Overview
Brickellia is a genus of about 1001] species of plants in the sunflower family (Asteraceae), known commonly as brickellbushes. They are found in North America, Mexico, and Central America.[1] Many species are native to the American southwest, especially Texas. Brickellia is among the more basal lineages of the Eupatorieae and should not be assigned to a subtribe pending further research.[2]
They are mostly woody perennial shrubs. Some species have a very strong pleasant scent, while others smell distasteful. All contain high amounts of essential oils, and Germacrene D, a natural insecticide, is found in B. veronicaefolia and probably other if not all species[3].
Regardless their chemical defense, brickellbushes are food for caterpillars of certain Lepidoptera. These include the noctuid moths Schinia buta (only known from California Brickellbush, B. californica), Schinia gracilenta (known from False Boneset, ), Schinia oleagina (only known from this genus) and Schinia trifascia.
Classification
The genera Brickelliastrum (United States and Mexico), Asanthus (United States and Mexico), Dyscritogyne (Mexico), and Steviopsis have been separated from Brickellia by many 20th century authors (and all four combined into Steviopsis by some). Their correct placement is still unclear.[4]
Selected species
- Brickellia amplexicaulis ? Earleaf Brickellbush
- Brickellia arguta ? Pungent Brickellbush
- Brickellia brachyphylla ? Plumed Brickellbush
- Brickellia californica ? California Brickellbush
- Brickellia coixtlahuaca[5]
- Brickellia cordifolia ? Flyr's Nemesis
- Brickellia coulteri ? Coulter's Brickellbush
- Brickellia cylindracea ? Gravelbar Brickellbush
- Brickellia desertorum ? Desert Brickellbush
- Brickellia eupatorioides ? False Boneset
- Brickellia grandiflora ? Tassel-flowered Brickellbush
- Brickellia greenei ? Greene's Brickellbush
- Brickellia huahuapana[5]
- Brickellia incana ? Woolly Brickellbush
- Brickellia knappiana ? Knapp Brickellbush
- Brickellia mosieri ? Florida Brickellbush
- Brickellia nevinii ? Nevin's Brickellbush
- Brickellia oblongifolia ? Mojave Brickellbush, Narrow-leaved Brickellbush
- Brickellia peninsularis
- Brickellia peninsularis var. amphithalassa
- Brickellia rusbyi ? Stinking Brickellbush
- Brickellia venosa ? Veiny Brickellbush
- Brickellia veronicaefolia
Footnotes
- ^ a < b>b "Brickellia". Flora of North America. http://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=104617.
- ^ Schmidt & Schilling (2000)
- ^ Rivero-Cruz et al. (2006)
- ^ "Brickelliastrum". Flora of North America. http://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=104618.
- ^ a b Turner, B. L. (2010). Two new species of Brickellia (Asteraceae: Eupatorieae) from Oaxaca, Mexico. Phytologia 92:1 15.
They are mostly woody perennial shrubs. Some species have a very strong pleasant scent, while others smell distasteful. All contain high amounts of essential oils, and Germacrene D, a natural insecticide, is found in B. veronicaefolia and probably other if not all species[3].
Regardless their chemical defense, brickellbushes are food for caterpillars of certain Lepidoptera. These include the noctuid moths Schinia buta (only known from California Brickellbush, B. californica), Schinia gracilenta (known from False Boneset, ), Schinia oleagina (only known from this genus) and Schinia trifascia.
Classification
The genera Brickelliastrum (United States and Mexico), Asanthus (United States and Mexico), Dyscritogyne (Mexico), and Steviopsis have been separated from Brickellia by many 20th century authors (and all four combined into Steviopsis by some). Their correct placement is still unclear.[4]
Selected species
- Brickellia amplexicaulis ? Earleaf Brickellbush
- Brickellia arguta ? Pungent Brickellbush
- Brickellia brachyphylla ? Plumed Brickellbush
- Brickellia californica ? California Brickellbush
- Brickellia coixtlahuaca[5]
- Brickellia cordifolia ? Flyr's Nemesis
- Brickellia coulteri ? Coulter's Brickellbush
- Brickellia cylindracea ? Gravelbar Brickellbush
- Brickellia desertorum ? Desert Brickellbush
- Brickellia eupatorioides ? False Boneset
- Brickellia greenei ? Greene's Brickellbush
- Brickellia huahuapana[5]
- Brickellia incana ? Woolly Brickellbush
- Brickellia knappiana ? Knapp Brickellbush
- Brickellia mosieri ? Florida Brickellbush
- Brickellia nevinii ? Nevin's Brickellbush
- Brickellia oblongifolia ? Mojave Brickellbush, Narrow-leaved Brickellbush
- Brickellia peninsularis
- Brickellia peninsularis var. amphithalassa
- Brickellia rusbyi ? Stinking Brickellbush
- Brickellia venosa ? Veiny Brickellbush
- Brickellia veronicaefolia
Footnotes
- ^
a b "Brickellia". Flora of North America. http://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=104617. - ^ Schmidt & Schilling (2000)
- ^ Rivero-Cruz et al. (2006)
- ^ "Brickelliastrum". Flora of North America. http://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=104618.
