Overview
The gobies form the family Gobiidae, which is one of the largest families of fish, with more than 2,000 species in more than 200 genera.1] Most are relatively small, typically less than 10 cm (4 in) in length. Gobies include some of the smallest vertebrates in the world, like species of the genera Trimmatom nanus and Pandaka pygmaea, which are under 1 cm (3/8 in) long when fully grown. There are some large gobies, such as some species of the genera or Periophthalmodon, that can reach over 30 cm (1 ft) in length, but that is exceptional. Although few are important as food for humans, they are of great significance as prey species for commercially important fish like cod, haddock, sea bass, and flatfish. Several gobies are also of interest as aquarium fish, such as the bumblebee gobies of the genus Brachygobius.
Description
The most distinctive aspects of goby morphology are the fused pelvic fins that form a disc-shaped sucker. This sucker is functionally analogous to the dorsal fin sucker possessed by the remoras or the pelvic fin sucker of the lumpsuckers, but is anatomically distinct: these similarities are the product of convergent evolution. Gobies can often be seen using the sucker to adhere to rocks and corals, and in aquariums they will stick to glass walls of the tank, as well.
Subfamilies
Gobiidae contains six subfamilies:
Amblyopinae
Members of the subfamily Amblyopinae are elongated mud-dwelling gobies commonly known as the eel gobies or worm gobies. Their two dorsal fins are connected by a membranous structure and their eyes are highly reduced. They are usually pink, red, or purple in coloration. Amblyopinae contains 12 genera and about 23 species.[2][3]
Benthophilinae
Members of Benthophilinae are endemic to the Ponto-Caspian region (including the Marmara, Black, Azov, Caspian, and Aral seas).[4] The representatives of the subfamily have fused pelvic fins and elongated dorsal and anal fins.[5] They are distinguished from the closely related subfamily Gobiinae by the absence of a swimbladder in adults and location of the uppermost rays of the pectoral fins within the fin membrane.[6] Its members include tadpole gobies, monkey gobies, and bighead gobies.
Gobiinae
Members of Gobiinae are known as true gobies. It is the most widespread and most diverse of the subfamilies under Gobiidae, containing around 2000 species and 150 genera.
Gobionellinae
Members of Gobionellinae mostly inhabit estuarine habitats, though some are freshwater. They are found in tropical and temperate regions around the world with the exception of the northeastern Atlantic Ocean, the Mediterranean Sea, and the Ponto-Caspian region. It includes around 370 species and 55 genera.[7]
Oxudercinae
Members of Oxudercinae are commonly known as mudskippers. They are highly specialized members of the family. They are able to survive for extended periods on land through a combination of behavioral and physiological adaptations, including pectoral fins that act as simple legs; the ability to breathe through their skins (like frogs); and the digging of damp burrows to avoid drying out. Mudskippers live in tidal areas, particularly on mudflats and in mangrove forests, and are only found in tropical and subtropical regions.
Sicydiinae
Sicydiinae is a small subfamily of freshwater gobies. It includes only nine genera.[8] They are usually found in fast-moving mountain streams in tropical islands.[9][10]
Ecology and biology
Gobies are primarily fish of shallow marine habitats including tide pools, coral reefs, and seagrass meadows; they are also very numerous in brackish water and estuarine habitats, including the lower reaches of rivers, mangrove swamps, and salt marshes. A small number of gobies (unknown exactly, but in the low hundreds) are also fully adapted to freshwater environments. These include the Asian river gobies (Rhinogobius spp.), the Australian desert goby (Chlamydogobius eremius), and the European freshwater goby Padogobius bonell i. Most gobies feed on small invertebrates, although some of the larger species eat other fish, and a few eat planktic algae.
Reproduction
Gobies attach their eggs to a substrate, such as vegetation, coral, or a rock surface. They can lay anywhere from five to a few hundred eggs, depending on species. After fertilizing the eggs, the male remains to guard them from predators and keep them free from detritus. The eggs hatch after a few days. The larvae are born transparent, developing their coloration after dispersing to find a suitable habitat. The larvae of many freshwater species are carried downstream to the brackish waters of estuaries, or even to the sea, and only return to fresh water weeks or months later.[11]
Gobies in warmer waters reach adulthood in a matter of months, while those in cooler environments may take up to two years. The total lifespan of gobies varies from a single year to up to ten years, again with the temperate species generally living longer.[11]
A few species of goby, such as the blackeye goby, are known to be able to change sex from female to male, although most do not do this. In such species, most individuals are born female, and the male must expend considerable effort in guarding the eggs of the multiple females with which he breeds.[11]
Symbiosis
Gobies sometimes form symbiotic relationships with other species.[12] Some goby species live in symbiosis with burrowing shrimps. The shrimp maintains a burrow in the sand in which both the shrimp and the goby live. The shrimp has poor eyesight compared to the goby, but if it sees or feels the goby suddenly swim into the burrow, it will follow. The goby and shrimp keep in contact with each other, the shrimp using its antennae, and the goby flicking the shrimp with its tail when alarmed. These gobies are thus sometimes known as watchmen or prawn gobies. Each party gains from this relationship: the shrimp gets a warning of approaching danger, and the goby gets a safe home and a place to lay its eggs. Only the alpha male and female reproduce, other fish in colony eat sparingly to resist being eaten by the alpha male or female. This way only the largest and fittest are able to reproduce.[citation needed]
Another example of symbiosis is demonstrated by the neon gobies (Elacatinus spp.). These gobies are known as "cleaner gobies", and remove parasites from the skin, fins, mouth, and gills of a wide variety of large fish. The most remarkable aspect of this symbiosis is that many of the fish that visit the
gobies' cleaning stations would otherwise treat such small fish as food (for example, groupers and snapper[disambiguation needed
]s). Again, this is a relationship where both parties gain: the gobies get a continual supply of food as bigger fish visit their cleaning stations, and the bigger fish leave the cleaning stations healthier than they were when they arrived.
