Humulus

(Genus)

Herbs, taprooted annuals or rhizomatous perennials, rightward-twining. Stems usually branched, armed with rigid 2-branched, stalked hairs that facilitate climbing. Leaves simple; petioles often twining, with 2-branched hairs.Leaf blade mostly cordate, palmately lobed, sometimes unlobed; surfaces abaxially resin-dotted and×or gland-dotted. Inflorescences: staminate inflorescences axillary and terminal, cymose panicles, erect to pendent, (10-) 20-100+-flowered, flowers small; pistillate axillary, spikes or racemes, flowers solitary or paired, short-pedicellate, subtended by bracts and bracteoles. Staminate and pistillate flowers usually on different plants. Achenes lenticular or terete, ensheathed by brownish or sometimes mottled persistent perianth; embryo coiled. x = 10.

Species 3: temperate Northern Hemisphere.^[1]

Photos

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Humulus japonicus Humulus japonicus Martin County, KY Photographer: Chris Evans
Location Taken: United States Copyright: © Chris Evans. Permissions for Use: see www.ForestryImages.org. ZipcodeZoo Photo ID: 176526

Humulus japonicus Humulus japonicus Embryo drawing of Humulus japonicus Photographer: Robert J. Gibbons ZipcodeZoo Photo ID: 148549

Humulus japonicus Humulus japonicus The down-curved bracts and sharp prickles help to identify this plant Photographer: Chris Evans
Location Taken: United States Copyright: © Chris Evans. Permissions for Use: see www.ForestryImages.org. ZipcodeZoo Photo ID: 176525

Humulus japonicus Humulus lupulus var. pubescens Photographer: The William and Lynda Steere Herbarium Date Taken: 2007
Location Taken: Grays Harbour, Grays Harbor Co, Washington, United States of America Copyright: © 2007 This image belongs to The New York Botanical Garden Virtual Herbarium. ZipcodeZoo Photo ID: 210420

Humulus japonicus Humulus lupulus var. lupuloides Photographer: The William and Lynda Steere Herbarium Date Taken: 2007
Location Taken: line: S of mouth of Venator Canyon ca 7 airmiles NNE of town of Alkali Lake, Lake-Harney Co, Oregon, United States of America Copyright: © 2007 This image belongs to The New York Botanical Garden Virtual Herbarium. ZipcodeZoo Photo ID: 210419

Humulus japonicus Humulus lupulus Hop, Fairlawne estate, W. Kent, England. 25 Aug. 2001 Photographer: David W. Mitchell Copyright: © David W. Mitchell ZipcodeZoo Photo ID: 293825

Map

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Taxonomy

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Domain: Eukaryota Whittaker & Margulis,1978 - eukaryotes
- Kingdom: Plantae Haeckel, 1866 - Plants
  - Subkingdom: Viridaeplantae Cavalier-Smith, 1981 - Green Plants
    - Phylum: Tracheophyta Sinnott, 1935 ex Cavalier-Smith, 1998 - Vascular Plants
      - Subphylum: Spermatophytina (auct.) Cavalier-Smith, 1998 - Seed Plants
        Infraphylum: Angiospermae auct.
        Class: Magnoliopsida Brongniart, 1843 - Dicotyledons
        Subclass: Dilleniidae Takhtajan, 1967
        Superorder: Urticanae Takhtajan ex Reveal, 1992
        Order: Urticales Dumortier, 1829
        Family: Cannabaceae Endlicher, 1837 - Hemp Family
        Subfamily: Rhododendroideae
        Tribe: Rhododendreae
        Genus: Humulus (HUM-uh-lus) Linnaeus, Sp. Pl. 2: 1028. 1753; Gen. Pl. ed. 5, 453, 1754. - Hop, houblon [Latin humulus, applied to hop plant]

The Genus Humulus is further organized into finer groupings including:

Species: ZipcodeZoo has pages for 49 species, subspecies, varieties, forms, and cultivars in the Genus Humulus: H. aculeatus · H. cordifolius · H. florissantellus · H. HC243 · H. japonicus (Japanese Hop) · H. japonicus 'Variegatus' (Variegated Japanese Hops) · H. luplus · H. lupulus (Common Hop) · H. lupulus 'Admiral' · H. lupulus 'Aureus' (Aureus Golden Hops) · H. lupulus 'Bramling Cross' · H. lupulus 'Cascade' (Hops) · H. lupulus 'Columbus' · H. lupulus 'First Gold' · H. lupulus 'Fuggle' (Hops) · H. lupulus 'Golden Tassels' · H. lupulus 'Hallertauer' (Hops) · H. lupulus 'Herald' · H. lupulus 'Hip-hop' · H. lupulus 'Northern Brewer' (Hops) · H. lupulus 'Nugget' (Hops) · H. lupulus 'Phoenix' · H. lupulus 'Pioneer' · H. lupulus 'Prima Donna' · H. lupulus 'Progress' · H. lupulus 'Sunbeam' (Golden Hops) · H. lupulus 'Taff's Variegated' · H. lupulus 'WGV' · H. lupulus 'Wye Challenger' · H. lupulus 'Wye Northdown' · H. lupulus 'Wye Target' · H. lupulus americanus (Common Hop) · H. lupulus Goldings Group · H. lupulus L. var. cordifolius Maxim. · H. lupulus L. var. lupuloides E.Small (Arizona Hops) · H. lupulus L. var. neomexicanus A.Nelson & Cockerell (Common Hop) · H. lupulus L. var. pubescens E.Small · H. lupulus var. fengxianensis · H. lupulus var. lupuloides · H. lupulus var. lupulus · H. lupulus var. neomexicanus (Common Hop) · H. lupulus var. pubescens · H. maericanus · H. neomexicanus · H. rotundatus · H. scabrellus · H. volubilis · H. vulgaris · H. yunnanensis

Bibliography

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Small, E. 1978. A numerical and nomenclatural analysis of morpho-geographic taxa of Humulus. Syst. Bot. 3: 37-76.
Small, E. 1980. The relationships of hop cultivars and wild variants of Humulus lupulus. Canad. J. Bot. 58: 676-686.

Footnotes

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"Humulus". in Flora of North America Vol. 3. Oxford University Press. Online at EFloras.org.

Germany 29,000 tonnes (63,934,056 pounds)
USA 26,180 tonnes (57,717,020 lbs)
China 20,000 tonnes (44,092,452 lbs)
Czech Republic 6,800 tonnes (14,991,434 lbs)
Poland 3,355 tonnes (7,396,509 lbs)
Australia 2,000 tonnes (4,409,245 lbs)
North Korea 2,000 tonnes (4,409,245 lbs)
UK 2,000 tonnes (4,409,245 lbs)
Slovenia 1,500 tonnes (3,306,934 lbs)
France 1,400 tonnes (3,086,472 lbs)

Sources

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Last Revised: May 30, 2008