Overview
Taxonomy
The Branch Protostomia is a member of the Subkingdom Bilateria. Here is the complete "parentage" of Protostomia:
- Domain: Eukaryota
Whittaker & Margulis,1978 - eukaryotes
- Kingdom: Animalia
C. Linnaeus, 1758 - animals
- Subkingdom: Bilateria
(Hatschek, 1888) Cavalier-Smith, 1983 - bilaterians
- Branch: Protostomia Grobben, 1908 - protostomes
- Subkingdom: Bilateria
(Hatschek, 1888) Cavalier-Smith, 1983 - bilaterians
- Kingdom: Animalia
C. Linnaeus, 1758 - animals
The Branch Protostomia is further organized into finer groupings including:
- Infrakingdom (9): Coelenterata · Ecdysozoa · Excavata · Haptista · Heterokonta · Lophotrochozoa · Platyzoa · Rhizaria · Spongiaria
- Superphylum (6): Articulata · Aschelminthes · Eutrochozoa · Gnathifera · Lophophorata · Panarthropoda
- Phylum (30): Acanthocephala · Acoelomorpha · Annelida · Arthropoda · Brachiopoda · Bryozoa · Chaetognatha · Cnidaria · Cycliophora · Echiura · Entoprocta · Euglenozoa · Gastrotricha · Gnathostomulida · Haptophyta · Kinorhyncha · Loricifera · Mollusca · Nematoda · Nematomorpha · Nemertea · Ony chophora · Oomycota · Platyhelminthes · Porifera · Priapulida · Radiozoa · Rotifera · Sipuncula · Tardigrada
Phyla
Acanthocephala
Acanthocephala (Greek , akanthos, thorn + ?efa??, kephale, head) is a phylum of parasitic worms known as acanthocephales, thorny-headed worms, or spiny-headed worms, characterized by the presence of an evertable proboscis, armed with spines, which it uses to pierce and hold the gut wall of its host. Acanthocephalans typically have complex life cycles, involving a number of hosts, including invertebrates, fishes, amphibians, birds, and mammals. About 1150 species have been described.[citation needed] [more]
Acoelomorpha
Annelida
The annelids (also called "ringed worms"), formally called Annelida (from French annel?s "ringed ones", ultimately from Latin anellus "little ring"), are a large phylum of segmented worms, with over 17,000 modern species including ragworms, earthworms and leeches. They are found in marine environments from tidal zones to hydrothermal vents, in freshwater, and in moist terrestrial environments. Although most textbooks still use the traditional division into polychaetes (almost all marine), oligochaetes (which include earthworms) and leech-like species, research since 1997 has radically changed this scheme, viewing leeches as a sub-group of oligochaetes and oligochaetes as a sub-group of polychaetes. In addition, the Pogonophora, Echiura and Sipuncula, previously regarded as separate phyla, are now regarded as sub-groups of polychaetes. Annelids are considered members of the Lophotrochozoa, a "super-phylum" of protostomes that also includes molluscs, brachiopods, flatworms and nemerteans. [more]
Arthropoda
An arthropod is an invertebrate animal having an exoskeleton (external skeleton), a segmented body, and jointed appendages. Arthropods are members of the phylum Arthropoda (from Greek ?rthron, "joint", and p?d?? pod?s "foot", which together mean "jointed feet"), and include the insects, arachnids, crustaceans, and others. Arthropods are characterized by their jointed limbs and cuticles, which are mainly made of a-chitin; the cuticles of crustaceans are also biomineralized with calcium carbonate. The rigid cuticle inhibits growth, so arthropods replace it periodically by molting. The arthropod body plan consists of repeated segments, each with a pair of appendages. It is so versatile that they have been compared to Swiss Army knives, and it has enabled them to become the most species-rich members of all ecological guilds in most environments. They have over a million described species, making up more than 80% of all described living animal species, and are one of only two animal groups that are very successful in dry environments ? the other being the amniotes. They range in size from microscopic plankton up to forms a few meters long. [more]
Brachiopoda
Brachiopods, phylum Brachiopoda, are marine animals that have hard "valves" (shells) on the upper and lower surfaces, unlike the left and right arrangement in bivalve molluscs. Brachiopod valves are hinged at the rear end, while the front can be opened for feeding or closed for protection. Two major groups are recognized, articulate and inarticulate. Articulate brachiopods have toothed hinges and simple opening and closing muscles, while inarticulate brachiopods have untoothed hinges and a more complex system of muscles used to keep the two halves aligned. In a typical brachiopod a stalk-like pedicle projects from an opening in one of the valves, known as the pedicle valve, attaching the animal to the seabed but clear of silt that would obstruct the opening. [more]
Bryozoa
