylogeny and Divergence Times in Pinaceae: Evidence from Three Genomes. Molecular Biology and Evolution 17:773-781. Available online
- ^ Gymnosperm database Cedrus.
- ^ NCBI Taxonomy Browser Cedrus.
- ^ Flora of China vol. 4
- ^ Qiao, C.-Y., Jin-Hua Ran, Yan Li and Xiao-Quan Wang (2007): Phylogeny and Biogeography of Cedrus (Pinaceae) Inferred from Sequences of Seven Paternal Chloroplast and Maternal Mitochondrial DNA Regions. Annals of Botany 100(3):573-580. Available online
- ^ Farjon, A. (2008). A Natural History of Conifers. Timber Press ISBN 0-88192-869-0.
- ^ Christou, K. A. (1991). The genetic and taxonomic status of Cyprus Cedar, Cedrus brevifolia (Hook.) Henry. Mediterranean Agronomic Institute of Chania, Greece.
- ^ GRIN Taxonomy for Plants Cedrus.
- ^ G?ner, A., ?zhatay, N., Ekim, T., & Baser, K. H. C. (ed.). 2000. Flora of Turkey and the East Aegean Islands 11 (Supplement 2): 5?6. Edinburgh University Press. ISBN 0-7486-1409-5
- ^ Eckenwalder, J. E. (2009). Conifers of the World: The Complete Reference. Timber Press ISBN 0-88192-974-3.
- ^ Sell, P. D. (1990). Some new combinations in the British Flora. Watsonia 18: 92.
- ^ ?dum, S. (1985). Report on frost damage to trees in Denmark after the severe 1981/82 and 1984/85 winters. H?rsholm Arboretum, Denmark.
- ^ Cedarwood oils
- ^ Meiggs, R. 1982. Trees and Timber in the Ancient Mediterranean World.
- ^ Andrews, A. C. 1961. Acclimatization of citrus fruits in the Mediterranean region. Agricultural History 35: 35?46.
- ^ a b Oxford English Dictionary.
- ^ Kelsey, H. P., & Dayton, W. A. (1942). Standardized Plant Names, second edition. American Joint Committee on Horticultural Nomenclature. Horace McFarland Company, Harrisburg, Pennsylvania.
Taxonomy
The Genus Cedrus is further organized into finer groupings including:
- Species: ZipcodeZoo has pages for 168 species, subspecies, varieties, forms, and cultivars in the Genus Cedrus: C. alternifolia · C. argentea · C. atlantica (Weeping Blue Atlas Cedar) · C. atlantica subsp. douglasii · C. atlantica 'Argentea Fastigiata' · C. atlantica 'Argentea' (Silver Atlas Cedar) · C. atlantica 'Aurea' (Golden Atlas Cedar) · C. atlantica 'Beacon Hill' · C. atlantica 'Blue Cascade' (Blue Cascade Weeping Atlas Cedar) · C. atlantica 'Cheltenham' (Atlas Cedar) · C. atlantica 'Fastigiata' (Blue Atlas Cedar) · C. atlantica 'Glauca Pendula' · C. atlantica 'Hillier's Hb' (Atlas Cedar) · C. atlantica 'Horstmann' (Horstmann Blue Atlas Cedar) · C. atlantica 'Pendula' · C. atlantica 'Sahara Frost' · C. atlantica 'Sahara Ice' (Blue Atlas Cedar) · C. atlantica 'Saphir Nymph' · C. atlantica 'Glauca' (Blue Atlas Cedar) · C. atlantica Glauca Group · C. atlanticus · C. brevifolia · C. brevifolia 'Epstein' · C. brevifolia 'Epsteinianum' (Cyprus Cedar) · C. brevifolia 'Hillier Compact' · C. brevifolia 'Horizon' · C. brevifolia 'Kenwith' · C. brevifolia 'Trevoron' · C. brevifolia (Hook.f.) A.Henry 'Compacta' · C. brevifolia (Hook.f.) A.Henry 'Horizontalis' · C. brevifolia (Hook.f.) A.Henry 'Nana' · C. deodar var. Kashmir (Kashmir Deodar Cedar) · C. deodara (California Christmas Tree) · C. deodara 'Albospica' (Deodar Cedar) · C. deodara 'Argentea' · C. deodara 'Aurea Pendula' (Deodar Cedar) · C. deodara 'Aurea' (Golden Deodar Cedar) · C. deodara 'Blue Dwarf' · C. deodara 'Blue Mountain Broom' · C. deodara 'Blue Snake' (Contorted Deodar Cedar) · C. deodara 