Overview
The bamboos are a group of perennial evergreen (except for certain temperate species) plants in the true grass family Poaceae, subfamily Bambusoideae, tribe Bambuseae. Some are giant bamboos, the largest members of the grass family. Bamboos are the fastest growing woody plants in the world. Their growth rate (up to 60 centimeters (24 in.)/day) is due to a unique rhizome-dependent system, but is highly dependent on local soil and climate conditions. They are of economic and high cultural significance in East Asia and South East Asia where they are used extensively in gardens, as a building material, and as a food source.
There are more than 70 genera divided into about 1,000 species.[1] They are found in diverse climates, from cold mountains to hot tropical regions. They occur across East Asia, from 50°N latitude in Sakhalin through to Northern Australia, and west to India and the Himalayas.[2] They also occur in sub-Saharan Africa, and in the Americas from the Mid-Atlantic United States[3] south to Argentina and Chile, reaching their southernmost point anywhere, at 47°S latitude. Major areas with no native bamboos include Europe and Antarctica.[4]
Ecology
Growth
Bamboo is the fastest-growing plant on Earth; it has been measured surging skyward as fast as 121 cm (47.6 inches) in a 24-hour period,[5] and can also reach maximal growth rate exceeding one meter (39 inches) per hour for short periods of time. Many prehistoric bamboos exceeded heights of 75 meters (250 feet). Primarily growing in regions of warmer climates during the Cretaceous, vast fields existed in what is now Asia. Modern bamboos can only sustain their maximal growth rate for short periods of time.[clarification needed]
Unlike trees, all bamboos grow to full height and girth in a single growing season of 3–4 months. During this first year the young shoots strike skyward supported by photosynthesis from the rest of the clump with no time to sprout their own branches and leaves. Over the next year the pulpy wall of each culm slowly dries and hardens, sprouting branches and leaves during the second year from juvenile sheathes that form from each node. Over the following year the culm hardens still further shedding its juvenile sheaths and commencing its life as a fully mature culm. Over the next 2–5 years depending on species, fungus and mould begin to form on the outside of the culm, eventually penetrating and overcoming the culm so that by around 5 – 8 years depending on species and climate the culms begin to collapse and decay. This brief life means culms are ready for harvest and suitable for use in construction from 3 – 7 years.
Mass Flowering
Although some bamboos flower every year, most species flower infrequently. In fact, many bamboos only flower at intervals as long as 60 or 120 years. These taxa exhibit mass flowering (or gregarious flowering), with all plants in the population flowering simultaneously. The longest mass flowering interval known is 130 years, and is found for all the species Phyllostachys bambusoides (Sieb. & Zucc.). In this species, all plants of the same stock flower at the same time, regardless of differences in geographic locations or climatic conditions, then the bamboo dies. The lack of environmental impact on the time of flowering indicates the presence of some sort of “alarm clock” in each cell of the plant which signals the diversion of all energy to flower production and the cessation of vegetative growth.[6] This mechanism, as well as the evolutionary cause behind it, is still largely a mystery.
One theory to explain the evolution of this semelparous mass flowering is the predator satiation hypothesis. This theory argues that by fruiting at the same time, a population increases the survival rate of their seeds by flooding the area with fruit so that even if predators eat their fill, there will still be seeds left over. By having a flowering cycle longer than the lifespan of the rodent predators, bamboos can regulate animal populations by causing starvation during the period between flowering events. Thus, according to this hypothesis, the death of the adult clone is due to resource exhaustion, as it would be more effective for parent plants to devote all resources to creating a large seed crop than to hold back energy for their own regeneration.[7]
A second theory, the fire cycle hypothesis, argues that periodic flowering followed by death of the adult plants has evolved as a mechanism to create disturbance in the habitat, thus providing the seedlings with a gap in which to grow. This hypothesis argues that the dead culms create a large fuel load, and also a large target for lightning strikes, increasing the likelihood of wildfire.[8] Because bamboos are very aggressive as early successional plants, the seedlings would be able to outstrip other plants and take over the space left by their parents.
However, both have been disputed for different reasons. The predator satiation theory does not explain why the flowering cycle is 10 times longer than the lifespan of the local rodents, something not predicted by the theory. The bamboo fire cycle theory is considered by a few scientists to be unreasonable because, as argued by [9] fires only result from humans and there is no natural fire in India. This notion is considered wrong based on distribution of lightning strike data during the dry season throughout India [10]
The mass fruiting also has direct economic and ecological consequences, however. The huge increase in available fruit in the forests often causes a boom in rodent populations, leading to increases in disease and famine in nearby human populations. For example, there are devastating consequences when the Melocanna bambusoides population flowers and fruits once every 30–35 years around the Bay of Bengal. The death of the bamboo plants following their fruiting means the local people lose their building material, and the large increase in bamboo fruit leads to a rapid increase in rodent populations. As the number of rodents increase, they consume all available food, including grain fields and stored food, sometimes leading to famine. These rats can also carry dangerous diseases such as typhus, typhoid, and bubonic plague, which can reach epidemic proportions as the rodents increase in number.[6][7]
Bamboo in Animal Diets
Soft bamboo shoots, stems, and leaves are the major food source of the Giant Panda of China. Rats will eat the fruits as described above. Mountain Gorillas of Africa also feed on bamboo and have been documented consuming bamboo sap which was fermented and alcoholic; chimps and elephants of the region also eat the stalks.[11]
Cultivation
Commercial Timber
Timber is harvested from cultivated and wild stands and some of the larger bamboos, particularly species in the genus Phyllostachys, are known as "timber bamboos".
Harvesting
If using bamboo for construction purposes it is critical to harvest the culms when at their greatest strength and when sugar levels in the sap are at their lowest, as high sugar content increases the ease and rate of pest infestation.
Harvesting of bamboo should be undertaken according to the following cycles.
