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Hipposideridae

(Family)

Overview

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Hipposideridae is a family of bats commonly known as the "Old World Leaf-nosed Bats". While it has often been seen as a subfamily, Hipposiderinae, of the family Rhinolophidae, it is now more generally classified as its own family.1] Nevertheless, it is most closely related to Rhinolophidae within the suborder Pteropodiformes (or Yinpterochiroptera).[2]

Hipposideridae contains ten living genera and more than 70 species, mostly in the widespread genus Hipposideros.[3] In addition, several fossil genera are known; the oldest fossils attributed to the family are from the middle Eocene of Europe.[4] In their 1997 , Malcolm C. McKenna and Susan K. Bell proposed a division of Hipposideridae (called Rhinonycterinae in their work) into three tribes, one with two subtribes,[5] but these tribes turned out to be non-monophyletic and have been abandoned.[1] A different classification was proposed by Hand and Kirsch in 2003.[6] More recently, Petr Benda and Peter Vallo (2009) proposed a separate tribe, Triaenopini, for the genera Triaenops, Paratriaenops, and possibly Cloeotis.[7]

Genera

The genera included in Hipposideridae are the following (species counts only include living species):[8]

List of species

Hipposideros lankadiva in Sri Lanka
Pseudorhinolophus antiquus sk ull and lower jaw at the Museum f?r Naturkunde, Berlin
Colony of Hipposideros lankadiva (or perhaps Hipposideros speori) in a cave in Sri Lanka
  • Family Hipposideridae ? Old World leaf-nosed bats
    • Genus Anthops
      • Flower-faced Bat, Anthops ornatus
    • Genus Asellia ? trident leaf-nosed bats
      • Patrizi's Trident Leaf-nosed Bat, Asellia patrizii
      • Trident Bat, Asellia tridens
    • Genus Aselliscus ? Tate's trident-nosed bats
      • Stoliczka's Trident Bat, Aselliscus stoliczkanus
      • Temminck's Trident Bat, Aselliscus tricuspidatus
    • Genus Cloeotis
      • Percival's Trident Bat, Cloeotis percivali
    • Genus Coelops ? tailless leaf-nosed bats
      • East Asian Talless Leaf-nosed Bat, Coelops frithii
      • Philippine Tailless Leaf-nosed Bat, Coelops hirsutus
      • Malayan Tailless Leaf-nosed Bat, Coelops robinsoni
    • Genus Hipposideros ? round leaf bats
      • Aba Roundleaf Bat, Hipposideros abae
      • Great Roundleaf Bat, Hipposideros armiger
      • Dusky Roundleaf Bat, Hipposideros ater
      • Benito Roundleaf Bat, Hipposideros beatus
      • Boeadi?s Roundleaf Bat, Hipposideros boeadii
      • Bicolored Roundleaf Bat, Hipposideros bicolor
      • Short-headed Roundleaf Bat, Hipposideros breviceps
      • Sundevall's Roundleaf Bat, Hipposideros caffer
      • Spurred Roundleaf Bat, Hipposideros calcaratus
      • Greater Roundleaf Bat, Hipposideros camerunensis
      • Fawn Roundleaf Bat, Hipposideros cervinus
      • Ashy Roundleaf Bat, Hipposideros cineraceus
      • Commerson's Roundleaf Bat, Hipposideros commersoni
      • Large Mindanao Roundleaf Bat, Hipposideros coronatus
      • Telefomin Roundleaf Bat, Hipposideros corynophyllus
      • Cox's Roundleaf Bat, Hipposideros coxi
      • Timor Roundleaf Bat, Hipposideros crumeniferus
      • Short-tailed Roundleaf Bat, Hipposideros curtus
      • Cyclops Roundleaf Bat, Hipposideros cyclops
      • Makira Roundleaf Bat, Hipposideros demissus
      • Diadem Roundleaf Bat, Hipposideros diadema
      • Fierce Roundleaf Bat, Hipposideros dinops
      • Borneo Roundleaf Bat, Hipposideros doriae
      • Khajuria's Leaf-nosed Bat, Hipposideros durgadasi
      • Dayak Roundleaf Bat, Hipposideros dyacorum
      • Hill's Roundleaf Bat, Hipposideros edwardshilli
      • Hipposideros fasensis
      • Sooty Roundleaf Bat, Hipposideros fuliginosus
      • Fulvus Roundleaf Bat, Hipposideros fulvus
      • Cantor's Roundleaf Bat, Hipposideros galeritus
      • Giant Roundleaf Bat, Hipposideros gigas
      • Grand Roundleaf Bat, Hipposideros grandis
      • Thailand Roundleaf Bat, Hipposideros halophyllus
      • Kolar Leaf-nosed Bat, Hipposideros hypophyllus
      • Crested Roundleaf Bat, Hipposideros inexpectatus
      • Arnhem Leaf-nosed Bat, Hipposideros inornatus
      • Jones's Roundleaf Bat, Hipposideros jonesi
      • Phou Khao Khouay Leaf-nosed Bat, Hipposideros khaokhouayensis
      • Khasian Leaf-nosed Bat, Hipposideros khasiana
      • Lamotte's Roundleaf Bat, Hipposideros lamottei
      • Indian Roundleaf Bat, Hipposideros lankadiva
      • Intermediate Roundleaf Bat, Hipposideros larvatus
      • Large Asian Roundleaf Bat, Hipposideros lekaguli
      • Shield-faced Roundleaf Bat, Hipposideros lylei
      • Big-eared Roundleaf Bat, Hipposideros macrobullatus
      • Maduran Leaf-nosed Bat, Hipposideros madurae
      • Maggie Taylor's Roundleaf Bat, Hipposideros maggietaylorae
      • Aellen's Roundleaf Bat, Hipposideros marisae
      • Ethiopian Large-eared Roundleaf Bat, Hipposideros megalotis
      • Fly River Roundleaf Bat, Hipposideros muscinus
      • Malayan Roundleaf Bat, Hipposideros nequam
      • Philippine Forest Roundleaf Bat, Hipposideros obscurus
      • Orbiculus Leaf-nosed Bat, Hipposideros orbiculus
      • Biak Roundleaf Bat, Hipposideros papua
      • Hipposideros parnabyi
      • Peleng Leaf-nosed Bat, Hipposideros pelingensis
      • Pomona Roundleaf Bat, Hipposideros pomona
      • Pratt's Roundleaf Bat, Hipposideros pratti
      • Philippine Pygmy Roundleaf Bat, Hipposideros pygmaeus
      • Ridley's Leaf-nosed Bat, Hipposideros ridleyi
      • Laotian Leaf-nosed Bat, Hipposideros rotalis
      • Noack's Roundleaf Bat, Hipposideros ruber
      • Hipposideros scutinares
      • Semon's Roundleaf Bat, Hipposideros semoni
      • Sorensen's Leaf-nosed Ba, Hipposideros sorenseni
      • Schneider's Leaf-nosed Bat, Hipposideros speoris
      • Narrow-eared Roundleaf Bat, Hipposideros stenotis
      • Sumban Leaf-nosed Bat, Hipposideros sumbae
      • Hipposideros tephrus
      • Sa? Tom? Leaf-nosed Bat, Hipposideros thomensis
      • Lesser Great Leaf-nosed Bat, Hipposideros turpis
      • Striped Leaf-nosed Bat, Hipposideros vittatus
      • Wollaston's Roundleaf Bat, Hipposideros wollastoni
    • Genus Paracoelops
      • Vietnam Leaf-nosed Bat, Paracoelops megalotis
    • Genus Rhinonicteris
      • Orange Leaf-nosed Bat, Rhinonicteris aurantia
    • Genus Paratriaenops
      • Grandidier's Trident Bat, Paratriaenops auritus
      • Trouessart's Trident Bat, Paratriaenops furculus
      • Paulian's Triple Leaf-nosed Bat, Paratriaenops pauliani
    • Genus Triaenops ? triple leaf-nosed bats
      • Triaenops afer
      • Triaenops menamena
      • Triaenops parvus
      • Persian Trident Bat (Rufous Trident Bat), Triaenops persicus (also known as Triaenops rufus)
ra and more than 70 species, mostly in the widespread genus Hipposideros.[3] In addition, several fossil genera are known; the oldest fossils attributed to the family are from the middle Eocene of Europe.[4] In their 1997 , Malcolm C. McKenna and Susan K. Bell proposed a division of Hipposideridae (called Rhinonycterinae in their work) into three tribes, one with two subtribes,[5] but these tribes turned out to be non-monophyletic and have been abandoned.[1] A different classification was proposed by Hand and Kirsch in 2003.[6] More recently, Petr Benda and Peter Vallo (2009) proposed a separate tribe, Triaenopini, for the genera Triaenops, Paratriaenops, and possibly Cloeotis.[7]