Commercial importance
Gobies have commercial importance in Ukraine. It is fished in the Sea of Azov, north-western Black Sea. Most important species are round goby, monkey goby, toad goby, and grass goby. The grass goby is also a commercial fish in Italy.
In aquariums
Several species of gobies are kept in aquariums. [13] Most captive gobies are species from saltwater, and make excellent additions to healthy reef or fish-only aquariums. Perhaps the most popular is the small but colorful neon goby. Most gobies stay toward the lower portion of the aquarium, hiding in the rockwork, but some species (most notably the shrimp gobies) prefer to dig themselves little burrows. Potential keepers of these striking fish should provide them with a fine grained substrate to prevent damage to their delicate undersides.
Commonly kept saltwater species include the Randall's shrimp goby, and watchman goby. The bumblebee gobies from the genus Brachygobius are perhaps the most widely traded freshwater species, being small, colorful, and easy to care for. They need tropical, hard and alkaline freshwater or slightly brackish conditions to do well.
Gobies are generally peaceful towards their tankmates, though territorial among themselves. Since most are small and few are predatory toward other fishes, they usually make good community fishes. Typically, the main problem with gobies is feeding them: with a few exceptions, the small species kept in aquariums prefer live on frozen foods rather than flake, and they are not very good at competing with active species such as cichlids. Another problem is that it is very common for them to jump out of the tank so having a tight fitting lid is a must. It is often recommended that gobies be kept on their own or with peaceful surface dwelling species such as halfbeaks and guppies.
See also
- Sleeper gobies are a closely related family (Eleotridae) that lack the fused pelvic fin sucker typical of most gobies, but are otherwise very similar in size, shape, and ecology.
- Blennies are a group of shallow water marine fish often confused with gobies.
- Dragonets are superficially similar to gobies and sometimes confused with them.
- Pholidichthys leucotaenia is commonly called the engineer goby or convict goby but is not a goby.
Gobiinae
Members of Gobiinae are known as true gobies. It is the most widespread and most diverse of the subfamilies under Gobiidae, containing around 2000 species and 150 genera.
Gobionellinae
Members of Gobionellinae mostly inhabit estuarine habitats, though some are freshwater. They are found in tropical and temperate regions around the world with the exception of the northeastern Atlantic Ocean, the Mediterranean Sea, and the Ponto-Caspian region. It includes around 370 species and 55 genera.[7]
Oxudercinae
Members of Oxudercinae are commonly known as mudskippers. They are highly specialized members of the family. They are able to survive for extended periods on land through a combination of behavioral and physiological adaptations, including pectoral fins that act as simple legs; the ability to breathe through their skins (like frogs); and the digging of damp burrows to avoid drying out. Mudskippers live in tidal areas, particularly on mudflats and in mangrove forests, and are only found in tropical and subtropical regions.
Sicydiinae
Sicydiinae is a small subfamily of freshwater gobies. It includes only nine genera.[8] They are usually found in fast-moving mountain streams in tropical islands.[9][10]
Ecology and biology
Gobies are primarily fish of shallow marine habitats including tide pools, coral reefs, and seagrass meadows; they are also very numerous in brackish water and estuarine habitats, including the lower reaches of rivers, mangrove swamps, and salt marshes. A small number of gobies (unknown exactly, but in the low hundreds) are also fully adapted to freshwater environments. These include the Asian river gobies (Rhinogobius spp.), the Australian desert goby (Chlamydogobius eremius), and the European freshwater goby Padogobius bonelli. Most gobies feed on small invertebrates, although some of the larger species eat other fish, and a few eat planktic algae.