1) Life cycle of the clump: As each individual culm goes through a 5-7 year life cycle, culms should ideally be allowed to reach this level of maturity prior to full capacity harvesting. The clearing out, thinning of culms particularly older decaying culms will help ensure adequate light and resources for new growth. Well maintained clumps may have a productivity 3-4 times that of an un harvested wild clump.
2) Life cycle of the culm: As per the life cycle described above, bamboo should be harvested from 2–3 years through to 5–7 years, depending on the species.
3) Annual cycle: As all growth of new bamboo occurs during the wet season, disturbing the clump during this phase will potentially damage the upcoming crop. Also during this high rain fall period sap levels are at their highest, with sap levels diminishing towards the dry season. Picking immediately prior to the wet/growth season may also damage new shoots. Hence harvesting is best at the end of the dry season, a few months prior to the start of the wet.
4) Daily cycle: During the height of the day, Photosynthesis is at its peak producing the highest levels of sugar in sap, making this the least ideal time of day to harvest. Many traditional practitioners believe that the best time to harvest is at dawn or dusk on a full moon. This practice makes sense in terms of both moon cycles, visibility and daily cycles.
Leaching
Leaching is the removal of sap post-harvest. In many areas of the world the sap levels in harvested bamboo are reduced either through leaching or post-harvest photosynthesis. Examples of this practice include:
- Cut bamboo is raised clear of the ground and leant against the rest of the clump for 1–2 weeks until leaves turn yellow to allow full consumption of sugars by the plant
- A similar method is undertaken but with the base of the culm standing in fresh water, either in a large drum or stream to leach out sap
- Cut culms are immersed in a running stream and weighted down for 3–4 weeks
- Water is pumped through the freshly cut culms forcing out the sap (this method is often used in conjunction with the injection of some form of treatment)
- Bamboo is very good for spanking people as well
In the process of water leaching, the bamboo is dried slowly and evenly in the shade to avoid cracking in the outer skin of the bamboo, thereby reducing opportunities for pest infestation.
Durability of bamboo in construction is directly related to how well it is handled from the moment of planting through harvesting, transportation, storage, design, construction and maintenance. Bamboo harvested at the correct time of year and then exposed to ground contact or rain, will break down just as quickly as incorrectly harvested material.
Ornamental Bamboos
Many bamboos are popular in cultivation as garden trees. There are two general patterns for the growth of bamboo: "clumping" (sympodial) and "running" (monopodial). Clumping bamboo species tend to spread slowly, as the growth pattern of the rhizomes is to simply expand the root mass gradually, similar to ornamental grasses. "Running" bamboos, on the other hand, need to be taken care of in cultivation because of their potential for aggressive behavior. They spread mainly through their roots and/or rhizomes, which can spread widely underground and send up new culms to break through the surface. Running bamboo species are highly variable in their tendency to spread; this is related to both the species and the soil and climate conditions. Some can send out runners of several meters a year, while others can stay in the same general area for long periods. If neglected, over time they can cause problems by moving into adjacent areas.
Bamboos seldom and unpredictably flower, and the frequency of flowering varies greatly from species to species. Once flowering takes place, a plant will decline and often die entirely. Although there are always a few species of bamboo in flower at any given time, collectors desiring to grow specific bamboo typically obtain their plants as divisions of already-growing plants, rather than waiting for seeds to be produced.
Regular maintenance will indicate major growth directions and locations. Once the rhizomes are cut, they are typically removed; however, rhizomes take a number of months to mature and an immature, severed rhizome will usually cease growing if left in-ground. If any bamboo shoots come up outside of the bamboo area afterwards, their presence indicates the precise location of the missed rhizome. The fibrous roots that radiate from the rhizomes do not grow up to be more bamboo so if they stay in the ground, that's not a problem.
The second way to control growth is by surrounding the plant or grove with a physical barrier. This method is very detrimental to ornamental bamboo as the bamboo within quickly becomes rootbound—showing all the signs of any unhealthy containerized plant. Symptoms include rhizomes escaping over the top, down underneath, and bursting the barrier. The bamboo within generally deteriorates in quality as fewer and fewer culms grow each year, culms live shorter periods, new culm diameter decreases, fewer leaves grow on the culms, and leaves turn yellow as the unnaturally contained rootmass quickly depletes the soil of nutrients, and curling leaves as the condensed roots cannot collect the water they need to sustain the foliage. Concrete and specially-rolled HDPE plastic are the usual materials used. This is placed in a 60-90 cm (2-3 feet) deep ditch around the planting, and angled out at the top to direct the rhizomes to the surface. (This is only possible if the barrier is installed in a straight line.) Strong rhizomes and tools can penetrate plastic barriers with relative ease, so great care must be taken. Barriers usually fail sooner or later, or the bamboo within suffers greatly. Casual observation of many failed barriers has shown bursting of 60 mil HDPE in 5–6 years, and rhizomes diving underneath in as few as 3 years post install. In small areas regular maintenance is the only perfect method of controlling the spreading bamboos. Bamboo in barriers is much more difficult to remove than free-spreading bamboo. Barriers and edging are unnecessary for clump-forming bamboos. Clump-forming bamboos may eventually need to have portions removed if they get too large.
The ornamental plant sold in containers and marketed as "lucky bamboo" is actually an entirely unrelated plant, Dracaena sanderiana. It is a resilient member of the lily family that grows in the dark, tropical rainforests of Southeast Asia and Africa. Lucky Bamboo has long been associated with the Eastern practice of Feng Shui. On a similar note, Japanese knotweed is also sometimes mistaken for a bamboo but it grows wild and is considred an invasive species.
U.S.A.s
Culinary
The shoots (new bamboo culms that come out of the ground) of bamboo are edible. They are used in numerous Asian dishes and broths, and are available in supermarkets in various sliced forms, both fresh and canne d version. The shoots of the giant bamboo contains cyanide. Despite this, the Golden Bamboo Lemur ingests many times the quantity of toxin that would kill a human.
The bamboo shoot in its fermented state (called khorisa) forms an important ingredient in the cuisine of Assam.