'Blue Surprise' · C. deodara 'Blue Triumph' · C. deodara 'Blue Velvet' (Deodar Cedar) · C. deodara 'Bush's Electra' · C. deodara 'Cream Puff' (Deodar Cedar) · C. deodara 'Dawn Mist' · C. deodara 'Deep Cove' (Deodar Cedar) · C. deodara 'Descanso Dwarf' (Deodar Cedar) · C. deodara 'Devinely Blue' (Deodar Cedar) · C. deodara 'Divinely Blue' (Divinely Blue Deodar Cedar) · C. deodara 'Eisregen' · C. deodara 'Electric Blue' (Deodar Cedar) · C. deodara 'Fastigiata' · C. deodara 'Feelin' Blue' (Feelin' Blue Deodar Cedar) · C. deodara 'Feeling Blue' · C. deodara 'Glacier Blue' (Deodar Cedar) · C. deodara 'Gold Cascade' (Deodar Cedar) · C. deodara 'Gold Cone' (Deodar Cedar) · C. deodara 'Gold Gowa' · C. deodara 'Gold Mound' · C. deodara 'Golden Horizon' (Deodar Cedar) · C. deodara 'Golden Jubilee' · C. deodara 'Inversa Pendula' · C. deodara 'Karl Fuchs' · C. deodara 'Kashmir' (Deodar Cedar) · C. deodara 'Klondyke' · C. deodara 'Lime Glow' · C. deodara 'Limeglow' · C. deodara 'MacPenny's Seedling' · C. deodara 'Mayford Grey' · C. deodara 'Miles High' · C. deodara 'Mountain Beauty' (Deodar Cedar) · C. deodara 'Nana Aurea' · C. deodara 'Nana' · C. deodara 'Nivea' · C. deodara 'Pendula' · C. deodara 'Polar Winter' · C. deodara 'Prostrata' (Weeping Deodar Cedar) · C. deodara 'Prostrate Beauty' (Prostrate Beauty Dwarf Deodar Cedar) · C. deodara 'Pygmaea' (Deodar Cedar) · C. deodara 'Pygmy' · C. deodara 'Raywood's Contorta' (Contorted Deodar Cedar) · C. deodara 'Raywood's Prostrate' · C. deodara 'Robusta' · C. deodara 'Roman Candle' · C. deodara 'Scott' · C. deodara 'Shalimar' (Deodar Cedar) · C. deodara 'Silver Mist' (Deodar Cedar) · C. deodara 'Silver Spring' · C. deodara 'Snow Sprite' (Snow Sprite Compact Deodor Cedar) · C. deodara 'Twisted Growth' (Contorted Deodar Cedar) · C. deodara 'Verticillata Glauca' · C. deodara 'Viridis Prostrata' (Weeping Deodar Cedar) · C. deodara 'White Imp' (Dwarf White Himalayan Cedar) · C. deodara (Roxb. ex D.Don) G.Don 'Compacta' · C. deodara (Roxb. ex D.Don) G.Don 'Crassifolia' · C. deodara (Roxb. ex D.Don) G.Don 'Erecta' · C. deodara (Roxb. ex D.Don) G.Don 'Feeling Blue' (Deodar Cedar) · C. deodara (Roxb. ex D.Don) G.Don 'Flava' · C. deodara (Roxb. ex D.Don) G.Don 'Fontinalis' · C. deodara (Roxb. ex D.Don) G.Don 'Glauca' · C. deodara (Roxb. ex D.Don) G.Don 'Golden Horizont' · C. deodara (Roxb. ex D.Don) G.Don 'Hesse' · C. deodara (Roxb. ex D.Don) G.Don 'Mutabilis' · C. deodara (Roxb. ex D.Don) G.Don 'Paktia' · C. deodara (Roxb. ex D.Don) G.Don 'Repandens' · C. deodara (Roxb. ex D.Don) G.Don 'Tristis' · C. deodara (Roxb. ex D.Don) G.Don 'Uncinata' · C. deodara (Roxb. ex D.Don) G.Don 'Variegata' · C. deodara (Roxb. ex D.Don) G.Don 'Verticillata' · C. deodara (Roxb. ex D.Don) G.Don 'Viridis' · C. deodara (Roxb. ex D.Don) G.Don 'Wiesemannii' · C. deodara Paktia Group · C. deodora · C. elegans · C. lebani · C. libanensis · C. libani (Cedar of Lebanon) · C. libani 'Aurea Prostrata' (Cedar of Lebanon) · C. libani 'Blue Angel' · C. libani 'Comte de Dijon' · C. libani 'De Creffe' · C. libani 'Eugene' (Cedar of Lebanon) · C. libani 'Fontaine' · C. libani 'Glauca Pendula' (Cedar of Lebanon) · C. libani 'Glauca' · C. libani 'Gold Tip' (Cedar of Lebanon) · C. libani 'Green Prince' (Cedar of Lebanon) · C. libani 'Home Park' · C. libani 'Katere' (Cedar of Lebanon) · C. libani 'May' · C. libani 'Nana' · C. libani 'Pampisford' · C. libani 'Pendula' (Cedar of Lebanon) · C. libani 'Purdue Hardy' · C. libani 'Saint Catherine' · C. libani 'Sargentii' (Cedar of Lebanon) · C. libani 'Taurus' · C. libani A.Rich. 