Reproduction
Gobies attach their eggs to a substrate, such as vegetation, coral, or a rock surface. They can lay anywhere from five to a fe w hundred eggs, depending on species. After fertilizing the eggs, the male remains to guard them from predators and keep them free from detritus. The eggs hatch after a few days. The larvae are born transparent, developing their coloration after dispersing to find a suitable habitat. The larvae of many freshwater species are carried downstream to the brackish waters of estuaries, or even to the sea, and only return to fresh water weeks or months later.[11]
Gobies in warmer waters reach adulthood in a matter of months, while those in cooler environments may take up to two years. The total lifespan of gobies varies from a single year to up to ten years, again with the temperate species generally living longer.[11]
A few species of goby, such as the blackeye goby, are known to be able to change sex from female to male, although most do not do this. In such species, most individuals are born female, and the male must expend considerable effort in guarding the eggs of the multiple females with which he breeds.[11]
Symbiosis
Gobies sometimes form symbiotic relationships with other species.[12] Some goby species live in symbiosis with burrowing shrimps. The shrimp maintains a burrow in the sand in which both the shrimp and the goby live. The shrimp has poor eyesight compared to the goby, but if it sees or feels the goby suddenly swim into the burrow, it will follow. The goby and shrimp keep in contact with each other, the shrimp using its antennae, and the goby flicking the shrimp with its tail when alarmed. These gobies are thus sometimes known as watchmen or prawn gobies. Each party gains from this relationship: the shrimp gets a warning of approaching danger, and the goby gets a safe home and a place to lay its eggs. Only the alpha male and female reproduce, other fish in colony eat sparingly to resist being eaten by the alpha male or female. This way only the largest and fittest are able to reproduce.[citation needed]
Another example of symbiosis is demonstrated by the neon gobies (Elacatinus spp.). These gobies are known as "cleaner gobies", and remove parasites from the skin, fins, mouth, and gills of a wide variety of large fish. The most remarkable aspect of this symbiosis is that many of the fish
that visit the
gobies' cleaning stations would otherwise treat such small fish as food (for example, groupers and snapper[disambiguation needed
]s). Again, this is a relationship where both parties gain: the gobies get a continual supply of food as bigger fish visit their cleaning stations, and the bigger fish leave the cleaning stations healthier than they were when they arrived.
Commercial importance
Gobies have commercial importance in Ukraine. It is fished in the Sea of Azov, north-western Black Sea. Most important species are round goby, monkey goby, toad goby, and grass goby. The grass goby is also a commercial fish in Italy.
In aquariums
Several species of gobies are kept in aquariums. [13] Most captive gobies are species from saltwater, and make excellent additions to healthy reef or fish-only aquariums. Perhaps the most popular is the small but colorful neon goby. Most gobies stay toward the lower portion of the aquarium, hiding in the rockwork, but some species (most notably the shrimp gobies) prefer to dig themselves little burrows. Potential keepers of these striking fish should provide them with a fine grained substrate to prevent damage to their delicate undersides.
Commonly kept saltwater species include the Randall's shrimp goby, and watchman goby. The bumblebee gobies from the genus Brachygobius are perhaps the most widely traded freshwater species, being small, colorful, and easy to care for. They need tropical, hard and alkaline freshwater or slightly brackish conditions to do well.
Gobies are generally peaceful towards their tankmates, though territorial among themselves. Since most are small and few are predatory toward other fishes, they usually make good community fishes. Typically, the main problem with gobies is feeding them: with a few exceptions, the small species kept in aquariums prefer live on frozen foods rather than flake, and they are not very good at competing with active species such as cichlids. Another problem is that it is very common for them to jump out of the tank so having a tight fitting lid is a must. It is often recommended that gobies be kept on their own or with peaceful surface dwelling species such as halfbeaks and guppies.
See also
- Sleeper gobies are a closely related family (Eleotridae) that lack the fused pelvic fin sucker typical of most gobies, but are otherwise very similar in size, shape, and ecology.
- Blennies are a group of shallow water marine fish often confused with gobies.
- Dragonets are superficially similar to gobies and sometimes confused with them.
- Pholidichthys leucotaenia is commonly called the engineer goby or convict goby but is not a goby.
References
- ^ Froese, Rainer, and Daniel Pauly, eds. (2006). "Gobiidae" in FishBas e. January 2006 version.
- ^ Edward Murdy (2011). "Systematics of Amblyopinae". In B.G. Kapoor. The Biology of Gobies. Science Publishers. pp. 107?118. doi:10.1201/b11397-10. ISBN 9781439862339.
- ^ WoRMS (2012). "Amblyopinae". In Nicolas Bailly. FishBase. World Register of Marine Species. http://www.marinespecies.org/aphia.php?p=taxdetails&id=267093. Retrieved February 27, 2012.
- ^ Simonovic P.D., Nikolic V.P., Sk?ra K.E. (1996) Vertebral number in Ponto-Caspian gobies: phylogenetic relevance. J. Fish Biol., 49: 1027?1029.
- ^ Miller P.J. (1986) Gobiidae. In: Whitehead P.J.P., Bauchot M.-L., Hureau J.-C., Nielsen J., Tortonese E. (eds.) Fishes of the North-eastern Atlantic and the Mediterranean, Vol. 3. UNESCO, Paris.
- ^ Pinchuk V.I. (1991) K voprosu o grupirovkakh vidov v predelakh roda Neogobius (Perciformes). Voprosy Ikhtiologii, 31: 380?393.
