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Hemichordata

(Phylum)

Overview

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Hemichordata is a of worm-shaped marine deuterostome animals, generally considered the sister group of the echinoderms. They date back to the Lower or Middle Cambrian and include an important class of fossils called graptolites, most of which became extinct in the Carboniferous. They seem to have a primitive form of notochord, formed from a diverticulum of the foregut called a stomochord, but this is most likely the result of convergent evolution rather than homology with the vertebrate notochord. A hollow neural tube exists among some species (at least in early life), probably a primitive trait they share with the common ancestor of chordata and the rest of the deuterostomes.

The bodies of Hemichordates are divided into three parts, proboscis, collar and trunk. They have open circulatory systems and a complete digestive tract but the musculature in their gut is very poorly developed, and food is mostly transported through it by using the cilia that cover its inside surface.

Hemichordata are divided into two classes: the Enteropneusta, commonly called acorn worms, and the Pterobranchia, which may include the graptolites. A third class, Planctosphaeroidea, is proposed based on a single species known only from larvae. The phylum contains about 100 living species. The exact taxonomic position of hemichordata and whether the group is monophyletic is currently under debate. One of the suggestions is that the pterobranchs are more basal deuterostomes, while the enteropneusts are an early offshoot of the lineage who are leading to Chordata.

Taxonomy

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The Phylum Hemichordata is further organized into finer groupings including:

Classes

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Enteropneusta

[more]

Graptolithina

Graptolites (Graptolithina) are colonial animals known chiefly from the Upper Cambrian through the Lower Carboniferous (Mississippian). A possible early graptolite, Chaunograptus, is known from the Middle Cambrian. [more]

Pterobranchia

Pterobranchia is a of small, worm-shaped animals. They belong to the hemichordata, and live in secreted tubes on the ocean floor. Pterobranchia feed by filtering plankton out of the water with the help of cilia attached to tentacles. There are about 30 known living species in the group. [more]

At least 33 species and subspecies belong to the Class Pterobranchia.

More info about the Class Pterobranchia may be found here.

References

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Sources

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Last Revised: September 22, 2009
2009/09/22 06:02:07