The Bryozoa, also known as Ectoprocta or commonly as moss animals[], are a phylum of aquatic invertebrate animals. Typically about 0.5 millimetres (0.020 in) long, they are filter feeders that sieve food particles out of the water using a retractable lophophore, a "crown" of tentacles lined with cilia. Most marine species live in tropical waters, but a few occur in oceanic trenches, and others are found in polar waters. One class lives only in a variety of freshwater environments, and a few members of a mostly marine class prefer brackish water. Over 4,000 living species are known. One genus is solitary and the rest colonial. [more]
Chaetognatha
Chaetognatha, meaning hair-jaws, and commonly known as arrow worms, are a phylum of predatory marine worms that are a major component of plankton worldwide. About 20% of the known species are benthic, that is belonging to the lowest zone of the ocean, or benthic zone, and can attach to algae and rocks. They are found in all marine waters, from surface tropical waters and shallow tide pools to the deep sea and polar regions. Most chaetognaths are transparent and are torpedo shaped, but some deep-sea species are orange. They range in size from 2 to 120 millimetres (0.079 to 4.7 in). [more]
Cnidaria
Cnidaria ( with a silent c) is a phylum containing over 10,000 species of animals found exclusively in aquatic and mostly marine environments. Their distinguishing feature is cnidocytes, specialized cells that they use mainly for capturing prey. Their bodies consist of mesoglea, a non-living jelly-like substance, sandwiched between two layers of epithelium that are mostly one cell thick. They have two basic body forms: swimming medusae and sessile polyps, both of which are radially symmetrical with mouths surrounded by tentacles that bear cnidocytes. Both forms have a single orifice and body cavity that are used for digestion and respiration. Many cnidarian species produce colonies that are single organisms composed of medusa-like or polyp-like zooids, or both. Cnidarians' activities are coordinated by a decentralized nerve net and simple receptors. Several free-swimming Cubozoa and Scyphozoa possess balance-sensing statocysts, and some have simple eyes. Not all cnidarians reproduce sexually. Many have complex lifecycles with asexual polyp stages and sexual medusae, but some omit either the polyp or the medusa stage. [more]
Cycliophora
Symbion is the name of a genus of aquatic animals, less than ? mm wide, found living attached to the bodies of cold-water lobsters. They have sac-like bodies, and three distinctly different forms in different parts of their two-stage life-cycle. They appear so different from other animals that they were assigned their own, new phylum Cycliophora shortly after they were discovered in 1995. This was the first new phylum of multicelled organism to be discovered since the Loricifera in 1983. [more]
Echiura
The Echiura, or spoon worms, are a small group of marine animals. They are often considered to be a group of annelids, although they lack the segmented structure found in other members of that group, and so may also be treated as a separate phylum. However, phylogenetic analyses of DNA sequences place echiurans and pogonophorans within the Annelida. The Echiura fossilise poorly and the earliest known specimen is from the Upper Carboniferous (called the Pennsylvanian in North America). However, U-shaped fossil burrows that could be Echiuran have been found dating back to the Cambrian. [more]
Entoprocta
Entoprocta, whose name means "anus inside", is a phylum of mostly sessile aquatic animals, ranging from 0.1 to 7 millimetres (0.0039 to 0.28 in) long. Mature individuals are goblet-shaped, on relatively long stalks. They have a "crown" of solid tentacles whose cilia generate water currents that draw food particles towards the mouth, and both the mouth and anus lie inside the "crown". The superficially similar Bryozoa (Ectoprocta) have the anus outside a "crown" of hollow tentacles. Most families of entoprocts are colonial, and all but 2 of the 150 species are marine. A few solitary species can move slowly. [more]
Euglenozoa
The Euglenozoa are a large group of flagellate protozoa. They include a variety of common free-living species, as well as a few important parasites, some of which infect humans. There are two main subgroups, the euglenids and kinetoplastids. Euglenozoa are unicellular, mostly around 15-40 ?m in size, although some euglenids get up to 500 ?m long. [more]
Gastrotricha
The gastrotrichs (from Greek ?ast??, gaster ["stomach"], and ????, thrix ["hair"]), often called hairy backs, are a phylum of microscopic (0.06-3.0 mm) animals abundant in fresh water and marine environments. Most fresh water species are part of the periphyton and benthos. Marine species are found mostly interstitially in between sediment particles, while terrestrial species live in the water films around grains of soil. [more]
Gnathostomulida