- ^ Frank Pezold (2011). "Systematics of the Family Gobionellidae". In Robert Patzner, James L. Van Tassell, Marcelo Kovacic, and B. G. Kapoor. The Biology of Gobies. Science Publishers, Inc.. ISBN 9781578084364. http://sci.tamucc.edu/~lsci/marb/uploads/FishLabPublications/Publication.pdf.
- ^ Keith, P.; Marquet, G.; Taillebois, L. (2011). "Discovery of the freshwater genusSicyopus(Teleostei: Gobioidei: Sicydiinae) in Madagascar, with a description of a new species and comments on regional dispersal". Journal of Natural History 45 (43?44): 2725. doi:10.1080/00222933.2011.602479. edit
- ^ Ronald E. Watson & Maurice Kottelat (2006). "Two new freshwater gobies from Halmahera, Maluku, Indonesia (Teleostei: Gobioidei: Sicydiinae)". Ichthyological Exploration of Freshwaters 17 (2): 121?128.
- ^ Ronald E. Watson, Philippe Keith, & G?rard Marquet (2007). "Akihito vanuatu, a new genus and new species of freshwater goby (Sicydiinae) from the South Paci?c". Cybium 31 (3): 341&?349. http://www.mnhn.fr/sfi/cybium/numeros/313/06.Watson%20448.pdf.
- ^ a b c Hoese, Douglas F. (1998). Paxton, J.R. & Eschmeyer, W.N.. ed. Encyclopedia of Fishes. San Diego: Academic Press. pp. 218?222. ISBN 0-12-547665-5.
- ^ G. S. Helfman, B. B. Colette & D. E. Facey (1997). "Chapter 21: Fishes as social animals". The Diversity of Fishes. Blackwell. ISBN 0-86542-256-7.
- ^ Frank Sch?fer (2005). Brackish-Water Fishes. Aqualog. ISBN 3-936027-82-X (English), ISBN 3-936027-81-1 (German).
External links
Taxonomy
The Family Gobiidae is further organized into finer groupings including:
- Subfamily (3): Amblyopinae · Gobiinae · Sicydiinae
- Genus (216): Aboma · Acanthogobius · Acentrogobius · Afurcagobius · Akihito · Akko · Alepideleotris · Amblugobius · Amblyeleotris · Amblygobius · Amblyopus · Amblyotrypauchen · Amoya · Aphia · Apocryptes · Apocryptodon · Aprolepis · Arcygobius · Arenigobius · Asterropteryx · Aulopareia · Awaous · Awous · Babka · Barbulifer · Bathygobius · Benthophiloides · Benthophilus · Boleophthalmus · Bolephthalmus · Bollmania · Bollmannia · Brachyamblyopus · Bryaninops · Cabillus · Caffrogobius · Callogobius · Caragobius · Chaenogobius · Chaeturichthys · Chaparrudo · Chasmichthys · Chloea · Chonophorus · Chriolepis · Chromogobius · Clariger · Clevelandia · Coryogalops · Coryphopterus · Cotylopus · Creisson · Cristatogobius · Cryptocentorus · Cryptocentroides · Cryptocentrus · Crystallogobius · Ctenogobiops · Ctenogobius · Ctenotrypauchen · Didogobius · Drombus · Elacatinus · Eleotriodes · Eleotroides · Enypnias · Eugnathogobius · Evermannichthys · Eviota · Evorthodus · Exyrias · Favinogobius · Favonigobius · Feia · Flabelligobius · Fusigobius · Gammogobius · Garmannia · Gillichthys · Gladiogobius · Glossogobius · Gnatholepis · Gobiichthys · Gobiidae · Gobiinae · Gobiodon · Gobioides · Gobionellus · Gobiopsis · Gobiopterus · Gobiosoma · Gobius · Gobiusculus · Gorogobius · Grallenia · Gymneleotris · Gymnogobius · Hazeus · Hemigobius · Hetereleotris · Illana · Istigobius · Kelloggella · Knipowitschia · Koumansetta · Larsonella · Latrunculus · Lebetus · Lentipes · Lepidogobius · Leptogobius · Lesueurigobius · Leucopsarion · Lioteres · Lizagobius · Lobulogobius · Lophiogobius · Lophogobius · Lythrypnus · Macrodontogobius · Mahidolia · Mapo · Matiidolia · Mesogobius · Microgobius · Minysicya · Mistichthys · Mugilagobius · Mugilogobius · Myersina · Neogobius · Nesogobius · Obliquogobius · Odontamblyopus · Oligolepis · Ophiogobius · Oplopomops · Oplopomopus · Oplopomus · Opua · Oxuderces · Oxyurichthys · Padogobius · Palutrus · Pandaka · Papillogobius · Paragobiodon · Paragobiopsis · Parapocryptes · Parasicydium · Paratrypauchen · Parella · Parrella · Pascua · Periophthalmodon · Periophthalmus · Pipidonia · Platygobiopsis · Platygobius · Pleurosicya · Pomatoschistus · Pomatotoschistus · Ponticola · Priolepis · Proterorhinus · Pseudapocryptes · Pseudogobiopsis · Pseudogobius · Psilotris · Pterogobius · Quietula · Redigobius · Rhinogobiops · Risor · Sagamia · Salarigobius · Satulinus · Scartelaos · Schismatogobius · Seychellea · Seychellia · Sicydium · Sicyopterus · Sicyopus · Signigobius · Silhouettea · Smaragdus · Stenogobius · Stigmatogobius · Stiphodon · Stonogobiops · Stupidogobius · Synechogobius · Taenioides · Tamanka · Thorogobius · Tigrigobius · Tomiyamichthys · Triaenopogon · Tridentiger · Trimma · Trypauchen · Trypauchenichthys · Trypauchenopsis · Tryssogobius · Tukugobius · Typhlogobius · Valenciennea · Vanderhorstia · Vanneaugobius · Varicus · Weberogobius · Yoga · Yongeichthys · Zappa · Zosterisessor
- Species: ZipcodeZoo has pages for 1,843 species and subspecies in the Family Gobiidae.