Genera

The genera included in Hipposideridae are the following (species counts only include living species):[8]

  • Archerops (Miocene of Australia)[9]
  • Anth ops (one species; Solomon Islands and Bougainville Island)
  • Asellia (two species; Africa and southwestern Asia; Miocene fossils from Europe)
  • Aselliscus (two species; southeastern Asia and Melanesia)
  • Brachipposideros (Oligocene to Miocene of Europe and Miocene of Australia; sometimes included in Hipposideros)
  • Brevipalatus (Miocene of Australia)[10]
  • Cloeotis (one species; Africa)
  • Coelops (at least two species; southeastern Asia; Miocene fossils from Africa)
  • Hipposideros (more than sixty species; Africa, southern Eurasia, and Australasia; oldest fossils from the Eocene of Europe; includes Pseudorhinolophus, sometimes considered a separate genus)
  • Miophyllorhina (Miocene of Australia)[11]
  • Palaeophyllophora (Eocene to Miocene of Europe)
  • Paracoelops (one species; Vietnam)
  • Paraphyllophora (Eocene or Oligocene to Miocene of Europe)
  • Paratriaenops (previously included in Triaenops; three species; Madagascar and Seychelles)[12]
  • Rhinonicteris (one species; Australia; earliest fossils from the Miocene)[11]
  • Riversleighia (Miocene of Australia)[11]
  • Triaenops (four species; Africa and southwestern Asia)[12]
  • Vaylatsia (Oligocene of Europe)[13]
  • Xenorhinos (Miocene of Australia)[11]