Gnathostomulids, or jaw worms, are a small phylum of nearly microscopic marine animals. They inhabit sand and mud beneath shallow coastal waters and can survive in relatively anoxic environments. They were first recognised and described in 1956. [more]
Haptophyta
The haptophytes, classified either as the Prymnesiophyta or Haptophyta, are a division of algae. [more]
Kinorhyncha
Kinorhyncha (Gr. ?????, kineo 'move' + ??????, rhynchos 'snout') is a phylum of small (1 mm or less) marine pseudocoelomate invertebrates that are widespread in mud or sand at all depths as part of the meiobenthos. They are also called mud dragons. [more]
Loricifera
Mollusca
The Mollusca (pronounced ), common name molluscs or mollusks (pronounced /'m?l?sks/), is a large phylum of invertebrate animals. There are around 85,000 recognized extant species of molluscs. Mollusca is the largest marine phylum, comprising about 23% of all the named marine organisms. Numerous molluscs also live in freshwater and terrestrial habitats. Molluscs are highly diverse, not only in size and in anatomical structure, but also in behaviour and in habitat. The phylum is typically divided into nine or ten taxonomic classes, of which two are entirely extinct. Cephalopod molluscs such as squid, cuttlefish and octopus are among the most neurologically advanced of all invertebrates ? and either the giant squid or the colossal squid is the largest known invertebrate species. The gastropods (snails and slugs) are by far the most numerous molluscs in terms of classified species, and account for 80% of the total. [more]
Nematoda
The nematodes () or roundworms (phylum Nematoda) are the most diverse phylum of pseudocoelomates, and one of the most diverse of all animals. Nematode species are very difficult to distinguish; over 28,000 have been described, of which over 16,000 are parasitic. The total number of nematode species has been estimated to be about 1,000,000. Unlike cnidarians or flatworms, roundworms have tubular digestive systems with openings at both ends. [more]
Nematomorpha
Nematomorpha (sometimes called Gordiacea, and commonly known as horsehair worms or Gordian worms) is a phylum of parasitic animals that are superficially morphologically similar to nematode worms, hence the name. They range in size in most species from 50 to 100 centimetres (20 to 39 in) long and can reach in extreme cases up to 2 metres, and 1 to 3 millimetres (0.039 to 0.12 in) in diameter. Horsehair worms can be discovered in damp areas such as watering troughs, streams, puddles, and cisterns. The adult worms are free living, but the larvae are parasitic on beetles, cockroaches, orthopterans, and crustaceans. About 351 species are known and a conservative estimate suggests that there may be about 2000 species worldwide. The name "Gordian" stems from the legendary Gordian knot. This relates to the fact that nematomorpha often tie themselves in knots. [more]
Nemertea
Nemertea is a phylum of invertebrate animals also known as "ribbon worms" or "proboscis worms". Alternative names for the phylum have included Nemertini, Nemertinea and Rhynchocoela. Although most are less than 20 centimetres (7.9 in) long, one specimen has been estimated at 54 metres (177 ft), which would make it the longest animal ever found. Most are very slim, usually only a few millimeters wide, although a few have relatively short but wide bodies. Many have patterns of yellow, orange, red and green coloration. [more]
Onychophora
The velvet worms (Onychophora ? literally "claw bearers", also known as Protracheata) are a minor ecdysozoan phylum. These obscurely segmented organisms have tiny eyes, antennae, multiple pairs of legs and slime glands. They have variously been compared to worms with legs, caterpillars and slugs. Most common in tropical regions of the Southern Hemisphere, they prey on smaller animals such as insects, which they catch by squirting an adhesive slime. In modern zoology, they are particularly renowned for their curious mating behaviour and for bearing live young. They are becoming increasingly popular as pets due to their bizarre appearance and eating habits.[] [more]
Oomycota
O?mycota or o?mycetes form a distinct phylogenetic lineage of fungus-like eukaryotic microorganisms. They are filamentous, microscopic, absorptive organisms that reproduce both sexually and asexually. Oomycetes occupy both saprophytic and pathogenic lifestyles ? and include some of the most notorious pathogens of plants, causing devastating diseases such as late blight of potato and sudden oak death. They are also often referred to as water molds (or water moulds), although the water-preferring nature which led to that name is not true of most species, which are terrestrial pathogens. [more]
Platyhelminthes