Genera
Aboma
Acanthogobius
Acanthogobius is a genus of fish in the family Gobiidae. [more]
Acentrogobius
The Malagasy mountain mullet (Acentrogobius therezieni) is a species of fish in the Gobiidae family and is in the monotypic genus Acentrogobius. It is endemic to Madagascar. Its natural habitat is rivers. It is threatened by habitat loss. [more]
Afurcagobius
Afurcagobius is a small genus of fish in the family Gobiidae. There are two described species. [more]
Akihito
Akko
Acre (Hebrew: ??, Akko; Arabic: ?????, ?Akka), is a city in the Western Galilee region of northern Israel at the northern extremity of Haifa Bay. Acre is one of the oldest continuously inhabited sites in the country. [more]
Alepideleotris
Amblugobius
Amblyeleotris
Amblyeleotris is a genus of colorful gobies found throughout the Indo-Pacific region. With 37 recognized species, this is the largest genus of the shrimp gobies or prawn gobies, so-called because of their symbiotic relationship with certain alpheid shrimps. The shrimp excavates and maintains a burrow used by both animals while the goby, which has far superior eyesight, acts as a lookout for predators. The shrimp maintains almost constant contact with the fish with an antenna. [more]
Amblygobius
Amblygobius is a genus of fish in the family Gobiidae. [more]
Amblyopus
A Genus in the Kingdom Animalia.[1] [more]
Amblyotrypauchen
Amblyotrypauchen fraseri is the only member of genus Amblyotrypauchen. [more]
Amoya
Amoya is a genus of fish in the family Gobiidae. [more]
Aphia
The transparent goby, Aphia minuta, is a fish species of the Gobiidae family. It is the only member of the genus Aphia. [more]
Apocryptes
A Genus in the Kingdom Animalia.[2] [more]
Apocryptodon
A Genus in the Kingdom Animalia.[3] [more]
Aprolepis
Arcygobius
Arenigobius
A Genus in the Kingdom Animalia.[4] [more]
Asterropteryx
A Genus in the Kingdom Animalia.[5] [more]
Aulopareia
A Genus in the Kingdom Animalia.[6] [more]
Awaous
Awaous is a genus of fish in the Gobiidae family. It contains the following species: [more]
Awous
Babka
Barbulifer
Bathygobius
Bathygobius burtoni is a species of in the Gobiidae family. It is endemic to Sao Tomé and Principe. [more]
Benthophiloides
Benthophiloides is a genus of Ponto-Caspian gobiid fishes. Costents two species, widespread in the basins of the Black Sea and the Caspian Sea: [more]
Benthophilus
A Genus in the Kingdom Animalia.[7] [more]
Boleophthalmus
A Genus in the Kingdom Animalia.[8] [more]
Bolephthalmus
Bollmania
Bollmannia
Brachyamblyopus
A Genus in the Kingdom Animalia.[9] [more]
Bryaninops
A Genus in the Kingdom Animalia.[10] [more]
Cabillus
A Genus in the Kingdom Animalia.[11] [more]
Caffrogobius
A Genus in the Kingdom Animalia.[12] [more]
Callogobius
A Genus in the Kingdom Animalia.[13] [more]
Caragobius
Chaenogobius
A Genus in the Kingdom Animalia.[14] [more]
Chaeturichthys
A Genus in the Kingdom Animalia.[15] [more]
Chaparrudo
Chasmichthys
Chloea
A Genus in the Kingdom Animalia.[16] [more]
Chonophorus
A Genus in the Kingdom Animalia.[17] [more]
Chriolepis
A Genus in the Kingdom Animalia.[18] [more]
Chromogobius
A Genus in the Kingdom Animalia.[19] [more]
Clariger
A Genus in the Kingdom Animalia.[20] [more]
Clevelandia
A Genus in the Kingdom unknown!.[21] [more]
Coryogalops
A Genus in the Kingdom Animalia.[22] [more]
Coryphopterus
Cotylopus
A Genus in the Kingdom Animalia.[23] [more]
Creisson
A Genus in the Kingdom Animalia.[24] [more]
Cristatogobius
A Genus in the Kingdom Animalia.[25] [more]
Cryptocentorus
Cryptocentroides
A Genus in the Kingdom Animalia.[26] [more]
Cryptocentrus
A Genus in the Kingdom Animalia.[27] [more]
Crystallogobius
A Genus in the Kingdom Animalia.[28] [more]
Ctenogobiops
Ctenogobiops is a genus of saltwater fish in the family Gobiidae (commonly known as gobies). [more]
Ctenogobius
A Genus in the Kingdom Animalia.[29] [more]
Ctenotrypauchen
A Genus in the Kingdom Animalia.[30] [more]
Didogobius
A Genus in the Kingdom Animalia.[31] [more]
Drombus
A Genus in the Kingdom Animalia.[32] [more]
Elacatinus
Elacatinus (formerly Gobiosoma) is a genus of marine gobies, often known collectively as the neon gobies. Although only one species, E. oceanops, is technically the "neon goby," because of their extremely similar appearance, other members of the genus are generally labeled neon gobies, as well. [more]
Eleotriodes
Eleotroides
A Genus in the Kingdom Animalia.[33] [more]
Enypnias
Eugnathogobius