- ^ a b Turner, B. L. (2010). Two new species of Brickellia (Asteraceae: Eupatorieae) from Oaxaca, Mexico. Phytologia 92:1 15.
References
- Rivero-Cruz, Blanca; Rivero-Cruz, Isabel; Rodriguez, Juan M.; Cerda-Garcia-Rojas, Carlos M. & Mata, Rachel (2006): Qualitative and quantitative analysis of the active components of the essential oil from Brickellia veronicaefolia by nuclear magnetic resonance spectroscopy. Journal of Natural Products 69(8): 1172-1176. doi:10.1021/np060180b S0163-3864(06)00180-7 (HTML abstract)
- S chmidt, Gregory J. & Schilling, Edward E. (2000): Phylogeny and biogeography of Eupatorium (Asteraceae: Eupatorieae) based on nuclear ITS sequence data. Am. J. Bot. 87(5): 716-726. doi:10.2307/2656858 PMID 10811796 PDF fulltext
External links
Taxonomy
The Genus Brickellia is further organized into finer groupings including:
- Species: ZipcodeZoo has pages for 207 species, subspecies, varieties, forms, and cultivars in the Genus Brickellia: B. adenocarpa · B. adenolepis · B. ambigens · B. amblyolepis · B. amplexicaulis (Ear-Leaf Brickellbush) · B. amplexicaulis B.L.Robins. var. amplexicaulis B.L.Robins. (Earleaf Brickellbush) · B. amplexicaulis B.L.Robins. var. lanceolata (Gray) B.L.Robins. (Earleaf Brickellbush) · B. amplexicaulis var. amplexicaulis (Earleaf Brickellbush) · B. amplexicaulis var. lanceolata (Earleaf Brickellbush) · B. anomala · B. aramberrana · B. arguta (Hollyleaf Brickellbush) · B. arguta odontolepis · B. arguta var. arguta (Pungent Brickellbush) · B. arguta var. odontolepis (Pungent Brickellbush) · B. argyrolepis · B. arsenei · B. atractyloides (Spear-Leaf Brickell-Bush) · B. atractyloides Gray var. atractyloides Gray (Spearleaf Brickellia) · B. atractyloides var. atractyloides (Spearleaf Brickellia) · B. atractyloides var. odontolepis · B. baccharidea (Baccharisleaf Brickellbush) · B. betonicaefolia · B. betonicifolia (Betony-Leaf Brickell-Bush) · B. betonicifolia var. conduplicata · B. betonicifolia var. elliptica · B. botterii · B. brachiata · B. brachiata glabrata · B. brachyphylla (Hinckleys Brickellbush) · B. brachyphylla var. brachyphylla (Plumed Brickellbush) · B. brachyphylla var. hinckleyi (Hinckley's Brickellia) · B. brachyphylla var. terlinguensis (Terlingua Brickellia) · B. brandegei · B. brasiliensis · B. californica (California Brickell-Bush) · B. californica (Torr. & Gray) Gray var. californica (Torr. & Gray) Gray · B. californica var. californica (California Brickellbush) · B. californica var. jepsonii (Jepson's Brickellbush) · B. cardiophylla · B. cavanillesii · B. chenopodina (Chenopod Brickell-Bush) · B. chlorolepis · B. coahuilensis · B. colimae · B. conduplicata (Southwestern Brickellbush) · B. cordifolia (Flyr's Brickell-Bush) · B. corymbosa · B. coulteri (Coulter's Brickell-Bush) · B. coulteri var. adenopoda · B. coulteri var. brachiata · B. cuspidata · B. cylindracea (Gravel-Bar Brickell-Bush) · B. cymulifera · B. dentata (Leafy Brickell-Bush) · B. desertorum (Desert Brickellbush) · B. diffusa · B. eupatorioides (Thoroughwort Brickellbush) · B. eupatorioides (L.) Shinners var. chlorolepis (Woot. & Standl.) B.L.Turner · B. eupatorioides (L.) Shinners var. corymbulosa (Torr. & Gray) Shinners (False Boneset) · B. eupatorioides (L.) Shinners var. texana (Shinners) Shinners · B. eupatorioides var. chlorolepis (False Boneset) · B. eupatorioides var. corymbulosa (False Boneset) · B. eupatorioides var. eupatorioides (False Boneset) · B. eupatorioides var. floridana (False Boneset) · B. eupatorioides var. gracilis (False Boneset) · B. eupatorioides var. gracillima (False Boneset) · B. eupatorioides var. texana (False Boneset) · B. extranea · B. fendleri · B. filipes · B. floribunda (Chihuahuan Brickellbush) · B. frutescens (Shrubby Brickellbush) · B. gentianoides · B. gentryi · B. glabrata · B. glandulosa · B. glomerata · B. glutinosa · B. grandiflora (Tasselflower Brickellbush) · B. grandiflora petiolaris · B. greenei (Greene Brickellbush) · B. guatemalensis · B. hastata · B. hebecarpa · B. hinckleyi · B. hinckleyi var. hinckleyi · B. hinckleyi var. terlinguensis (Terlingu Brickellia) · B. hintoniorum · B. hymenochlaena · B. incana (Woolly Brickellbush) · B. jaliscensis · B. kellermanii · B. knappiana (Knapp's Brickellbush) · B. laccata · B. laciniata (Splitleaf Brickellbush) · B. lanata · B. lanata microdonta · B. lancifolia · B. laxiflora · B. lemmonii (Lemmon's Brickellbush) · B. lemmonii Gray var. wootonii (Greene) B.L.Robins. · B. lemmonii var. conduplicata · B. lemmonii var. lemmonii (Lemmon's Brickellbush) · B. lemmonii var. wootonii (Lemmon's Brickellbush) · B. leptophylla · B. lewisii · B. linifolia · B. linifolia var. linifolia · B. longifolia (Long Leaf Brickellbush) · B. longifolia var. longifolia (Longleaf Brickellbush) · B. longifolia var. multiflora (Longleaf Brickellbush) · B. longifolia var. multiflora (Kellogg) Cronq. (Longleaf Brickellbush) · B. macromera · B. magnifica · B. mcdonaldii · B. megalodonta · B. megaphylla · B. microphylla (Littleleaf Brickellbush) · B. microphylla scabra (Littleleaf Brickellbush) · B. microphylla var. microphylla (Littleleaf Brickellbush) · B. microphylla var. scabra (Rough Brickellbush) · B. microphylla var. watsonii (Watson's Brickellbush) · B. mohavensis · B. monocephala · B. multiflora · B. multiflora var. multiflora · B. nelsonii (Nelson's Brickellbush) · B. nesomii · B. nevinii (Nevin's Brickellbush) · B. nutans · B. nutanticeps · B. oblongifolia (Narrow-Leaf Brickell-Bush) · B. oblongifolia abbreviata · B. oblongifolia linifolia (Narrow Leaf Brickellbush) · B. oblongifolia var. abbreviata · B. oblongifolia var. linifolia (Narrow Leaf Brickellbush) · B. oblongifolia var. oblongifolia (Narrowleaf Brickellbush) · B. odontophylla · B. oligadena · B. oliganthes · B. oreithales · B. orizabaensis · B. orizabensis · B. pacayensis · B. palmeri · B. palmeri var. amphothrix · B. paniculata · B. parryi · B. parvula (Mt. Davis Brickellbush) · B. paucidentata · B. pavonii · B. pedunculosa · B. pendula · B. peninsularis · B. petrophila · B. petrophila umbratilis · B. pinifolia · B. pringlei (Pringle's Brickellbush) · B. problematica · B. pulcherrima · B. recondita · B. reniformis · B. rhomboidea · B. riddellii · B. robinsoniana · B. rosmarinifolia · B. rusbyi (Stinking Brickell-Bush) · B. saltillensis · B. scabra · B. schaffneri · B. scoparia · B. scoparia subauriculata · B. scoparia var. subauriculata · B. secundiflora · B. seemannii · B. shineri · B. simplex (Sonoran Brickell-Bush) · B. solidaginifolia · B. sonorana · B. spinulosa · B. spinulosa asperata · B. squamulosa · B. squarrosa · B. stolonifera · B. subsessilis · B. subuligera · B. tenera · B. thyrsiflora · B. tomentella · B. umbellata · B. urolepis · B. venosa (Veined Brickell-Bush) · B. vernicosa · B. veronicaefolia · B. veronicifolia (Veronica-Leaved Brickell-Bush) · B. veronicifolia var. petrophila (Veronicaleaf Brickellbush) · B. veronicifolia var. senilis · B. veronicifolia var. veronicifolia (Veronicaleaf Brickellbush) · B. viejensis (Tierra Vieja Brickellbush) · B. villarrealii · B. vollmeri · B. watsonii · B. wendtii · B. wislizeni · B. worthingtonii · B. wrightii
References
- Rivero-Cruz, Blanca; Rivero-Cruz, Isabel; Rodriguez, Juan M.; Cerda-Garcia-Rojas, Carlos M. & Mata, Rachel (2006): Qualitative and quantitative analysis of the active components of the essential oil from Brickellia veronicaefolia by nuclear magnetic resonance spectroscopy. Journal of Natural Products 69(8): 1172-1176. doi:10.1021/np060180b S0163-3864(06)00180-7 (HTML abstract)
- Schmidt, Gregory J. & Schilling, Edward E. (2000): Phylogeny and biogeography of Eupatorium (Asteraceae: Eupatorieae) based on nuclear ITS sequence data. Am. J. Bot. 87(5): 716-726. doi:10.2307/2656858 PMID 10811796 PDF fulltext
External link s
Footnotes
Sources
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