List of species

Hipposideros lankadiva in Sri Lanka
Pseudorhinolophus antiquus skull and lower jaw at the Museum f?r Naturkunde, Berlin
Colony of Hipposideros lankadiva (or perhaps Hipposideros speori) in a cave in Sri Lanka
  • Family Hipposideridae ? Old World leaf-nosed bats
    • Genus Anthops
      • Flower-faced Bat, Anthops ornatus
    • Genus Asellia ? trident leaf-nosed bats
      • Patrizi's Trident Leaf-nosed Bat, Asellia patrizii
      • Trident Bat, Asellia tridens
    • Genus Aselliscus ? Tate's trident-nosed bats
      • Stoliczka's Trident Bat, Aselliscus stoliczkanus
      • Temminck's Trident Bat, Aselliscus tricuspidatus
    • Genus Cloeotis
      • Percival's Trident Bat, Cloeotis percivali
    • Genus Coelops ? tailless leaf-nosed bats
      • East Asian Talless Leaf-nosed Bat, Coelops frithii
      • Philippine Tailless Leaf-nosed Bat, Coelops hirsutus
      • Malayan Tailless Leaf-nosed Bat, Coelops robinsoni
    • Genus Hipposideros ? round leaf bats
      • Aba Roundleaf Bat, Hipposideros abae
      • Great Roundleaf Bat, Hipposideros armiger
      • Dusky Roundleaf Bat, Hipposideros ater
      • Benito Roundleaf Bat, Hipposideros beatus
      • Boeadi?s Roundleaf Bat, Hipposideros boeadii
      • Bicolored Roundleaf Bat, Hipposideros bicolor
      • Short-headed Roundleaf Bat, Hipposideros breviceps
      • Sundevall's Roundleaf Bat, Hipposideros caffer
      • Spurred Roundleaf Bat, Hipposideros calcaratus
      • Greater Roundleaf Bat, Hipposideros camerunensis
      • Fawn Roundleaf Bat, Hipposideros cervinus
      • Ashy Roundleaf Bat, Hipposideros cineraceus
      • Commerson's Roundleaf Bat, Hipposideros commersoni
      • Large Mindanao Roundleaf Bat, Hipposideros coronatus
      • Telefomin Roundleaf Bat, Hipposideros corynophyllus
      • Cox's Roundleaf Bat, Hipposideros coxi
      • Timor Roundleaf Bat, Hipposideros crumeniferus
      • Short-tailed Roundleaf Bat, Hipposideros curtus
      • Cyclops Roundleaf Bat, Hipposideros cyclops
      • Makira Roundleaf Bat, Hipposideros demissus
      • Diadem Roundleaf Bat, Hipposideros diadema
      • Fierce Roundleaf Bat, Hipposideros dinops
      • Borneo Roundleaf Bat, Hipposideros doriae
      • Khajuria's Leaf-nosed Bat, Hipposideros durgadasi
      • Dayak Roundleaf Bat, Hipposideros dyacorum
      • Hill's Roundleaf Bat, Hipposideros edwardshilli
      • Hipposideros fasensis
      • Sooty Roundleaf Bat, Hipposideros fuliginosus
      • Fulvus Roundleaf Bat, Hipposideros fulvus
      • Cantor's Roundleaf Bat, Hipposideros galeritus
      • Giant Roundleaf Bat, Hipposideros gigas
      • Grand Roundleaf Bat, Hipposideros grandis
      • Thailand Roundleaf Bat, Hipposideros halophyllus
      • Kolar Leaf-nosed Bat, Hipposideros hypophyllus
      • Crested Roundleaf Bat, Hipposideros inexpectatus
      • Arnhem Leaf-nosed Bat, Hipposideros inornatus
      • Jones's Roundleaf Bat, Hipposideros jonesi
      • Phou Khao Khouay Leaf-nosed Bat, Hipposideros khaokhouayensis
      • Khasian Leaf-nosed Bat, Hipposideros khasiana
      • Lamotte's Roundleaf Bat, Hipposideros lamottei
      • Indian Roundleaf Bat, Hipposideros lankadiva
      • Intermediate Roundleaf Bat, Hipposideros larvatus
      • Large Asian Roundleaf Bat, Hipposideros lekaguli
      • Shield-faced Roundleaf Bat, Hipposideros lylei
      • Big-eared Roundleaf Bat, Hipposideros macrobullatus
      • Maduran Leaf-nosed Bat, Hipposideros madurae
      • Maggie Taylor's Roundleaf Bat, Hipposideros maggietaylorae
      • Aellen's Roundleaf Bat, Hipposideros marisae
      • Ethiopian Large-eared Roundleaf Bat, Hipposideros megalotis
      • Fly River Roundleaf Bat, Hipposideros muscinus
      • Malayan Roundleaf Bat, Hipposideros nequam
      • Philippine Forest Roundleaf Bat, Hipposideros obscurus
      • Orbiculus Leaf-nosed Bat, Hipposideros orbiculus
      • Biak Roundleaf Bat, Hipposideros papua
      • Hipposideros parnabyi
      • Peleng Leaf-nosed Bat, Hipposideros pelingensis
      • Pomona Roundleaf Bat, Hipposideros pomona
      • Pratt's Roundleaf Bat, Hipposideros pratti
      • Philippine Pygmy Roundleaf Bat, Hipposideros pygmaeus
      • Ridley's Leaf-nosed Bat, Hipposideros ridleyi
      • Laotian Leaf-nosed Bat, Hipposideros rotalis
      • Noack's Roundleaf Bat, Hipposideros ruber
      • Hipposideros scutinares
      • Semon's Roundleaf Bat, Hipposideros semoni
      • Sorensen's Leaf-nosed Ba, Hipposideros sorenseni
      • Schneider's Leaf-nosed Bat, Hipposideros speoris
      • Narrow-eared Roundleaf Bat, Hipposideros stenotis
      • Sumban Leaf-nosed Bat, Hipposideros sumbae
      • Hipposideros tephrus
      • Sa? Tom? Leaf-nosed Bat, Hipposideros thomensis
      • Lesser Great Leaf-nosed Bat, Hipposideros turpis
      • Striped Leaf-nosed Bat, Hipposideros vittatus
      • Wollaston's Roundleaf Bat, Hipposideros wollastoni
    • Genus Paracoelops
      • Vietnam Leaf-nosed Bat, Paracoelops megalotis
    • Genus Rhinonicteris
      • Orange Leaf-nosed Bat, Rhinonicteris aurantia
    • Genus Paratriaenops
      • Grandidier's Trident Bat, Paratriaenops auritus
      • Trouessart's Trident Bat, Paratriaenops furculus
      • Paulian's Triple Leaf-nosed Bat, Paratriaenops pauliani
    • Genus Triaenops ? triple leaf-nosed bats
      • Triaenops afer
      • Triaenops menamena
      • Triaenops parvus
      • Persian Trident Bat (Rufous Trident Bat), Triaenops persicus (also known as Triaenops rufus)