Evermannichthys
Eviota
Eviota is a genus of reef-associated gobies currently with 48 described species distributed throughout the Indo-Pacific region. These are very small fishes (females of most species are sexually mature at a standard length of 15 mm or less) and many species are commonly called pygmy gobies, while some are known as fringefin gobies. [more]
Evorthodus
Exyrias
Exyrias is a genus of marine with four described species found throughout the Indian and Pacific Oceans. These are relatively large gobies, typically found in turbid estuarine waters (although the recently described Exyrias akihito is a species of clearer water associated with coral reefs). [more]
Favinogobius
Favonigobius
Favonigobius is a of gobies in the family Gobiidae. [more]
Feia
A Genus in the Kingdom Animalia.[34] [more]
Flabelligobius
A Genus in the Kingdom Animalia.[35] [more]
Fusigobius
Fusigobius is a genus of coral reef inhabiting gobies found throughout the Indian and Pacific Oceans. [more]
Gammogobius
Garmannia
A Genus in the Kingdom Animalia.[36] [more]
Gillichthys
Gillichthys is a of gobies also known as "mudsuckers". [more]
Gladiogobius
A Genus in the Kingdom Animalia.[37] [more]
Glossogobius
Glossogobius is a genus of in the Gobiidae family. It contains the following species: [more]
Gnatholepis
Gobiichthys
Gobiidae
Gobiinae
Gobiodon
Gobioides is a genus of gobies also known as coral gobies or "clown gobies" (which can also mean the related genus ). Generally, coral gobies, unlike the rest of the Gobiidae family, are not burrowers, but instead prefer to inhabit the branches of certain acropora or similar hard corals. [more]
Gobioides
Gobionellus
Gobiopsis
Gobiopsis is a of gobies of the family Gobiidae. [more]
Gobiopterus
A Genus in the Kingdom Animalia.[38] [more]
Gobiosoma
Gobius
Gobius is a genus of fish in the Gobiidae family. It contains the typical gobies, being the type genus of its subfamily (Gobiinae) and family, and the namesake genus of its suborder (Gobioidei). [more]
Gobiusculus
A Genus in the Kingdom Animalia.[39] [more]
Gorogobius
A Genus in the Kingdom Animalia.[40] [more]
Grallenia
Gymneleotris
A Genus in the Kingdom Animalia.[41] [more]
Gymnogobius
A Genus in the Kingdom Animalia.[42] [more]
Hazeus
A Genus in the Kingdom Animalia.[43] [more]
Hemigobius
A Genus in the Kingdom Animalia.[44] [more]
Hetereleotris
A Genus in the Kingdom Animalia.[45] [more]
Illana
A Genus in the Kingdom Animalia. [more]
Istigobius
A Genus in the Kingdom Animalia.[46] [more]
Kelloggella
A Genus in the Kingdom Animalia.[47] [more]
Knipowitschia
Knipowitschia is a genus of in the Gobiidae family. It contains the following species: [more]
Koumansetta
Larsonella
Latrunculus
Lebetus
A Genus in the Kingdom Animalia.[48] [more]
Lentipes
Lentipes is a genus of in the Gobiidae family. It contains the following species: [more]
Lepidogobius
Leptogobius
Lesueurigobius
A Genus in the Kingdom Animalia.[49] [more]
Leucopsarion
A Genus in the Kingdom Animalia.[50] [more]
Lioteres
Lizagobius
Lobulogobius
A Genus in the Kingdom Animalia.[51] [more]
Lophiogobius
Lophogobius
Lythrypnus
Macrodontogobius
A Genus in the Kingdom Animalia.[52] [more]
Mahidolia
A Genus in the Kingdom Animalia.[53] [more]
Mapo
A Genus in the Kingdom Animalia. [more]
Matiidolia
Mesogobius
Mesogobius is a genus of in the Gobiidae family. It contains the following species: [more]
Microgobius
Minysicya
Mistichthys
A Genus in the Kingdom Animalia.[54] [more]
Mugilagobius
Mugilogobius
Mugilogobius is a genus of in the Gobiidae family. It contains the following species: [more]
Myersina
A Genus in the Kingdom Animalia.[55] [more]
Neogobius
Neogobius is a genus of in the Gobiidae family. It contains the following species: [more]
Nesogobius
A Genus in the Kingdom Animalia.[56] [more]
Obliquogobius
Odontamblyopus
Odontamblyopus is a genus of with five described species found in mud bottom coastal and estuarine habitats from Pakistan eastward to Japan. They have long, slender bodies and are commonly called eel gobies or worm gobies. [more]
Oligolepis
Oligolepis is a genus of in the Gobiidae family. It contains the following species: [more]
Ophiogobius
A Genus in the Kingdom Animalia.[57] [more]
Oplopomops
A Genus in the Kingdom Animalia.[58] [more]
Oplopomopus
Oplopomus
A Genus in the Kingdom Animalia.[59] [more]
Opua
A Genus in the Kingdom Animalia.[60] [more]
Oxuderces
A Genus in the Kingdom Animalia.[61] [more]
Oxyurichthys
Padogobius
