Overview
Asterales is an of dicotyledonous flowering plants that includes the composite family (Asteraceae) and its related families.
The order is a cosmopolite(a cosmopolitan plant), and includes mostly herbaceous species, although a small number of trees (such as some members of the genus Lobelia) and shrubs are also present.
The Asterales can be characterized on the morphological and molecular level. Synapomorphies(a character that is shared by 2 or more groups through growth) include the oligosaccharide inulin, a nutrient storage molecule, and unique stamen morphology. The stamens are usually found around the style, either aggregated densely or fused into a tube, probably an adaptation in association with the plunger (or secondary) pollination that is common among the families of the order.
Asterales belong to organisms that has one common ancestor. Their families are linked because of their genes, chemical characteristics,and having a special mechanism called plunger pollinatio n or secondary pollen presentation.
The order Asterales includes about eleven families, the largest of which are the Asteraceae, with about 25,000 species, and the Campanulaceae, with about 2,000 species. The remaining families count together for less than 500 species. The two large families are cosmopolitan, with many of their species found in the northern hemisphere, and the smaller families are usually confined to Australia and the adjacent areas, or sometimes South America.
Under the Cronquist system, Asteraceae was the only family in the group, but newer systems (e. g. APG II) have expanded it.
Pyrethrum (plant in the chrysanthemum family) is an insecticide that does not cause environmental problems. Chriysanthemum is a specie that has an environmental impact and grows from woormwood(a bitter plant used as an ingredient in absinthe). Absinthium is a source that is poisonous oil used to give the linqueur ab sinthe a different character.
Evolution and biogeography
The Asterales order probably originated in the Cretaceous on the supercontinent Gondwana, in the area that is now Australia and Asia. Although most extant species are herbaceous, the examination of the basal families in the order suggests that the common ancestor of the order was an arborescent plant.
Fossil evidence of the Asterales is rare and belongs to rather recent epochs, so the precise estimation of the order's age is quite difficult. An Oligocene pollen is known for Asteraceae and Goodeniaceae, and seeds from Oligocene and Miocene are known for Menyanthaceae and Campanulaceae respectively.
(Bremer and Gustafsson, 1997)
Economical importance
The Asteraceae include some species grown for food, including sunflower (Helianthus annuus), lettuce (Lactuca sativa) and ch icory (Cichorium). Many spices and medicinal herbs are also present.
Of horticultural importance are the Asteraceae (e. g. chrysanthemum) and Campanulaceae.
b>Asterales is an of dicotyledonous flowering plants that includes the composite family (Asteraceae) and its related families.The order is a cosmopolite(a cosmopolitan plant), and includes mostly herbaceous species, although a small number of trees (such as some members of the genus Lobelia) and shrubs are also present.
The Asterales can be characterized on the morphological and molecular level. Synapomorphies(a character that is shared by 2 or more groups through growth) include the oligosaccharide inulin, a nutrient storage molecule, and unique stamen morphology. The stamens are usually found around the style, either aggregated densely or fused into a tube, probably an adaptation in association with the plunger (or secondary) pollination that is common among the families of the order.
Asterales belong to o rganisms that has one common ancestor. Their families are linked because of their genes, chemical characteristics,and having a special mechanism called plunger pollination or secondary pollen presentation.
The order Asterales includes about eleven families, the largest of which are the Asteraceae, with about 25,000 species, and the Campanulaceae, with about 2,000 species. The remaining families count together for less than 500 species. The two large families are cosmopolitan, with many of their species found in the northern hemisphere, and the smaller families are usually confined to Australia and the adjacent areas, or sometimes South America.
Under the Cronquist system, Asteraceae was the only family in the group, but newer systems (e. g. APG II) have expanded it.
Pyrethrum (plant in the chrysanthemum family) is an insecticide that does not cause environmental problems. Chriysanthemum is a specie that has an environmental impact and grows from woormwood(a bitter plant used as an ingredient in absinthe). Absinthium is a source that is poisonous oil used to give the linqueur absinthe a different character.
Evolution and biogeography
The Asterales order probably originated in the Cretaceous on the supercontinent Gondwana, in the area that is now Australia and Asia. Although most extant species are herbaceous, the examination of the basal families in the order suggests that the common ancestor of the order was an arborescent plant.
Fossil evidence of the Asterales is rare and belongs to rather recent epochs, so the precise estimation of the order's age is quite difficult. An Oligocene pollen is known for Asteraceae and Goodeniaceae, and seeds from Oligocene and Miocene are known for Menyanthaceae and Campanulaceae respectively.
(Bremer and Gustafsson, 1997)
Economical importance
The Asteraceae include some species grown for food, including sunflower (Helianthus annuus), lettuce (Lactuca sativa) and chicory (Cichorium). Many spices and medicinal herbs are also present.
Of horticultural importance are the Asteraceae (e. g. chrysanthemum) and Campanulaceae.
References
- K. Bremer, M. H. G. Gustafsson (1997). East Gondwana ancestry of the sunflower alliance of families. Proceedings of the National Academy of Sciences U.S.A. 94, 9188-9190. (Available online: Abstract | Full text (HTML) | Full text (PDF))
- W. S. Judd, C. S. Campbell, E. A. Kellogg, P. F. Stevens, M. J. Donoghue (2002). Plant Systematics: A Phylogenetic Approach, 2nd editio n. pp. 476?486 (Asterales). Sinauer Associates, Sunderland, Massachusetts. ISBN 0-87893-403-0.
