Overview
Trees or shrubs, deciduous. Winter buds large, viscid resinous or not, with several pairs of imbricate scales; scales abaxially glabrous or sparsely puberulent. Leaf blade 5-11-foliolate; leaflet blades without scattered, conspicuous glands, margin crenate to serrate or compoundly so. Thyrse cylindric or conic; branches simple; bracts absent. Flowers often large and showy. Sepals connate to form a tubular to campanulate calyx tube. Petals often unequal, base clawed, limb obovate, oblong, oblanceolate, or spatulate. Ovary without a gynophore; style long, slender; stigma depressed globose, entire or obscurely lobed. Capsule depressed globose to pyriform, without a long gynophore, often 1-seeded; pericarp usually smooth, often dotted, rarely verrucose or prickly. Seeds depressed globose to pyriform, large (2-7 cm) ; testa brown; hilum large, pale, occupying 1/3-1/2 of seed. x = 20.
Twelve species: mainly in North America (United States, Baja California in Mexico) and Asia from the Himalayas to Japan (Afghanistan, Bangladesh, Bhutan, China, India, Japan, Kashmir, Laos, Myanmar, Nepal, Pakistan, Thailand, Vietnam), one species in SE Europe (Albania, Bulgaria, Greece, former Yugoslavia) ; four species (one endemic, two introduced) in China.
Fang (Acta Sci. Nat. Univ. Szechuan. 1960(3) : 77-125. 1962) published a revision of the Hippocastanaceae in China, including 12 new species, for which six of the names were not validly published, either because no Latin description was provided, no type was indicated, or more than one type was indicated. Fang later (in FRPS) identified three of these entities as species of Araliaceae: "Aesculus chingsiensis" as Acanthopanax evodiifolius Franchet, "Aesculus chinpingensis" as Brassaiopsis glomerulata (Blume) Regel, and "Aesculus kwangsiensis" as Schefflera octophylla (Loureiro) Harms.
The leaves, flowers, and large seeds contain the poisonous compound aesculin.[1]
Photos
Taxonomy
- Domain: Eukaryota
Whittaker & Margulis,1978 - eukaryotes
- Kingdom: Plantae
Haeckel, 1866
- Subkingdom: Viridaeplantae
Cavalier-Smith, 1981 - Green Plants
- Phylum: Tracheophyta
Sinnott, 1935 Ex Cavalier-Smith, 1998 - Vascular Plants
- Subphylum: Euphyllophytina
- Infraphylum: Radiatopses
Kenrick & Crane, 1997
- Class: Magnoliopsida
Brongniart, 1843 - Dicotyledons
- Subclass: Rosidae
Takhtajan, 1967
- Superorder: Rutanae
Takhtajan, 1967
- Order: Sapindales
Dumortier, 1829
- Family: Hippocastanaceae
(ES-kew-lus)
A.p. De Candolle, 1824
- Subfamily: Hippocastanoideae
- Family: Hippocastanaceae
(ES-kew-lus)
A.p. De Candolle, 1824
- Order: Sapindales
Dumortier, 1829
- Superorder: Rutanae
Takhtajan, 1967
- Subclass: Rosidae
Takhtajan, 1967
- Class: Magnoliopsida
Brongniart, 1843 - Dicotyledons
- Infraphylum: Radiatopses
Kenrick & Crane, 1997
- Subphylum: Euphyllophytina
- Phylum: Tracheophyta
Sinnott, 1935 Ex Cavalier-Smith, 1998 - Vascular Plants
- Subkingdom: Viridaeplantae
Cavalier-Smith, 1981 - Green Plants
- Kingdom: Plantae
Haeckel, 1866
The Genus Aesculus is further organized into finer groupings including:
- Species: ZipcodeZoo has pages for 195 species, subspecies, varieties, forms, and cultivars in the Genus Aesculus: A. 'Autumn Splendor' (Ohio Buckeye) · A. 'Dallimorei' · A. alba · A. antiquorum · A. arguta · A. arguta var. arguta · A. arnoldiana (Arnold's Chestnut) · A. assamica · A. austrina · A. bushii (Bush's Chestnut) · A. californica (California Buckeye) · A. californica 'Blue Haze' · A. californica 'Canyon Pink' · A. canadensis · A. carnea (Pink Horse Chestnut) · A. castanea · A. chekiangensis · A. chinensis (Chinese Horsechestnut) · A. chinensis var. chekiangensis · A. chinensis var. wilsonii · A. chuniana · A. dallimorei · A. discolor · A. discolor Pursh var. mollis (Raf.) Sarg. · A. discolor var. flavescens · A. dissimilis · A. du-pontii · A. echinata · A. flava (Yellow Buckeye) · A. flava f. vestita · A. flava × pavia · A. georgiana · A. georgina · A. glabra (Texas Buckeye) · A. glabra 'April Wine' (Buckeye) · A. glabra 'Autumn Blaze' · A. glabra 'Fall Red' (Ohio Buckeye) · A. glabra 'Klein's Weeping' (Ohio Buckeye) · A. glabra 'October Red' · A. glabra arguta · A. glabra f. pallida · A. glabra forma pallida (Texas Buckeye) · A. glabra subsp. arguta · A. glabra var. arguta (Western Buckeye) · A. glabra var. glabra (Ohio Buckeye) · A. glabra var. Nana · A. glabra Willd. var. glabra · A. glabra Willd. var. leucodermis Sarg. · A. glabra Willd. var. monticola Sarg. · A. glabra Willd. var. sargentii