Padogobius is a genus of in the Gobiidae family. It contains the following species: [more]
Palutrus
Pandaka
A Genus in the Kingdom Animalia.[62] [more]
Papillogobius
Papillogobius is a genus of in the Gobiidae family. It contains the following species: [more]
Paragobiodon
A Genus in the Kingdom Animalia.[63] [more]
Paragobiopsis
Parapocryptes
A Genus in the Kingdom Animalia.[64] [more]
Parasicydium
Paratrypauchen
Parella
A Genus in the Kingdom Animalia. [more]
Parrella
A Genus in the Kingdom Animalia.[65] [more]
Pascua
Periophthalmodon
A Genus in the Kingdom Animalia.[66] [more]
Periophthalmus
Pipidonia
Platygobiopsis
A Genus in the Kingdom Animalia.[67] [more]
Platygobius
Pleurosicya
A Genus in the Kingdom Animalia.[68] [more]
Pomatoschistus
Pomatoschistus is a genus of fish in the Gobiidae family. It contains 12 species: [more]
Pomatotoschistus
Ponticola
Priolepis
Priolepis is a genus of in the Gobiidae family. It contains the following species: [more]
Proterorhinus
Proterorhinus is a small genus of in the Gobiidae family. [more]
Pseudapocryptes
A Genus in the Kingdom Animalia.[69] [more]
Pseudogobiopsis
A Genus in the Kingdom Animalia.[70] [more]
Pseudogobius
Psilotris
A Genus in the Kingdom Animalia.[71] [more]
Pterogobius
A Genus in the Kingdom Animalia.[72] [more]
Quietula
Redigobius
Redigobius is a genus of in the Gobiidae family. It contains the following species: [more]
Rhinogobiops
A Genus in the Kingdom Animalia.[73] [more]
Risor
Sagamia
A Genus in the Kingdom Animalia.[74] [more]
Salarigobius
Satulinus
Scartelaos
A Genus in the Kingdom Animalia.[75] [more]
Schismatogobius
Schismatogobius is a genus of freshwater gobies currently with 10 described species found on Indo-Pacific islands from Sri Lanka to Fiji. [more]
Seychellea
Seychellia
Sicydium
A Genus in the Kingdom unknown!.[76] [more]
Sicyopterus
Sicyopterus is a genus of in the Gobiidae family. It contains the following species: [more]
Sicyopus
Sicyopus is a genus of in the Gobiidae family. It contains the following species: [more]
Signigobius
A Genus in the Kingdom Animalia.[77] [more]
Silhouettea
Silhouettea is a genus of in the Gobiidae family. [more]
Smaragdus
Stenogobius
Stenogobius is a genus of in the Gobiidae family. It contains the following species: [more]
Stigmatogobius
A Genus in the Kingdom Animalia.[78] [more]
Stiphodon
Stiphodon is a genus of amphidromous freshwater gobies. They inhabit swift, clear streams close to the sea and are found in large parts of Asia and Oceania. Many of the 30 currently recognized species have extremely restricted distributions on single islands or even single streams. the fish are common around Japan, Samoa, Cook Island and the Philippines. [more]
Stonogobiops
A Genus in the Kingdom Animalia.[79] [more]
Stupidogobius
Synechogobius
Taenioides
Taenioides is a genus of in the Gobiidae family. It contains the following species: [more]
Tamanka
Tamanka is a genus of in the Gobiidae family. It contains the following species: [more]
Thorogobius
A Genus in the Kingdom Animalia.[80] [more]
Tigrigobius
Tomiyamichthys
A Genus in the Kingdom Animalia.[81] [more]
Triaenopogon
Tridentiger
Trimma
A Genus in the Kingdom Animalia.[82] [more]
Trypauchen
A Genus in the Kingdom Animalia.[83] [more]
Trypauchenichthys
A Genus in the Kingdom Animalia.[84] [more]
Trypauchenopsis
Tryssogobius
Tukugobius
A Genus in the Kingdom Animalia.[85] [more]
Typhlogobius
Valenciennea
Valenciennea is a genus of small, bottom-dwelling fish from the family Gobiidae. The members of the genus tend to rest directly on the substrate for extended periods of time. While this is a common behavior for members of the family, the Valenciennea also float motionless directly above the substrate, which is why they are sometimes called hover gobies. Their resting behavior has resulted in the vernacular name "sleeper gobies", which invites confusion with the related family Eleotridae. [more]
Vanderhorstia
A Genus in the Kingdom Animalia.[86] [more]
Vanneaugobius
A Genus in the Kingdom Animalia.[87] [more]
Varicus
A Genus in the Kingdom Animalia.[88] [more]
Weberogobius
Weberogobius is a genus of in the Gobiidae family. It contains the following species: [more]
Yoga
Yongeichthys
Zappa
A Genus in the Kingdom Animalia.[89] [more]
Zosterisessor
Zosterisessor is a genus of in the Gobiidae family. It contains the following species: [more]
At least 4 species and subspecies belong to the Genus Zosterisessor.