In Indonesia, they are sliced thin and then boiled with santan (thick coconut milk) and spices to make a dish named gulai rebung. Other recipes using bamboo shoots are sayur lodeh (mixed vegetables in coconut milk) and lun pia (sometimes written lumpia: fried wrapped bamboo shoots with vegetables). The shoots of some species contain toxins that need to be leached or boiled out before they can be eaten safely.
Pickled bamboo, used as a condiment, may also be made from the pith of the young shoots.
The sap of young stalks tapped during the rainy season may be fermented to make ulanzi (a sweet wine) or simply made into a soft drink. Bamboo leaves are also used as wrappers for steamed dumplings which usually contains glutinous rice and other ingredients.
The empty hollow in the stalks of larger bamboo is often used to cook food in many Asian cultures. Soups are boiled and rice is cooked in the hollows of fresh stalks of bamboo directly over a flame. Similarly, steamed tea is sometimes rammed into bamboo hollows to produce compressed forms of Pu-erh tea. Cooking food in bamboo is said to give the food a subtle but distinctive taste.
In Sambalpur, India, the tender shoots are grated into juliennes and fermented to prepare kardi. The name is derived from the Sanskrit word for bamboo shoot, "karira". This fermented bamboo shoot is used in various culinary preparations, notably "amil", a sour vegetable soup. It is also made into pancakes using rice flour as a binding agent. The shoots that have turned a little fibrous are fermented, dried, and ground to sand size particles to prepare a garnish known as "hendua". It is also cooked with tender pumpkin leaves to make sag green leaves.
In addition, bamboo is frequently used for cooking utensils within many cultures. In modern times, some see bamboo tools as an eco-friendly alternative to other manufactured utensils.[12]
Medicine
Bamboo is used in Chinese medicine for treating infections and healing.
It is a low-calorie source of potassium. It is known for its sweet taste and as a good source of nutrients and protein.
In Ayurveda, the Indian system of traditional medicine, the silicious concretion found in the culms of the bamboo stem is called banslochan. It is known as tabashir or tawashir in Unani-Tibb the Indo-Persian system of medicine. In English it is called "bamboo manna". This concretion is said to be a tonic for the respiratory diseases. It was earlier obtained from Melocanna bambusoides and is very hard to get; it has been largely replaced by synthetic silicic acid. In most Indian literature, Bambusa arundinacea is described as the source of bamboo manna.[13]
Construction
In its natural form, bamboo as a construction material is traditionaly associated with the cultures of East Asia and the South Pacific, to some extent in Central and South America and by extention in the aesthetic of Tiki culture. In China, bamboo was used to hold up simple suspension bridges, either by making cables of split bamboo or twisting whole culms of sufficiently pliable bamboo together. One such bridge in the area of Qian-Xian is referenced in writings dating back 960 A.D. and may have stood since as far back as the 3rd centureny B.C., due largely to continuious maintinence.[14] It has long been used as scaffolding; the practice has been banned in China for buildings over six stories but is still in continuious use for scyscrapers in Hong Kong.[15] In the Philipines, the Nipa Hut is a fairly typical example of the most basic sort of housing that bamboo is used for; the walls are split and woven bamboo and bamboo slats and poles may be used as its support. In Japanese architecture, bamboo is used primarily as a supplimental and/or decorative element in buildings such as fencing, fountians, grates and gutters, largely due to the ready abundence of quality timber wood.[16]
Various structural shapes may be made by training the bamboo to assume them as it grows. Squared sections of bamboo are created by compressing the growing stalk within a square form.[17] Arches may similarly be created by forcing the bamboo's growth with the desired form and costs many times less than it would to assume the same shape in regular wood timber. More traditional forming methoods such as the application of heat and preasure may also be used to curve or flatten the cut stalks.[18]
Bamboo may also be cut and laminated into sheets and planks. This process involves cutting stalks into thin strips, planing them flat, boiling and drying the strips which are then glued, pressed and finished.[19] Generaly long practiced in China and Japan, enterprauneers started developing and selling laminated bamboo flooring in the West during the mid 1990s[19]; products made from bamboo laminate including flooring, cabinetry, furniture and even decoritive use are currently surging in popularity, transitioning from the botique market to mainstream providers such as Home Depot. The bamboo goods industry (which also includes small goods, fabric, etc) is expected to be worth $25 billion by the year 2012.[20] The quality of bamboo laminate varries between manufactuerers and the maturity of the plant from which it was harvested (6 years being considered the optimum); the sturdiest products fufill their claims of being up to three times harder than oak hardwood but others may be softer than standard hardwood.[19]
Bamboo intended for use in construction should be treated to resist insects and rot. The most common solution for this purpose is a mixture of borax and boric acid.[21] Another process involves boiling cut bamboo in order to remove the starches that bugs are attracted to.[19].
Bamboo has been used as reenforcement for concrete in those areas where it is plentiful, though dispute exists over its effectiveness in the various studies done on the subject. Bamboo does have the neciccary strength to fufull this function, but untreated bamboo will swell from the absorbsion of water from the concrete, causing it to crack. Several procedures must be followed to overcome this shortcomming.[22]
Several institutes, businesses, and universities are working on the bamboo as an ecological construction material. In the United States and France, it is possible to get houses made entirely of bamboo, which are earthquake and cyclone-resistant and internationally certified. In Bali Indonesia there is a International primary school, named the Green School, which is constructed entirely of bamboo, due to its beauty, and advantages as a sustainable resource. There are three ISO standards for bamboo as a construction material.