Notes

  1. ^ This name technically has priority over Hipposiderinae Lydekker, 1891, and some have consequently used "Rhinonycteridae" or "Rhinonycterinae" for this (sub)family; however, Hipposideridae/inae has been in common use since 1907 and is currently retained pending action by the International Commission on Zoological Nomenclature.[1]
ass="reference">[8]

    < li>Archerops (Miocene of Australia)[9]
  • Anthops (one species; Solomon Islands and Bougainville Island)
  • Asellia (two species; Africa and southwestern Asia; Miocene fossils from Europe)
  • Aselliscus (two species; southeastern Asia and Melanesia)
  • Brachipposideros (Oligocene to Miocene of Europe and Miocene of Australia; sometimes included in Hipposideros)
  • Brevipalatus (Miocene of Australia)[10]
  • Cloeotis (one species; Africa)
  • Coelops (at least two species; southeastern Asia; Miocene fossils from Africa)
  • Hipposideros (more than sixty species; Africa, southern Eurasia, and Australasia; oldest fossils from the Eocene of Europe; includes Pseudorhinolophus, sometimes considered a separate genus)
  • Miophyllorhina (Miocene of Australia)[11]
  • Palaeophyllophora (Eocene to Miocene of Europe)
  • Paracoelops (one species; Vietnam)
  • Paraphyllophora (Eocene or Oligocene to Miocene of Europe)
  • Paratriaenops (previously included in Triaenops; three species; Madagascar and Seychelles)[12]
  • Rhinonicteris (one species; Australia; earliest fossils from the Miocene)[11]
  • Riversleighia (Miocene of Australia)[11]
  • Triaenops (four species; Africa and southwestern Asia)[12]
  • Vaylatsia (Oligocene of Europe)[13]
  • Xenorhinos (Miocene of Australia)[11]