'Aurea' · C. libani A.Rich. 'Decidua' · C. libani A.Rich. 'Denudata' · C. libani A.Rich. 'Golden Dwarf' · C. libani A.Rich. 'Multicaulis' · C. libani A.Rich. 'Nana Pyramidata' · C. libani A.Rich. 'Nana' (Dwarf Lebanon Cedar) · C. libani A.Rich. 'Stricta' · C. libani A.Rich. 'Tortuosa' · C. libani A.Rich. 'Viridis' · C. libani atlantica · C. libani atlantica 'Glauca' · C. libani brevifolia · C. libani libani (Cedar of Lebanon) · C. libani Nana Group · C. libani stenocoma var. stenocoma · C. libani var. brevifolia · C. libani var. libani (Cedar of Lebanon) · C. libani var. stenocoma (Turkish Cedar) · C. libanii 'Green Prince' · C. libanii 'Hedgehog' · C. libanitica stenocoma · C. libanotica · C. mahogani (Small-Leaved Mahogany) · C. montis libani · C. odorata · C. sempervirens · C. torulosa · C. verticillatus 'Glauca' · C. libani var. libani
References
- ^ a b c d Farjon, A. (1990). Pinaceae. Drawings and Descriptions of the Genera. Koeltz Scientific Books ISBN 3-87429-298-3.
- ^ Frankis, M. & Lauria, F. (1994). The maturation and dispersal of cedar cones and seeds. International Dendrology Society Yearbook 1993: 43?46.
- ^ Liston A., D.S. Gernandt, T.F. Vining, C.S. Campbell, D. Pi?ero. 2003. Molecular Phylogeny of Pinaceae and Pinus. In Mill, R.R. (ed.): Proceedings of the 4th Conifer Congress. Acta Hort 615: Pp. 107-114.
- ^ Wang, X.-Q., Tank, D. C. and Sang, T. (2000): Phylogeny and Divergence Times in Pinaceae: Evidence from Three Genomes. Molecular Biology and Evolution 17:773-781. Available online
- ^ Gymnosperm database Cedrus.
- ^ NCBI Taxonomy Browser Cedrus.
- ^ Flora of China vol. 4
- ^ Qiao, C.-Y., Jin-Hua Ran, Yan Li and Xiao-Quan Wang (2007): Phylogeny and Biogeography of Cedrus (Pinaceae) Inferred from Sequences of Seven Paternal Chloroplast and Maternal Mitochondrial DNA Regions. Annals of Botany 100(3):573-580. Available online
- ^ Farjon, A. (2008). A Natural History of Conifers. Timber Press ISBN 0-88192-869-0.
- ^ Christou, K. A. (1991). The genetic and taxonomic status of Cyprus Cedar, Cedrus brevifolia (Hook.) Henry. Mediterranean Agronomic Institute of Chania, Greece.
- ^ GRIN Taxonomy for Plants Cedrus.
- ^ G?ner, A., ?zhatay, N., Ekim, T., & Baser, K. H. C. (ed.). 2000. Flora of Turkey and the East Aegean Islands 11 (Supplement 2): 5?6. Edinburgh University Press. ISBN 0-7486-1409-5
- ^ Eckenwalder, J. E. (2009). Conifers of the World: The Complete Reference. Timber Press ISBN 0-88192-974-3.
- ^ Sell, P. D. (1990). Some new combinations in the British Flora. Watsonia 18: 92.
- ^ ?dum, S. (1985). Report on frost damage to trees in Denmark after the severe 1981/82 and 1984/85 winters. H?rsholm Arboretum, Denmark.
- ^ Cedarwood oils
- ^ Meiggs, R. 1982. Trees and Timber in the Ancient Mediterranean World.
- ^ Andrews, A. C. 1961. Acclimatization of citrus fruits in the Mediterranean region. Agricultural History 35: 35?46.
- ^ a b Oxford English Dictionary.
- ^ Kelsey, H. P., & Dayton, W. A. (1942). Standardized Plant Names, second edition. American Joint Committee on Horticultural Nomenclature. Horace McFarland Company, Harrisburg, Pennsylvania.
Sources
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