Textiles
There are two methoods by which bamboo may be processed into fiber for fabric, both developed in China. The first is a mechanical process similar to that used to process flax or hemp; the stalks are crushed and natural enzymes break them down further, allowing fibers to be combed out.[23] The other follows the process by which rayon is made where the fibers are broken down with chemicals and extruded through mechanical spineretes; the chemicals include lye, carbon disulfide and strong acids.[19] Retailers have sold both end products as "bamboo fabric" to cash in on bamboo's current eco-friendly cachet, however the Canadian Competition Bureau[24] and the US Federal Trade Commission[25], as of mid-2009, are cracking down on the practice of labeling bamboo rayon as natural bamboo fabric. Under the guidelines of both agencies these products must be labeled as rayon with the optional qualifier "from bamboo". Bamboo fabric is known for its softness and boasts strong absorbancy and anti-microbial properties, though the chemical process in bamboo rayon destroyes any anti-microbial quality.[25]
In addition, the fiber of bamboo has been used to make paper in China since early times. A high quality hand-made paper is still produced in small quantities. Coarse bamboo paper is still used to make spirit money in many Chinese communities.
Musical Instruments
Bamboo's natural hollow form makes it an obvious choice for many instruments, particularly wind and percussion. There are numerous types of bamboo flute made all over the world, such as the dizi, xiao, shakuhachi, palendag, jinghu, angklung. The Bamboo Organ of Las Piñas, Philippines has pipes made of bamboo culms. Four of the instruments used in Polynesia for traditional hula are made of bamboo: nose flute, rattle, stamping pipes and the Jew's harp. Bamboo may be used in the construction of the Austrialian didgeridoo instead of the more traditional eucalyptus wood. In Indonesia, bamboo has been used for making various kinds of musical instruments including the kolintang and the angklung.
Bamboo in Asian Culture
Bamboo's long life makes it a Chinese symbol of longevity, while in India it is a symbol of friendship. The rarity of its blossoming has led to the flowers' being regarded as a sign of impending famine. This may be due to rats feeding upon the profusion of flowers, then multiplying and destroying a large part of the local food supply. The most recent flowering began in May 2006 (see Mautam). Bamboo is said to bloom in this manner only about every 50 years (see 28–60 year examples in FAO: 'gregarious' species table).
In Chinese culture, the bamboo, plum blossom, orchid, and chrysanthemum (often known as méi lán zhú jú ????) are collectively referred to as the Four Noble Ones. These four plants also represent the four seasons and, in Confucian ideology, four aspects of the junzi ("prince" or "noble one").
The pine tree, the bamboo, and the plum blossom (song zhú méi ???) are also admired for their perseverance under harsh conditions, and are together known as the "Three Friends in Winter" (????). The "Three Friends" is traditionally used as a system of ranking in Japan, for example in sushi sets or accommodations at a traditional Ryokan (inn). Pine (matsu ?) is of the first rank, bamboo (také ?) is of second rank, and plum (ume ?) is of the third.
In Japan, a bamboo forest sometimes surrounds a Shinto shrine as part of a sacred barrier against evil. Many Buddhist temples also have bamboo groves.
In northern Indian state of Assam, the fermented bamboo paste known as khorisa is known locally as a folk remedy for the treatment of impotence, infertility, and menstrual pains.
Bamboo symbolizes the spirit of Vovinam (a Vietnamese martial arts): "cuong nhu ph?i tri?n" (coordination between hard and soft (martial arts)). Bamboo also symbolizes the Vietnamese hometown and Vietnamese soul: the gentlemanlike, straightforwardness, hard working, optimism, unity and adaptableness. Furthermore, some scientists even regard that Vietnamese culture is bamboo culture. A Vietnamese proverb says: "When the bamboo is old, the bamboo sprouts appear", the meaning being Vietnam will never be annihilated; if the previous generation dies, the children take their place. Therefore the Vietnam nation and Vietnamese value will be maintained and developed eternally. Traditional Vietnamese villages are surrounded by thick bamboo hedges (luy tre).
The ethnic group known as the Bozo of West Africa, take their name from the Bambara phrase bo-so, which means "bamboo house".
The Song Dynasty (960-1279 AD) Chinese scientist and polymath Shen Kuo (1031-1095) used the evidence of underground petrified bamboo found in the dry northern climate of Yan'an, Shanbei region, Shaanxi province to support his geological theory of gradual climate change.[26][27]
Myths and Legends
Several Asian cultures, including that of the Andaman Islands, believe that humanity emerged from a bamboo stem. In the Philippine creation myth, legend tells that the first man and the first woman each emerged from split bamboo stems on an island created after the battle of the elemental forces (Sky and Ocean). In Malaysian legends a similar story includes a man who dreams of a beautiful woman while sleeping under a bamboo plant; he wakes up and breaks the bamboo stem, discovering the woman inside. The Japanese folktale "Tale of the Bamboo Cutter" (Taketori Monogatari) tells of a princess from the Moon emerging from a shining bamboo section. Hawaiian bamboo ('ohe) is a kinolau or body form of the Polynesian creator god Kane Milohai.
Bamboo cane is also the weapon of Vietnamese legendary hero Saint Giong- who had grown up immediately and magically since the age of 3 years old because of his national liberating wish against Ân invaders.
An ancient Vietnamese legend (The Hundred-knot Bamboo Tree) tells of a poor, young farmer who fell in love with his landlord's beautiful daughter. The farmer asked the landlord for his daughter's hand in marriage, but the proud landlord would not allow her to be bound in marriage to a poor farmer. The landlord decided to foil the marriage with an impossible deal; the farmer must bring him a "bamboo tree of one-hundred nodes". But Buddha (B?t) appeared to the farmer and told him that such a tree could be made from one-hundred nodes from several different trees. B?t gave to him four magic words to attach the many nodes of bamboo: "Kh?c nh?p, kh?c xu?t", which means "joined together immediately, fell apart immediately". The triumphant farmer returned to the landlord and demanded his daughter. Curious to see such a long bamboo, the landlord was magically joined to the bamboo when he touched it as the young farmer said the first two magic words. The story ends with the happy marriage of the farmer and the landlord's daughter after the landlord agreed to the marriage and asked to be separated from the bamboo.