More info about the Genus Zosterisessor may be found here.
References
- ^ Froese, Rainer, and Daniel Pauly, eds. (2006). "Gobiidae" in FishBase. January 2006 version.
- ^ Edward Murdy (2011). "Systematics of Amblyopinae". In B.G. Kapoor. The Biology of Gobies. Science Publishers. pp. 107?118. doi:10.1201/b11397-10. ISBN 9781439862339.
- ^ WoRMS (2012). "Amblyopinae". In Nicolas Bailly. FishBase. World Register of Marine Species. http://www.marinespecies.org/aphia.php?p=taxdetails&id=267093. Retrieved February 27, 2012.
- ^ Simonovic P.D., Nikolic V.P., Sk?ra K.E. (1996) Vertebral number in Ponto-Caspian gobies: phylogenetic relevance. J. Fish Biol., 49: 1027?1029.
- ^ Miller P.J. (1986) Gobiidae. In: Whitehead P.J.P., Bauchot M.-L., Hureau J.-C., Nielsen J., Tortonese E. (eds.) Fishes of the North-eastern Atlantic and the Mediterranean, Vol. 3. UNESCO, Paris.
- ^ Pinchuk V.I. (1991) K voprosu o grupirovkakh vidov v predelakh roda Neogobius (Perciformes). Voprosy Ikhtiologii, 31: 380?393.
- ^ Frank Pezold (2011). "Systematics of the Family Gobionellidae". In Robert Patzner, James L. Van Tassell, Marcelo Kovacic, and B. G. Kapoor. The Biology of Gobies. Science Publishers, Inc.. ISBN 9781578084364. http://sci.tamucc.edu/~lsci/marb/uploads/FishLabPublications/Publication.pdf.
- ^ Keith, P.; Marquet, G.; Taillebois, L. (2011). "Discovery of the freshwater genusSicyopus(Teleostei: Gobioidei: Sicydiinae) in Madagascar, with a description of a new species and comments on regional dispersal". Journal of Natural History 45 (43?44): 2725. doi:10.1080/00222933.2011.602479. edit
- ^ Ronald E. Watson & Maurice Kottelat (2006). "Two new freshwater gobies from Halmahera, Maluku, Indonesia (Teleostei: Gobioidei: Sicydiinae)". Ichthyological Exploration of Freshwaters 17 (2): 121?128.
- ^ Ronald E. Watson, Philippe Keith, & G?rard Marquet (2007). "Akihito vanuatu, a new genus and new species of freshwater goby (Sicydiinae) from the South Paci?c". Cybium 31 (3): 341&?349. http://www.mnhn.fr/sfi/cybium/numeros/313/06.Watson%20448.pdf.
- ^ a b c Hoese, Douglas F. (1998). Paxton, J.R. & Eschmeyer, W.N.. ed. Encyclopedia of Fishes. San Diego: Academic Press. pp. 218?222. ISBN 0-12-547665-5.
- ^ G. S. Helfman, B. B. Colette & D. E. Facey (1997). "Chapter 21: Fishes as social animals". The Diversity of Fishes. Blackwell. ISBN 0-86542-256-7.
- ^ Frank Sch?fer (2005). Brackish-Water Fishes. Aqualog. ISBN 3-936027-82-X (English), ISBN 3-936027-81-1 (German).
Bibliography
- Guo YX, Kawasaki M, The Journal of neuroscience : the official journal of the Society for Neuroscience. 1997 Mar;17(5):1761-8.
Footnotes
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Sources
- The text on this page is licensed under the GNU Free Documentation License. It includes material from Wikipedia retrieved Wednesday, April 25, 2012.
- The distribution map on the Distribution tab comes from the Global Biodiversity Information Facility and is used with permission.
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