The flatworms, known in scientific literature as Platyhelminthes or Plathelminthes (from the Greek p?at?, platy, meaning "flat" and ?????? (root: ??????-), helminth-, meaning worm) are a phylum of relatively simple bilaterian, unsegmented, soft-bodied invertebrate animals. Unlike other bilaterians, they have no body cavity, and no specialized circulatory and respiratory organs, which restricts them to flattened shapes that allow oxygen and nutrients to pass through their bodies by diffusion. [more]
Porifera
Sponges are animals of the phylum Porifera (; meaning "pore bearer").They are multicellular organisms which have bodies full of pores and channels allowing water to circulate through them, consist of jelly-like mesohyl sandwiched between two thin layers of cells. While all animals have unspecialized cells that can transform into specialized cells, sponges are unique in having some specialized cells, but can also have specialized cells that can transform into other types, often migrating between the main cell layers and the mesohyl in the process. Sponges do not have nervous, digestive or circulatory systems. Instead, most rely on maintaining a constant water flow through their bodies to obtain food, oxygen and remove wastes. The shapes of their bodies are adapted for maximal efficiency of water flow. Water enters through the central cavity, deposits nutrients, and leaves through a hole called the osculum. All sponges are sessile aquatic animals. Although there are freshwater species, the great majority are marine (salt water) species, ranging from tidal zones to depths exceeding 8,800 metres (5.5 mi). [more]
Priapulida
Priapulida (priapulid worms or penis worms, from Gr. p???p??, priapos 'Priapus' + Lat. -ul-, diminutive) is a phylum of marine worms. They are named for their extensible spiny proboscis, which, in some species, may have a shape like that of a human penis. They live in the mud, which they eat, in comparatively shallow waters up to 90 metres (300 ft). [more]
Radiozoa
Radiolarians (also radiolaria) are amoeboid protozoa (diameter 0.1-0.2 mm) that produce intricate mineral skeletons, typically with a central capsule dividing the cell into inner and outer portions, called endoplasm and ectoplasm. They are found as zooplankton throughout the ocean, and their skeletal remains cover large portions of the ocean bottom as radiolarian ooze. Due to their rapid turn-over of species, they represent an important diagnostic fossil found from the Cambrian onwards. Some common radiolarian fossils include , Heliosphaera and Hexadoridium. [more]
Rotifera
The rotifers (Rotifera, commonly called wheel animals) make up a phylum of microscopic and near-microscopic pseudocoelomate animals. They were first described by Rev. John Harris in 1696, and other forms were described by Anton van Leeuwenhoek in 1703. Most rotifers are around 0.1?0.5 mm long (although their size can range from 50 ?m to over 2 millimeters), and are common in freshwater environments throughout the world with a few saltwater species; for example, those of genus . Some rotifers are free swimming and truly planktonic, others move by inchworming along a substrate, and some are sessile, living inside tubes or gelatinous holdfasts that are attached to a substrate. About 25 species are colonial (e.g., Sinantherina semibullata), either sessile or planktonic. Rotifers are an important part of the freshwater zooplankton, being a major foodsource and with many species also contributing to the decomposition of soil organic matter. Most species of the rotifers are cosmopolitan, but there are also some endemic species, like Cephalodella vittata to Lake Baikal. Recent barcoding evidence, however, suggests that some 'cosmopolitan' species, such as Brachionus plicatilis, B. calyciflorus, Lecane bulla, among others, are actually species complexes. [more]
Sipuncula
The Sipuncula or Sipunculida (common names sipunculid worms or peanut worms) is a group containing 144-320 species (estimates vary) of bilaterally symmetrical, unsegmented marine worms. Traditionally considered a phylum, molecular work suggests that they might be a subgroup of phylum Annelida. [more]
Tardigrada
Tardigrades (commonly known as waterbears or moss piglets) form the phylum Tardigrada, part of the superphylum Ecdysozoa. They are small, water-dwelling, segmented animals with eight legs. Tardigrades were first described by Johann August Ephraim Goeze in 1773 (kleiner Wasserb?r = little water bear). The name Tardigrada means "slow walker" and was given by Lazzaro Spallanzani in 1777. The name water bear comes from the way they walk, reminiscent of a bear's gait. The biggest adults may reach a body length of 1.5 millimetres (0.059 in), the smallest below 0.1 mm. Freshly hatched larvae may be smaller than 0.05 mm. [more]
At least 1,107 species and subspecies belong to the Phylum Tardigrada.
More info about the Phylum Tardigrada may be found here.
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