Bamboo As a Writing Material
Bamboo was in widespread use in early China as a medium for written documents. The earliest surviving examples of such documents, written in ink on string-bound bundles of bamboo strips (or "slips"), date from the 5th c. BC during the Warring States period. However, references in earlier texts surviving on other media make it clear that some precursor of these Warring States period bamboo slips was in use as early as the late Shang period (from about 1250 BC).
Bamboo or wooden strips were the standard writing material during the Han dynasty and excavated examples have been found in abundance. [28] Subsequently, paper began to displace bamboo and wooden strips from mainstream uses, and by the 4th c. AD bamboo had been largely abandoned as a medium for writing in China.
Gallery
Photos
Taxonomy
The Tribe Bambuseae is a member of the Subfamily Bambusoideae. Here is the complete "parentage" of Bambuseae:
- Domain: Eukaryota
Whittaker & Margulis,1978 - eukaryotes
- Kingdom: Plantae
Haeckel, 1866
- Subkingdom: Viridaeplantae
Cavalier-Smith, 1981 - Green Plants
- Phylum: Tracheophyta
Sinnott, 1935 Ex Cavalier-Smith, 1998 - Vascular Plants
- Subphylum: Euphyllophytina
- Infraphylum: Radiatopses
Kenrick & Crane, 1997
- Class: Liliopsida
Scopoli, 1760 - Monocotyledons
- Subclass: Commelinidae
Takhtajan, 1967
- Superorder: Poanae
(Small, 1903) Takhtajan, 1997 Ex Reveal & Doweld, 1999
- Order: Poales
Small, 1903
- Family: Poaceae
(poh-AY-see-ee)
(R. Brown) Barnhart, 1895
- Subfamily: Bambusoideae
- Tribe: Bambuseae
- Subfamily: Bambusoideae
- Family: Poaceae
(poh-AY-see-ee)
(R. Brown) Barnhart, 1895
- Order: Poales
Small, 1903
- Superorder: Poanae
(Small, 1903) Takhtajan, 1997 Ex Reveal & Doweld, 1999
- Subclass: Commelinidae
Takhtajan, 1967
- Class: Liliopsida
Scopoli, 1760 - Monocotyledons
- Infraphylum: Radiatopses
Kenrick & Crane, 1997
- Subphylum: Euphyllophytina
- Phylum: Tracheophyta
Sinnott, 1935 Ex Cavalier-Smith, 1998 - Vascular Plants
- Subkingdom: Viridaeplantae
Cavalier-Smith, 1981 - Green Plants
- Kingdom: Plantae
Haeckel, 1866
The Tribe Bambuseae is further organized into finer groupings including:
- Subtribe (1): Chusqueinae
- Genus (17): Arthrostylidium · Arundinaria · Bambusa · Banisteria · Chimonobambusa · Chusquea · Cotoneaster · Dendrocalamus · Gigantochloa · Guadua · Neurolepis · Oxytenanthera · Phyllostachys · Pseudosasa · Sasa · Schizostachyum · Semiarundinaria
- Species: ZipcodeZoo has pages for 245 species, subspecies, varieties, forms, and cultivars in the Tribe Bambuseae.
Genera
Arthrostylidium
Arthrostylidium is a of bamboo (tribe Bambuseae of the family Poaceae), comprising 20 species found in the New World. They are climbing bamboos. [more]
Arundinaria
Small to arborescent bamboos, spreading or loosely clumped. Rhizomes leptomorph. Culms diffuse to pluricaespitose, suberect to drooping, 1-7(-13) m tall, 0.5-4(-6) cm thick; internodes terete to flattened on one side above branches. Branch buds tall, with or without promontory, within 2-keeled prophyll, always open at front. Branches (1 or) 2-5(-7), subequal. Lateral branch axes always subtended by sheaths, without replication of lateral branches. Culm sheaths deciduous to persistent, blade usually recurved or reflexed, lanceolate, articulate. Leaf sheaths persistent; blade oblong-lanceolate or narrowly lanceolate, small to medium-sized, without marginal necrosis in winter, arrangement random, transverse veins distinct. Inflorescence an open panicle or raceme, flowering branches usually subtended by tiny bracts. Spikelets several to many flowered, slender; rachilla internodes extended, disarticulating. Glumes 1 or 2, mucronate; lemma similar to glumes; palea 2-keeled, apex obtuse; lodicules 3. Stamens 3; filaments free, slender; anthers yellow. Style usually very short; stigmas 2 or 3, plumose. Caryopsis dry, oblong. New shoots May-Jun.[1] [more]
Bambusa
Arborescent bamboos, occasionally shrubby or scrambling, 1-20 m. Rhizomes short necked, pachymorph. Culms unicaespitose, erect to pendulous, rarely subscandent; internodes terete; nodes not raised. Branches several to many, often 1-3 dominant (subequal in Bambusa subg. Lingnania), branchlets of lower branches sometimes forming tough or weak thorns. Culm sheaths deciduous, rarely persistent; auricles usually conspicuous, always with marginal oral setae; blade usually erect. Leaf blade variable in size, transverse veins inconspicuous. Inflorescence iterauctant, fully bracteate, subtended by a broad 2-keeled prophyll; pseudospikelets rarely solitary, usually several to many clustered to capitate on flowering branches. Pseudospikelets prophyllate; florets 2 to many, terminal floret sterile or imperfect, sessile; fertile glumes preceded by 1 or more gemmiferous, glumaceous, or spathaceous bracts and/or 1-3 empty glumes; rachilla internodes usually distinct and usually disarticulating with florets, falling separately; lemma broad, many veined; palea 2-keeled, apex acute or shortly bifid; lodicules 3 or 2. Stamens 6; filaments free. Ovary usually stalked, apex thickened and hairy; style solid, usually short; stigmas (1-) 3, long, hairy, plumose. Caryopsis terete, apex hairy; pericarp slightly thickened.[2] [more]
Banisteria
Chimonobambusa