List of species

Hipposideros lankadiva in Sri Lanka
Pseudorhinolophus a ntiquus skull and lower jaw at the Museum f?r Naturkunde, Berlin
Colony of Hipposideros lankadiva (or perhaps Hipposideros speori) in a cave in Sri Lanka
  • Family Hipposideridae ? Old World leaf-nosed bats
    • Genus Anthops
      • Flower-faced Bat, Anthops ornatus
    • Genus Asellia ? trident leaf-nosed bats
      • Patrizi's Trident Leaf-nosed Bat, Asellia patrizii
      • Trident Bat, Asellia tridens
    • Genus Aselliscus ? Tate's trident-nosed bats
      • Stoliczka's Trident Bat, Aselliscus stoliczkanus
      • Temminck's Trident Bat, Aselliscus tricuspidatus
    • Genus Cloeotis
      • Percival's Trident Bat, Cloeotis percivali
    • Genus Coelops ? tailless leaf-nosed bats
      • East Asian Talless Leaf-nosed Bat, Coelops frithii
      • Philippine Tailless Leaf-nosed Bat, Coelops hirsutus
      • Malayan Tailless Leaf-nosed Bat, Coelops robinsoni
    • Genus Hipposideros ? round leaf bats
      • Aba Roundleaf Bat, Hipposideros abae
      • Great Roundleaf Bat, Hipposideros armiger
      • Dusky Roundleaf Bat, Hipposideros ater
      • Benito Roundleaf Bat, Hipposideros beatus
      • Boeadi?s Roundleaf Bat, Hipposideros boeadii
      • Bicolored Roundleaf Bat, Hipposideros bicolor
      • Short-headed Roundleaf Bat, Hipposideros breviceps
      • Sundevall's Roundleaf Bat, Hipposideros caffer
      • Spurred Roundleaf Bat, Hipposideros calcaratus
      • Greater Roundleaf Bat, Hipposideros camerunensis
      • Fawn Roundleaf Bat, Hipposideros cervinus
      • Ashy Roundleaf Bat, Hipposideros cineraceus
      • Commerson's Roundleaf Bat, Hipposideros commersoni
      • Large Mindanao Roundleaf Bat, Hipposideros coronatus
      • Telefomin Roundleaf Bat, Hipposideros corynophyllus
      • Cox's Roundleaf Bat, Hipposideros coxi
      • Timor Roundleaf Bat, Hipposideros crumeniferus
      • Short-tailed Roundleaf Bat, Hipposideros curtus
      • Cyclops Roundleaf Bat, Hipposideros cyclops
      • Makira Roundleaf Bat, Hipposideros demissus
      • Diadem Roundleaf Bat, Hipposideros diadema
      • Fierce Roundleaf Bat, Hipposideros dinops
      • Borneo Roundleaf Bat, Hipposideros doriae
      • Khajuria's Leaf-nosed Bat, Hipposideros durgadasi
      • Dayak Roundleaf Bat, Hipposideros dyacorum
      • Hill's Roundleaf Bat, Hipposideros edwardshilli
      • Hipposideros fasensis
      • Sooty Roundleaf Bat, Hipposideros fuliginosus
      • Fulvus Roundleaf Bat, Hipposideros fulvus
      • Cantor's Roundleaf Bat, Hipposideros galeritus
      • Giant Roundleaf Bat, Hipposideros gigas
      • Grand Roundleaf Bat, Hipposideros grandis
      • Thailand Roundleaf Bat, Hipposideros halophyllus
      • Kolar Leaf-nosed Bat, Hipposideros hypophyllus
      • Crested Roundleaf Bat, Hipposideros inexpectatus
      • Arnhem Leaf-nosed Bat, Hipposideros inornatus
      • Jones's Roundleaf Bat, Hipposideros jonesi
      • Phou Khao Khouay Leaf-nosed Bat, Hipposideros khaokhouayensis
      • Khasian Leaf-nosed Bat, Hipposideros khasiana
      • Lamotte's Roundleaf Bat, Hipposideros lamottei
      • Indian Roundleaf Bat, Hipposideros lankadiva
      • Intermediate Roundleaf Bat, Hipposideros larvatus
      • Large Asian Roundleaf Bat, Hipposideros lekaguli
      • Shield-faced Roundleaf Bat, Hipposideros lylei
      • Big-eared Roundleaf Bat, Hipposideros macrobullatus
      • Maduran Leaf-nosed Bat, Hipposideros madurae
      • Maggie Taylor's Roundleaf Bat, Hipposideros maggietaylorae
      • Aellen's Roundleaf Bat, Hipposideros marisae
      • Ethiopian Large-eared Roundleaf Bat, Hipposideros megalotis
      • Fly River Roundleaf Bat, Hipposideros muscinus
      • Malayan Roundleaf Bat, Hipposideros nequam
      • Philippine Forest Roundleaf Bat, Hipposideros obscurus
      • Orbiculus Leaf-nosed Bat, Hipposideros orbiculus
      • Biak Roundleaf Bat, Hipposideros papua
      • Hipposideros parnabyi
      • Peleng Leaf-nosed Bat, Hipposideros pelingensis
      • Pomona Roundleaf Bat, Hipposideros pomona
      • Pratt's Roundleaf Bat, Hipposideros pratti
      • Philippine Pygmy Roundleaf Bat, Hipposideros pygmaeus
      • Ridley's Leaf-nosed Bat, Hipposideros ridleyi
      • Laotian Leaf-nosed Bat, Hipposideros rotalis
      • Noack's Roundleaf Bat, Hipposideros ruber
      • Hipposideros scutinares
      • Semon's Roundleaf Bat, Hipposideros semoni
      • Sorensen's Leaf-nosed Ba, Hipposideros sorenseni
      • Schneider's Leaf-nosed Bat, Hipposideros speoris
      • Narrow-eared Roundleaf Bat, Hipposideros stenotis
      • Sumban Leaf-nosed Bat, Hipposideros sumbae
      • Hipposideros tephrus
      • Sa? Tom? Leaf-nosed Bat, Hipposideros thomensis
      • Lesser Great Leaf-nosed Bat, Hipposideros turpis
      • Striped Leaf-nosed Bat, Hipposideros vittatus
      • Wollaston's Roundleaf Bat, Hipposideros wollastoni
    • Genus Paracoelops
      • Vietnam Leaf-nosed Bat, Paracoelops megalotis
    • Genus Rhinonicteris
      • Orange Leaf-nosed Bat, Rhinonicteris aurantia
    • Genus Paratriaenops
      • Grandidier's Trident Bat, Paratriaenops auritus
      • Trouessart's Trident Bat, Paratriaenops furculus
      • Paulian's Triple Leaf-nosed Bat, Paratriaenops pauliani
    • Genus Triaenops ? triple leaf-nosed bats
      • Triaenops afer
      • Triaenops menamena
      • Triaenops parvus
      • Persian Trident Bat (Rufous Trident Bat), Triaenops persicus (also known as Triaenops rufus)
ra and more than 70 species, mostly in the widespread genus Hipposideros.[3] In addition, several fossil genera are known; the oldest fossils attributed to the family are from the middle Eocene of Europe.[4] In their 1997 , Malcolm C. McKenna and Susan K. Bell proposed a division of Hipposideridae (called Rhinonycterinae in their work) into three tribes, one with two subtribes,[5] but these tribes turned out to be non-monophyletic and have been abandoned.[1] A different classification was proposed by Hand and Kirsch in 2003.[6] More recently, Petr Benda and Peter Vallo (2009) proposed a separate tribe, Triaenopini, for the genera Triaenops, Paratriaenops, and possibly Cloeotis.[7]