- J. Lindley (1833). Nixus Plantarum, 20. Londini.
- Smissen, R. D. (December 2002). Asterales (Sunflower). In: Nature Encyclopedia of Life Sciences. Nature Publishing Group, London. (Available online: DOI | ELS site)
- "Asterales (plant Order) -- Britannica Online Encyclopedia." Encyclopedia - Britannica Online Encyclopedia. Web. 19 Jan. 2012. <http://www.britannica.com/EBchecked/topic/39703/Asterales>.
- "Asterales - Definition and More from the Free Merriam-Webster Dictionary." Dictionary and Thesaurus - Merriam-Webster Online. Web. 19 Jan. 2012. <http://www.merriam-webster.com/di ctionary/asterales>.
Taxonomy
The Order Asterales is further organized into finer groupings including:
- Family (12): Alseuosmiaceae · Argophyllaceae · Asteraceae · Calyceraceae · Campanulaceae · Compositae · Goodeniaceae · Menyanthaceae · Pentaphragmataceae · Phellinaceae · Rousseaceae · Stylidiaceae
- Species: ZipcodeZoo has pages for 89,879 species, subspecies, varieties, forms, and cultivars in the Order Asterales.
Families
Alseuosmiaceae
Argophyllaceae
The Argophyllaceae is a family of shrubs or small trees belonging to the order Asterales. The family includes two genera Argophyllum and Corokia. Members of the family are native to eastern Australia, New Zealand, Lord Howe Island, New Caledonia, and Rapa Iti. [more]
Asteraceae
The Asteraceae or Compositae (commonly referred to as the aster, daisy, or sunflower family), is an exceedingly large and widespread family of vascular plants. The group has more than 22,750 currently accepted species, spread across 1620 genera and 12 subfamilies. Along with the Orchidaceae, this makes it one of the two largest flowering plant families in the world. However, selecting which of the two families is larger has yet to be been done conclusively, owing to the uncertainty around exactly how many species are in each group. The largest composite genera are Senecio (1,000 species), Vernonia (1,000 species), Centaurea (700 species), Cousinia (600 species), Helichrysum (550 species), and Artemesia (550 species). [more]
Calyceraceae
Calyceraceae is a plant family in the order Asterales. [more]
Campanulaceae
The family Campanulaceae (also bellflower family), of the order Asterales, contains about 2000 species in 70 genera of herbaceous plants, shrubs, and rarely small trees, often with milky non-toxic sap. Among them are the familiar garden plants Campanula (bellflower), Lobelia, and Platycodon (balloonflower). [more]
Compositae
Goodeniaceae
Goodeniaceae are a family of flowering plants in the order Asterales. It contains about 404 species in twelve genera. The family is distributed mostly in Australia, except for the genus Scaevola, which is pantropical. Its species are found across most of Australia, being especially common in arid and semi-arid climates. [more]
Menyanthaceae
Menyanthaceae are a family of aquatic and wetland plants in the order Asterales. There are approximately 60-70 species in five genera distributed worldwide. The simple or compound leaves arise alternately from a creeping rhizome. In the submersed aquatic genus Nymphoides, leaves are floating and support a lax, umbellate or racemose inflorescence. In other genera the inflorescence is erect and consists of one (e.g., Liparophyllum) to many flowers. The sympetalous, insect-pollinated flowers are five-parted and either yellow or white. The petals are ciliate or adorned with lateral wings. Fruit type is a capsule. [more]
Pentaphragmataceae
Phellinaceae
Rousseaceae
Stylidiaceae
The family Stylidiaceae is a taxon of dicotyledonous flowering plants. It consists of five genera with over 240 species, most of which are endemic to Australia and New Zealand. Members of Stylidiaceae are typically grass-like herbs or small shrubs and can be perennials or annuals. Most species are free standing or self-supporting, though a few can be climbing or scrambling (Stylidium scandens uses leaf tips recurved into hooks to climb). [more]
At least 644 species and subspecies belong to the Family Stylidiaceae.
More info about the Family Stylidiaceae may be found here.
References
- K. Bremer, M. H. G. Gustafsson (1997). East Gondwana ancestry of the sunflower alliance of families. Proceedings of the National Academy of Sciences U.S.A. 94, 9188-9190. (Available online: Abstract | Full text (HTML) | Full text (PDF))
- W. S. Judd, C. S. Campbell, E. A. Kellogg, P. F. Stevens, M. J. Donoghue (2002). Plant Systematics: A Phylogenetic Approach, 2nd edition. pp. 476?486 (Asterales). Sinauer Associates, Sunderland, Massachusetts. ISBN 0-87893-403-0.
- J. Lindley (1833). Nixus Plantarum, 20. Londini.
- Smissen, R. D. (December 2002). Asterales (Sunflower). In: Nature Encyclopedia of Life Sciences. Nature Publishing Group, London. (Available online: DOI | ELS site)
- "Asterales (plant Order) -- Brita nnica Online Encyclopedia." Encyclopedia - Britannica Online Encyclopedia. Web. 19 Jan. 2012. <http://www.britannica.com/EBchecked/topic/39703/Asterales>.
- "Asterales - Definition and More from the Free Merriam-Webster Dictionary." Dictionary and Thesaurus - Merriam-Webster Online. Web. 19 Jan. 2012. <http://www.merriam-webster.com/dictionary/asterales>.
Sources
- The text on this page is licensed under the GNU Free Documentation License. It includes material from Wikipedia retrieved Wednesday, April 25, 2012.
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