Shrubby bamboos, rarely subarborescent. Rhizomes leptomorph, with running underground stems. Culms usually diffuse, sometimes tillering (pluricaespitose), erect; internodes terete or 4-angled, often basally grooved above branches; nodes prominent to very prominent, basal nodes often with a ring of sparse or dense root thorns; sheath scars usually with a ring of pubescence or persistent base of culm sheath. Branches 3(-7 on upper culm), subequal, buds ovate-triangular, open at front, prophyll reduced. Culm sheaths deciduous and leathery, or sometimes persistent and papery; auricles minute or absent; blade reduced, to 1 cm, narrow. Leaves (1 or) 2-5 per ultimate branch; blade lanceolate, base cuneate. Inflorescence fully bracteate, weakly iterauctant, 1-3 single pseudospikelet racemes loosely fasciculate, subtended by gradually enlarged bracts. Spikelets several to many flowered, sessile. Rachilla disarticulating. Glumes usually 1-3, frequently one subtending a bud; lemma papery or membranous; palea membranous, 2-keeled, obtuse; lodicules 3, membranous. Stamens 3; filaments free. Ovary ellipsoid; style 1, short; stigmas 2 or 3, plumose. Fruit a nutlike caryopsis with a hardened pericarp. New shoots Apr-Nov.[3] [more]
Chusquea
Chusquea is a of bamboo with about 120 species. Most of them are mountain clumping bamboos native from southern Mexico to southern Chile and Argentina. They are sometimes referred to as South American mountain bamboos. Unlike most other bamboos, the stems of these species are solid, not hollow. [more]
Cotoneaster
Shrubs, rarely small trees, erect, decumbent, or prostrate, deciduous, semievergreen, or evergreen. Branchlets mostly terete, rarely slightly angulate, unarmed. Winter buds small; scales several, imbricate, exposed. Leaves alternate, simple, shortly petiolate; stipules caducous, usually subulate, small; margin of leaf blade entire, venation camptodromous. Inflorescences terminal or axillary, cymose or corymbose, sometimes flowers several fascicled or solitary. Hypanthium turbinate or campanulate, rarely cylindric, adnate to ovary. Sepals 5, persistent, short. Petals 5, erect or spreading, imbricate in bud, white, pink, or red. Stamens 10-20(-22), inserted in mouth of hypanthium. Ovary inferior or semi-inferior, 2-5-loculed; carpels 2-5, connate abaxially, free adaxially; ovules 2 per carpel, erect; styles 2-5, free; stigmas dilated. Fruit a drupe-like pome, red, brownish red, or orange to black, with persistent, incurved, fleshy sepals, containing pyrenes; pyrenes (1 or) 2-5, bony, 1-seeded; seeds compressed; cotyledons plano-convex.[4] [more]
Dendrocalamus
Arborescent bamboos, large-sized; clumps dense. Rhizomes short necked, pachymorph. Culms unicaespitose, erect, or occasionally scrambling, apex usually pendulous; internodes terete. Branches several to many, none to 3 dominant. Culm sheaths deciduous; ligule conspicuous; auricles often absent or small; blade usually recurved, or erect. Leaf blades usually large; ligule conspicuous; auricles usually absent; venation not tessellate. Inflorescence iterauctant, fully bracteate, subtended by a narrow single-keeled prophyll, pseudospikelets clustered in soft or spiky globose mass at nodes of leafless flowering branches. Pseudospikelets prophyllate, (1 or) 2-8-flowered, with or without rachilla extension and rudiment, sessile, fertile glumes preceded by 1 or more gemmiferous bracts and 0-2 empty glumes. Prophyll narrow, single-keeled. Rachilla internodes usually abbreviated and not disarticulating; florets falling together. Lemma broad, nearly equal to palea, many veined, sometimes long mucronate; palea of lower florets 2-keeled, but rounded or imperfectly keeled in terminal floret if rachilla extension small or absent, apex acute or shortly bifid; lodicules absent or variably 1-3. Stamens 6; filaments usually free, rarely united into a loose tube. Ovary stalked, apex thickened and hairy; style very short, solid; stigmas 1(-3), long, hairy, plumose. Caryopsis terete, apex hairy; pericarp slightly thickened. 2n = 76*.[5] [more]
Gigantochloa
Arborescent bamboos, large-sized; clumps dense. Rhizomes short necked, pachymorph. Culms unicaespitose, erect, pendulous at apex; internodes green initially, often with yellow stripes, terete. Branches several, 1 dominant. Culm sheaths deciduous, very broad, densely hairy; ligule conspicuous; auricles absent or small; blade recurved or erect. Leaves usually large, base cuneate; auricles usually absent; ligule conspicuous; blade cuneate at base, venation not tessellate. Inflorescence iterauctant, fully bracteate, subtended by a narrow single-keeled prophyll, pseudospikelets clustered in soft or spiky globose mass at nodes of leafless flowering branches. Pseudospikelets sessile, prophyllate; florets (1 or) 2-5, with a sterile terminal floret with lemma only, sessile. Fertile glumes preceded by 1 or more gemmiferous bracts and 0-2 empty glumes; rachilla very short, obscure, not disarticulating; lemma broad, many veined; palea strongly 2-keeled; lodicules absent. Stamens 6; filaments united into a firm tube; anthers apiculate. Ovary stalked, apex thickened and hairy; stigma 1, long, hairy, plumose. Caryopsis terete, apex hairy; pericarp slightly thickened. 2n = 76*.[6] [more]
Guadua
Guadua is a genus of thorny clumping bamboo, ranging from moderate to very large species. Physically, Guadua angustifolia is noted for being the largest Neotropics bamboo. The genus is similar to Bambusa and is sometimes included in Bambusa. [more]
Neurolepis
Neurolepis is a of bamboo. It has 9 species, found in South America (from Venezuela to Peru). It is one of two genera in the subtribe Chusqueinae, the other being Chusquea; unlike that genus the stems of Neurolepis are hollow, like other bamboos. Many of the species are running, rather than clumping, bamboos, and are found cascading down hillsides. [more]