Genera

The genera included in Hipposideridae are the following (species counts only include living species):[8]

  • Archerops (Miocene of Australia)[9]
  • Anth ops (one species; Solomon Islands and Bougainville Island)
  • Asellia (two species; Africa and southwestern Asia; Miocene fossils from Europe)
  • Aselliscus (two species; southeastern Asia and Melanesia)
  • Brachipposideros (Oligocene to Miocene of Europe and Miocene of Australia; sometimes included in Hipposideros)
  • Brevipalatus (Miocene of Australia)[10]
  • Cloeotis (one species; Africa)
  • Coelops (at least two species; southeastern Asia; Miocene fossils from Africa)
  • Hipposideros (more than sixty species; Africa, southern Eurasia, and Australasia; oldest fossils from the Eocene of Europe; includes Pseudorhinolophus, sometimes considered a separate genus)
  • Miophyllorhina (Miocene of Australia)[11]
  • Palaeophyllophora (Eocene to Miocene of Europe)
  • Paracoelops (one species; Vietnam)
  • Paraphyllophora (Eocene or Oligocene to Miocene of Europe)
  • Paratriaenops (previously included in Triaenops; three species; Madagascar and Seychelles)[12]
  • Rhinonicteris (one species; Australia; earliest fossils from the Miocene)[11]
  • Riversleighia (Miocene of Australia)[11]
  • Triaenops (four species; Africa and southwestern Asia)[12]
  • Vaylatsia (Oligocene of Europe)[13]
  • Xenorhinos (Miocene of Australia)[11]