Rehder · A. glaucescens · A. harbisonii · A. hickeyi · A. hippocastan · A. hippocastaneum · A. hippocastanum (European Horse Chestnut) · A. hippocastanum 'Aureomarginata' · A. hippocastanum 'Baumannii' (Baumanns Horse Chestnut) · A. hippocastanum 'Digitata' · A. hippocastanum 'Flore Pleno' · A. hippocastanum 'Glabra' · A. hippocastanum 'Globosa' · A. hippocastanum 'Hampton Court Gold' · A. hippocastanum 'Honiton Gold' · A. hippocastanum 'Laciniata' (Horse Chestnut) · A. hippocastanum 'Monstrosa' · A. hippocastanum 'Pyramidalis' · A. hippocastanum 'Umbraculifera' · A. hippocastanum 'Variegata' · A. hippocastanum 'Wisselink' · A. hippocastanum f. memmingeri · A. hippocastanum L. 'Henkelii' · A. hippocastanum L. 'Incisa' · A. hippocastanum L. 'Monstrosum' · A. hippocastanum L. 'Plena' · A. hippocastanum L. var. asplenifolia · A. hippocastanum L. var. pumila Dippel · A. humilis · A. hybrida (Hybrid Chestnut) · A. hyppocastanum · A. incisa · A. indica (Indian Horse Chesnut) · A. indica 'Sydney Pearce' · A. indica var. concolor · A. intermedia · A. khassyana · A. lantsangensis · A. livida · A. lucida · A. lutea · A. lyonii · A. macrostachya · A. macrostachys · A. marylandica (Maryland Chestnut) · A. maxima · A. megaphylla · A. memmingeri · A. mexicana · A. michauxii · A. microcarpa · A. mississippiensis · A. mollis · A. montanus · A. muricata · A. mutabilis (Chestnut) · A. neglecta (Chestnut) · A. neglecta var. tomentosa · A. ochroleuca · A. octandra · A. octandra var. hybrida · A. odorata · A. ohioensis · A. pallida · A. parryi · A. parviflora (Small-Flowered Buckeye) · A. parviflora f. serotina (Bottlebrush Buckeye) · A. parviflora var. serotina (Bottlebrush Buckeye) · A. parviflora var. serotina 'Rogers' (Bottlebrush Buckeye) · A. pavia (Firecracker Plant) · A. pavia 'Atrosanguinea' · A. pavia 'Humilis' · A. pavia 'Penduliflora' · A. pavia 'Purple Spring' · A. pavia 'Rosea Nana' · A. pavia L. 'Arguta' · A. pavia L. var. discolor (Pursh) Torr. & Gray · A. pavia L. var. discolor (Pursh) Torr. & Gray 'Koehnei' · A. pavia L. var. flavescens (Sarg.) Correll (Red Buckeye) · A. pavia L. var. nana Dippel · A. pavia L. var. pavia · A. pavia L. var. whitleyi hort. · A. pavia var. atrosanguinea · A. pavia var. discolor 'Koehnei' · A. pavia var. flavescens (Yellow Buckeye) · A. pavia var. humilis · A. pavia var. nana · A. pavia var. pavia (Red Buckeye) · A. pawia · A. plantierensis · A. polyneura · A. polyneura var. dongchuanensis · A. procera · A. pubescens · A. punduana · A. rosea · A. rubella · A. rubicunda · A. rubra · A. septenata · A. silvatica · A. sinensis · A. splendens · A. sylvatica (Oainted Buckeye) · A. sylvatica W.Bartram var. lanceolata (Sarg.) Sarg. · A. sylvatica W.Bartram var. pubescens (Sarg.) Sarg. · A. sylvatica W.Bartram var. tomentosa Sarg. · A. tsiangii · A. turbinata (Japanese Horse Chestnut) · A. turbinata var. pubescens · A. versicolor · A. wangii · A. wangii var. rupicola · A. whitleyi · A. wilsonii · A. woerlitzensis · A. woerlitzensis Koehne var. ellwangeri Rehder · A. woerlitzensis var. ellwangeri · A. worlitzensis (Worlitz's Chestnut) · A. × arnoldiana (Arnold's Chestnut) · A. × arnoldiana 'Autumn Splendor' · A. x carnea (Pink Horse Chestnut) · A. x carnea 'Aureomarginata' · A. x carnea 'Briotii' (Ruby Red Horsechestnut) · A. x carnea 'Fort Mcnair' (Fort Mcnair Red Horse Chestnut) · A. × carnea 'Marginata' · A. x carnea 'O'Neill Red' (O'neill Red Horse Chestnut) · A. × carnea 'O'Neill' · A. x carnea 'O'neill's Red' (Bottlebrush Buckeye) · A. x carnea 'Plantierensis' (Bottlebrush Buckeye) · A. x carnea 'Rosea' (Bottlebrush Buckeye) · A. × carnea 'Rubro-petiolata' · A. × carnea 'Variegata' · A. × dallimorei · A. x dupontii · A. × glaucescens · A. × mississippiensis · A. × mutabilis 'Harbisonii' · A. x mutabilis 'Induta' (Chestnut) · A. × mutabilis 'Penduliflora' · A. × neglecta 'Autumn Fire' · A. x neglecta 'Erythroblastos' (Yellow Horsechestnut) · A. × plantierensis · A. x woerlitzensis · A. x worlitzensis (Worlitz's Chestnut) · A. xmarylandica
Bibliography
- Fang Wen-pei. 1981. Hippocastanaceae. In: Fang Wen-pei, ed., Fl. Reipubl. Popularis Sin. 46: 274-289
- Lo Hsien-shui & Chen Te-chao. 1985. Handeliodendron. In: Law Yuh-wu & Lo Hsien-shui, eds., Fl. Reipubl. Popularis Sin. 47(1): 61-62.
Footnotes
- Nianhe Xia & Nicholas J. Turland "Aesculus". in Flora of China Vol. 12 Page 1, 2, 6. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org.
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