Oxytenanthera
Phyllostachys
Arborescent or shrubby bamboos. Rhizomes leptomorph, with running underground stems. Culms diffuse; internodes profoundly flattened or grooved on one side above branches; nodes 2-ridged. Branches 2, subequal, rarely with a much smaller, central or lateral 3rd branch. Culm sheaths deciduous, papery to subleathery; ligule usually conspicuous; usually auricled with long bristles; blade usually recurved or reflexed. Leaf blade with distinct transverse veins, usually abaxially pilose proximally. Inflorescence bracteate, partially iterauctant, composed of 1-7-spikeleted racemes gathered into fascicles or globose mass subtended by a tiny, membranous, 2-keeled prophyll, 0 or 1 gemmiferous bract, 2-6, gradually enlarged scaly bracts, and 2-7 spathiform bracts. Spikelets with 2-7 florets, terminal sterile. Glumes absent to 1(-3). Rachilla extending beyond uppermost floret, disarticulating just below fertile florets. Lemma variable in size and texture; palea 2-keeled, apex bifid; lodicules 3, ciliate. Stamens 3. Style long; stigmas (1-) 3, plumose. Caryopsis elliptical to linear-lanceolate, dorsally grooved.[7] [more]
Pseudosasa
Plants small, shrublike, or arborescent, spreading and densely clumped; rhizomes leptomorph. Culms pluricaespitose, erect to drooping, 0.5-13 m tall, to 6 cm thick; internodes terete or moderately grooved; nodes not greatly swollen; supra-nodal ridge not evident. Branch buds tall, prophylls 2-keeled, initially closed at front, without promontory. Branches erect, initially 1-3 per node, short or long, central slightly dominant with basal nodes compressed, branches always fully sheathed, without replication of lateral branches, sheaths and prophylls ± glabrous. Culm sheaths deciduous to very persistent, tough; blade erect or reflexed, narrowly triangular to strap-shaped. Leaf sheaths persistent; blades moderately large for size of culm, without marginal necrosis in winter, arrangement random, transverse veins distinct. Inflorescence an open raceme or panicle; branching subtended by large or small bracts. Spikelets 2-20 cm; rachilla sinuous, disarticulation below florets; florets 3-30. Glumes 2, shorter than first lemma; lemma to 1 cm. Palea 2-keeled. Stamens 3. Stigmas 3.[8] [more]
Sasa
Schizostachyum
Schizostachyum is a tall or shrub-like genus of bamboo. The genus name comes from Greek schistos ("cleft") and stachys ("spike"), referring to the spacing of spikelets. [more]
Semiarundinaria
Shrubby bamboo, sometimes subarborescent. Rhizomes leptomorph, with running underground stems. Culms densely pluricaespitose, erect; internodes flattened or grooved above branches, glabrous (pubescent in S. densiflora) ; nodes prominent. Branches (3-) 5-9(-13), subequal, buds initially open at front. Culm sheaths deciduous, leathery or thickly papery; ligule conspicuous; blade recurved or reflexed. Leaves 3-7(-10) per ultimate branch; blade with distinct transverse veins. Inflorescence lateral, racemose to paniculate, fully bracteate, partially iterauctant, prophyllate; pseudospikelets subtended by a spathiform prophyll and 2 or 3 gradually enlarged bracts. Spikelets sessile, 2-7-flowered. Rachilla articulate, internodes extended (short in S. densiflora). Glumes absent to 3; lemma papery, acuminate; palea about as long as or longer than lemma, 2-keeled abaxially, apex rounded, ciliolate; lodicules 3(or 4). Stamens 3; filaments free; anthers exserted. Ovary ellipsoid, ovoid, or globose; style 1; stigmas 3, plumose. Fruit a caryopsis.[9] [more]
At least 40 species and subspecies belong to the Genus Semiarundinaria.
More info about the Genus Semiarundinaria may be found here.
References
- ^ Gratani, Loretta; Maria Fiore Crescente, Laura Varone, Giuseppe Fabrini, and Eleonora Digiulio (2008). "Growth pattern and photosynthetic activity of different bamboo species growing in the Botanical Garden of Rome". Flora 203: 77–84. .
- ^ Bystriakova, N.; N. Bystriakova, V. Kapos, I. Lysenko and C.M.A. Stapleton (September 2003). "Distribution and conservation status of foret bamboo biodiversity in the Asia-Pacific Region". Biodiversity and Conservation 12 (9): 1833–1841. doi:
- ^ "Arundinaria gigantea (Walt.) Muhl. giant cane". PLANTS Database. USDA. http://plants.usda.gov/java/profile?symbol=ARGI.
- ^ Huxley, ed (1992). New RHS Diction ary of Gardening. Macmillan New RHS Dictionary of Gardening. ISBN 0-333-47494-5.
- ^ Farrelly, David (1984). The Book of Bamboo. Sierra Club Books. ISBN 087156825X.
- ^ a b Thomas R. Soderstrom; Cleofe E. Calderon; Thomas R. Soderstrom; Cleofe E. Calderon; T.R. Soderstrom, C.E. Calderon (1979). "A Commentary on the Bamboos (Poaceae: Bambusoideae)". Biotropica 11 (3): 161–172. doi:
- ^ a b Janzen, DH. (1976). "Why Bamboos Wait so Long to Flower". Annual Review of Ecology and Systematics 7: 347–391. doi:
- ^ "Mast flowering and semelparity in bamboos: The bamboo fire cycle hypothesis". American Naturalist (154): 383–391. 1999.
- ^ "The Bamboo Fire Cycle Hypothesis: A Coment". The American Naturalist 6 (158): 659–663. 2001.
- < b>^ "On incorporating fire into our thinking about natural ecosystems: A response to Saha and Howe". American Naturalist (158): 664–670. 2001.
- ^ "Gorillas get drunk on bamboo sap". Telegraph.co.uk. 23 March 2009. http://www.telegraph.co.uk/earth/earthpicturegalleries/5037343/Gorillas-get-drunk-on-bamboo-sap.html. Retrieved on 12 August 2009.