List of species

Hipposideros lankadiva in Sri Lanka
Pseudorhinolophus antiquus skull and lower jaw at the Museum f?r Naturkunde, Berlin
Colony of Hipposideros lankadiva (or perhaps Hipposideros speori) in a cave in Sri Lanka
  • Family Hipposideridae ? Old World leaf-nosed bats
    • Genus Anthops
      • Flower-faced Bat, Anthops ornatus
    • Genus Asellia ? trident leaf-nosed bats
      • Patrizi's Trident Leaf-nosed Bat, Asellia patrizii
      • Trident Bat, Asellia tridens
    • Genus Aselliscus ? Tate's trident-nosed bats
      • Stoliczka's Trident Bat, Aselliscus stoliczkanus
      • Temminck's Trident Bat, Aselliscus tricuspidatus
    • Genus Cloeotis
      • Percival's Trident Bat, Cloeotis percivali
    • Genus Coelops ? tailless leaf-nosed bats
      • East Asian Talless Leaf-nosed Bat, Coelops frithii
      • Philippine Tailless Leaf-nosed Bat, Coelops hirsutus
      • Malayan Tailless Leaf-nosed Bat, Coelops robinsoni
    • Genus Hipposideros ? round leaf bats
      • Aba Roundleaf Bat, Hipposideros abae
      • Great Roundleaf Bat, Hipposideros armiger
      • Dusky Roundleaf Bat, Hipposideros ater
      • Benito Roundleaf Bat, Hipposideros beatus
      • Boeadi?s Roundleaf Bat, Hipposideros boeadii
      • Bicolored Roundleaf Bat, Hipposideros bicolor
      • Short-headed Roundleaf Bat, Hipposideros breviceps
      • Sundevall's Roundleaf Bat, Hipposideros caffer
      • Spurred Roundleaf Bat, Hipposideros calcaratus
      • Greater Roundleaf Bat, Hipposideros camerunensis
      • Fawn Roundleaf Bat, Hipposideros cervinus
      • Ashy Roundleaf Bat, Hipposideros cineraceus
      • Commerson's Roundleaf Bat, Hipposideros commersoni
      • Large Mindanao Roundleaf Bat, Hipposideros coronatus
      • Telefomin Roundleaf Bat, Hipposideros corynophyllus
      • Cox's Roundleaf Bat, Hipposideros coxi
      • Timor Roundleaf Bat, Hipposideros crumeniferus
      • Short-tailed Roundleaf Bat, Hipposideros curtus
      • Cyclops Roundleaf Bat, Hipposideros cyclops
      • Makira Roundleaf Bat, Hipposideros demissus
      • Diadem Roundleaf Bat, Hipposideros diadema
      • Fierce Roundleaf Bat, Hipposideros dinops
      • Borneo Roundleaf Bat, Hipposideros doriae
      • Khajuria's Leaf-nosed Bat, Hipposideros durgadasi
      • Dayak Roundleaf Bat, Hipposideros dyacorum
      • Hill's Roundleaf Bat, Hipposideros edwardshilli
      • Hipposideros fasensis
      • Sooty Roundleaf Bat, Hipposideros fuliginosus
      • Fulvus Roundleaf Bat, Hipposideros fulvus
      • Cantor's Roundleaf Bat, Hipposideros galeritus
      • Giant Roundleaf Bat, Hipposideros gigas
      • Grand Roundleaf Bat, Hipposideros grandis
      • Thailand Roundleaf Bat, Hipposideros halophyllus
      • Kolar Leaf-nosed Bat, Hipposideros hypophyllus
      • Crested Roundleaf Bat, Hipposideros inexpectatus
      • Arnhem Leaf-nosed Bat, Hipposideros inornatus
      • Jones's Roundleaf Bat, Hipposideros jonesi
      • Phou Khao Khouay Leaf-nosed Bat, Hipposideros khaokhouayensis
      • Khasian Leaf-nosed Bat, Hipposideros khasiana
      • Lamotte's Roundleaf Bat, Hipposideros lamottei
      • Indian Roundleaf Bat, Hipposideros lankadiva
      • Intermediate Roundleaf Bat, Hipposideros larvatus
      • Large Asian Roundleaf Bat, Hipposideros lekaguli
      • Shield-faced Roundleaf Bat, Hipposideros lylei
      • Big-eared Roundleaf Bat, Hipposideros macrobullatus
      • Maduran Leaf-nosed Bat, Hipposideros madurae
      • Maggie Taylor's Roundleaf Bat, Hipposideros maggietaylorae
      • Aellen's Roundleaf Bat, Hipposideros marisae
      • Ethiopian Large-eared Roundleaf Bat, Hipposideros megalotis
      • Fly River Roundleaf Bat, Hipposideros muscinus
      • Malayan Roundleaf Bat, Hipposideros nequam
      • Philippine Forest Roundleaf Bat, Hipposideros obscurus
      • Orbiculus Leaf-nosed Bat, Hipposideros orbiculus
      • Biak Roundleaf Bat, Hipposideros papua
      • Hipposideros parnabyi
      • Peleng Leaf-nosed Bat, Hipposideros pelingensis
      • Pomona Roundleaf Bat, Hipposideros pomona
      • Pratt's Roundleaf Bat, Hipposideros pratti
      • Philippine Pygmy Roundleaf Bat, Hipposideros pygmaeus
      • Ridley's Leaf-nosed Bat, Hipposideros ridleyi
      • Laotian Leaf-nosed Bat, Hipposideros rotalis
      • Noack's Roundleaf Bat, Hipposideros ruber
      • Hipposideros scutinares
      • Semon's Roundleaf Bat, Hipposideros semoni
      • Sorensen's Leaf-nosed Ba, Hipposideros sorenseni
      • Schneider's Leaf-nosed Bat, Hipposideros speoris
      • Narrow-eared Roundleaf Bat, Hipposideros stenotis
      • Sumban Leaf-nosed Bat, Hipposideros sumbae
      • Hipposideros tephrus
      • Sa? Tom? Leaf-nosed Bat, Hipposideros thomensis
      • Lesser Great Leaf-nosed Bat, Hipposideros turpis
      • Striped Leaf-nosed Bat, Hipposideros vittatus
      • Wollaston's Roundleaf Bat, Hipposideros wollastoni
    • Genus Paracoelops
      • Vietnam Leaf-nosed Bat, Paracoelops megalotis
    • Genus Rhinonicteris
      • Orange Leaf-nosed Bat, Rhinonicteris aurantia
    • Genus Paratriaenops
      • Grandidier's Trident Bat, Paratriaenops auritus
      • Trouessart's Trident Bat, Paratriaenops furculus
      • Paulian's Triple Leaf-nosed Bat, Paratriaenops pauliani
    • Genus Triaenops ? triple leaf-nosed bats
      • Triaenops afer
      • Triaenops menamena
      • Triaenops parvus
      • Persian Trident Bat (Rufous Trident Bat), Triaenops persicus (also known as Triaenops rufus)

Notes

  1. ^ This name technically has priority over Hipposiderinae Lydekker, 1891, and some have consequently used "Rhinonycteridae" or "Rhinonycterinae" for this (sub)family; however, Hipposideridae/inae has been in common use since 1907 and is currently retained pending action by the International Commission on Zoological Nomenclature.[1]

References

  1. ^ a b c Simmons, 2005, p. 365
  2. ^ Hutcheon and Kirsch, 2006
  3. ^ Simmons, 2005, pp. 365?379
  4. ^ McKenna and Bell, 1997, p. 306
  5. ^ McKenna and Bell, 1997, pp. 306?307
  6. ^ Hand and Kirsch, 2003, table 3
  7. ^ Benda and Vallo, 2009, p. 33
  8. ^ Simmons, 2005, pp. 365?379; McKenna and Bell, 1997, pp. 306?307; other sources cited for specific genera
  9. ^ Hand and Kirsch, 2003
  10. ^ Hand and Archer, 2005
  11. ^ a b c d Archer et al., 2006, p. 7
  12. ^ a b Benda and Vallo, 2009, p. 34
  13. ^ Ziegler, 2000, p. 652; Hand and Kirsch, 2003, table 3; cf. McKenna and Bell, 1997, p. 305 (excluded from Rhinonycterinae)