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- ^ Puri, H. S. (2003). Rasayana ayurvedic herbs for longevity and rejuvenation. London: Taylor & Francis. pp. 71–73. ISBN 0203216563. http://books.google.com/books?id=aQh25X9mzjAC&lpg=PP1&dq=Rasa yana%20Ayurvedic%20Herbs%20for%20Longevity%20and%20Rejuvenation&pg=PP1#v=onepage&q=&f=false. Retrieved on 12 August 2009.
- ^ Peters, Tom F. (1987). Transitions in Engineering: Guillaume Henri Dufour and the Early 19th Century Cable Suspension Bridges. Birkhauser. ISBN 3764319291. http://books.google.com/books?id=73JPiTuDYscC.
- ^ Landler, Mark (27 March 2002). "Hong Kong Journal; For Raising Skyscrapers, Bamboo Does Nicely". New York Times. http://www.nytimes.com/2002/03/27/world/hong-kong-journal-for-raising-skyscrapers-bamboo-does-nicely.html. Retrieved on 12 August 2009.
- ^ Bamboo In Japan. Kodansha International. 1987. p. 101. ISBN 4770025106. http://books.google.com/books?id=UukQ2LaaP0wC&pg=PA101&lpg=PA101.
- ^ Roger Lewis (1 July 2006). "Square Bamboo". LewisBamboo.com. http://www.lewisbamboo.com/square.html. Retrieved on 12 August 2009.
- ^ CASSANDRA ADAMS. "Bamboo Architecture and Construction with Oscar Hidalgo". Natural Building Colloquium. http://www.networkearth.org/naturalbuilding/bamboo.html. Retrieved on 11 August 2009.
- ^ a b c d e {cite web |title=Bamboo Boom: Is This Material for You? |url=http://www.scientificamerican.com/article.cfm?id=bamboo-boom| author= Michelle Nijhuis |date= June 2009 |work=Scientific American Earth 3.0 special |publisher=Scientific American |accessdate=11 August 2009}}
- ^ Jonathan Bardelline (9 July 2009). "Growing the Future of Bamboo Products". GreenBiz.com. http://www.greenbiz.com/feature/2009/07/09/growing-future-bamboo-products. Retrieved on 11 August 2009.
- ^ "Bamboo Construction". CD3WD. http://www.fastonline.org/CD 3WD_40/VITA/BAMBOO/EN/BAMBOO.HTM. Retrieved on 11 August 2009.
- ^ Bamboo as a Building Material. Washington D.C.: US Department of Agriculture. 1981. pp. 7–11. http://www.eric.ed.gov/ERICDocs/data/ericdocs2sql/content_storage_01/0000019b/80/34/73/06.pdf. Retrieved on 11 August 2009.
- ^
"Bamboo Fiber: Greenwash or Treasure?". Feelgood Style. 26 June 2008. http://feelgoodstyle.com/2008/06/26/bamboo-fiber-greenwash-or-treasure/. Retrieved on 12 August 2009. - ^ "Competition Bureau Calls on Textile Dealers to Accurately Label Textile Articles Derived from Bamboo". Reuters. 11 March 2009. http://www.reuters.com/article/pressRelease/idUS175478+11-Mar-2009+MW20090311. Retrieved on 12 August 2009.
- ^ a b "Four Companies Charged with Labeling Rayon Clothing As Bamboo". GreenBiz.com. 11 August 2009. http://www.greenbiz.com/news/2009/08/11/companies-label-rayon-clothing-bamboo. Retrieved on 12 August 2009.
- ^ Chan, Alan Kam-leung and Gregory K. Clancey, Hui-Chieh Loy (2002). Historical Perspectives on East Asian Science, Technology and Medicine. Singapore: Singapore University Press. ISBN 9971692597. p. 15.
- ^ Needham, Joseph (1986). Science and Civilization in China: Volume 3, Mathematics and the Sciences of the Heavens and the Earth. Taipei: Caves Books, Ltd. p. 614.
- ^ Loewe, Michael (1997). "Wood and bamboo administrative documents of the Han period". in Edward L. Shaughnessy. New Sources of Early Chinese History. Society for the Study of Early China. pp. 161–192. ISBN 1-55729-058-X.
Bibliography
- Yü Te-tsun, Lu Ling-ti, Ku Tsue-chih, Li Chao-luan, Kuan Ke-chien & Chiang Wan-fu. 1974, 1985, 1986. Rosaceae. In: Yü Te-tsun, ed., Fl. Reipubl. Popularis Sin. 36: 1443; 37: 1516; 38: 1133.
Footnotes
- Zheng-de Zhu, De-Zhu Li & Chris Stapleton "Arundinaria". in Flora of China Vol. 22 Page 9, 112. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org.
- Nianhe Xia, Liang-zhi Jia, De-Zhu Li & Chris Stapleton "Bambusa". in Flora of China Vol. 22 Page 8, 9, 41, 121, 148. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org.
- De-Zhu Li & Chris Stapleton "Chimonobambusa". in Flora of China Vol. 22 Page 9, 100, 152. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org.
- Lu Ling-ti, Anthony R. Brach "Cotoneaster". in Flora of China Vol. 9 Page 85. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org.
- De-Zhu Li & Chris Stapleton "Dendrocalamus". in Flora of China Vol. 22 Page 9, 37, 39, 98, 100. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org.
- De-Zhu Li & Chris Stapleton "Gigantochloa". in Flora of China Vol. 22 Page 9, 46, 48. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org.
- Zheng-ping Wang & Chris Stapleton "Phyllostachys". in Flora of China Vol. 22 Page 9, 114, 163. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org.
- Zheng-de Zhu, De-Zhu Li & Chris Stapleton "Pseudosasa". in Flora of China Vol. 22 Page 9, 115, 124. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org.
- De-Zhu Li & Chris Stapleton "Semiarundinaria". in Flora of China Vol. 22 Page 9, 151. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org.
Sources
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