Literature cited

  • Archer, M., Arena, D.A., Bassarova, M., Beck, R.M.D., Black, K., Boles, W.E., Brewer, P., Cooke, B.N., Crosby, K., Gillespie, A., Godthelp, H. , Hand, S.J., Kear, B.P., Louys, J., Morrell, A., Muirhead, J., Roberts, K.K., Scanlon, J.D., Travouillon, K.J. and Wroe, S. 2006. Current status of species-level representation in faunas from selected fossil localities in the Riversleigh World Heritage Area, northwestern Queensland. Alcheringa Special Issue 1:1-17. ISBN 0975789457
  • Benda, P. and Vallo, P. 2009. Taxonomic revision of the genus Triaenops (Chiroptera: Hipposideridae) with description of a new species from southern Arabia and definitions of a new genus and tribe. Folia Zoologica 58(Monograph 1):1?45.
  • Hand, S.J. and Archer, M. 2005. A new hipposiderid genus (Microchiroptera) from an early Miocene bat community in Australia. Palaeontology 48(2):371?383.
  • Hand, S.J. and Kirsch, J.A.W. 2003. Archerops, a new annectent hipposiderid genus (Mammalia: Microchiroptera) from the Australian Miocene. Journal of Paleontology 77(6):1139?1151.
  • Hutcheon, J.M. and Kirsch, J.A.W. 2006. A moveable face: deconstructing the Microchiroptera and a new classification of extant bats. Acta Chiropterologica 8(1):1?10.
  • McKenna, M.C. and Bell, S.K. 1997. Classification of Mammals: Above the species level. New York: Columbia University Press, 631 pp. ISBN 978-0-231-11013-6
  • Simmons, N.B. 2005. Order Chiroptera. Pp. 312?529 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: a taxonomic and geographic reference. 3rd ed. Baltimore: The Johns Hopkins University Press, 2 vols., 2142 pp. ISBN 9780801882210
  • Ziegler, R. 2000. The bats (Chiroptera, Mammalia) from the Late Oligocene fissure fillings Herrlingen 8 and Herrlingen 9 near Ulm (Baden-W?rttemberg). Senckenbergiana Lethaea 80(2):647?683.

Taxonomy

The Family Hipposideridae is further organized into finer groupings including:

Genera

Anthops

The Flower-faced Bat (Anthops ornatus) is a species of bat in the family Hipposideridae. It is in the monotypic genus Anthops.[] It is found in Papua New Guinea and the Solomon Islands. [more]

Archerops

[more]

Aselliscus

Aselliscus is a genus of bat in the family Hipposideridae. It contains the following species: [more]

Cloeotis

Percival's Trident Bat (Cloeotis percivali) is a species of in the Hipposideridae family. It is monotypic within the genus Cloeotis. It is found in Botswana, Democratic Republic of the Congo, Kenya, Mozambique, South Africa, Swaziland, Tanzania, Zambia, and Zimbabwe. Its natural habitats are subtropical or tropical dry forests, dry savanna, and caves. [more]

Coelops

Coelops is a genus of in the Hipposideridae family. It contains the following species: [more]

Rhinonicteris

The Orange Leaf-nosed Bat, Rhinonicteris aurantia, is a species of in the Hipposideridae family. It is monotypic within the genus Rhinonicteris. It is endemic to Australia. [more]

More info about the Genus Rhinonicteris may be found here.

References

  1. ^ a b c Simmons, 2005, p. 365
  2. ^ Hutcheon and Kirsch, 2006
  3. ^ Simmons, 2005, pp. 365?379
  4. ^ McKenna and Bell, 1997, p. 306
  5. ^ McKenna and Bell, 1997, pp. 306?307
  6. ^ Hand and Kirsch, 2003, table 3
  7. ^ Benda and Vallo, 2009, p. 33
  8. ^ Simmons, 2005, pp. 365?379; McKenna and Bell, 1997, pp. 306?307; other sources cited for specific genera
  9. ^ Hand and Kirsch, 2003
  10. ^ Hand and Archer, 2005
  11. ^ a b c d Archer et al., 2006, p. 7
  12. ^ a b Benda and Vallo, 2009, p. 34
  13. ^ Ziegler, 2000, p. 652; Hand and Kirsch, 2003, table 3; cf. McKenna and Bell, 1997, p. 305 (excluded from Rhinonycterinae)

Footnotes

  1. ^ This name technically has priority over Hipposiderinae Lydekker, 1891, and some have consequently used "Rhinonycteridae" or "Rhinonycterinae" for this (sub)family; however, Hipposideridae/inae has been in common use since 1907 and is currently retained pending action by the International Commission on Zoological Nomenclature.[1]

Sources

Last Revised: August 24, 2012
2012/08/24 13:33:36