Overview
The pocket gophers are burrowing rodents of the family Geomyidae. These are the "true" gophers, though several ground squirrels of the family Sciuridae are often called gophers as well. The name "pocket gopher" on its own may be used to refer to any of a number of subspecies of the family.
Pocket gophers are widely distributed in North America, extending into Central America.
Appearance
Gophers are heavily built, and most are 12 to 30 cm (4.7 to 12 in) long, weighing a few hundred grams. A few species reach weights approaching 1 kg (2.2 lb). Within any species, the males are larger than the females and can be nearly double their weight.[1] Most gophers have brown fur that often closely matches the color of the soil in which they live. Their most characte ristic features are their large cheek pouches, from which the word "pocket" in their name derives. These pouches are fur-lined, and can be turned inside out. They extend from the side of the mouth well back onto the shoulders. They have small eyes and a short, hairy tail, which they use to feel around tunnels when they walk backwards.
Behavior
All pocket gophers are burrowers. They are larder hoarders, and their cheek pouches are used for transporting food back to their burrows. Gophers can collect large hoards. Their presence is unambiguously announced by the appearance of mounds of fresh dirt about 20 cm (7.9 in) in diameter. These mounds will often appear in vegetable gardens, lawns, or farms, as gophers like moist soil (see Soil biomantle). They also enjoy feeding on vegetables. For this reason, some species are considered agricultural pests. They may also damage trees in forests. Although they will attempt to flee w hen threatened, they may attack other animals, including cats and humans, and can inflict serious bites with their long, sharp teeth.
Pocket gophers are solitary outside of the breeding season, aggressively maintaining territories that vary in size depending on the resources available. Males and females may share some burrows and nesting chambers if their territories border each other, but in general, each pocket gopher inhabits its own individual tunnel system.
Depending on the species and local conditions, pocket gophers may have a specific annual breeding season, or may breed repeatedly through the year. Each litter typically consists of two to five young, although this may be much higher in some species. The young are born blind and helpless, and are weaned at around forty days.[2]
Classification
There has been much debate among taxonomists about which races of pocket gopher should be recognised as full species, and the following list cannot be regarded as definitive.
- Family Geomyidae
- Genus Cratogeomys; some authors treat this genus as a subgenus of Pappogeomys.
- Yellow-faced Pocket Gopher (Cratogeomys castanops)
- Oriental Basin Pocket Gopher (C. fulvescens)
- Smoky Pocket Gopher (C. fumosus)
- Llano Pocket Gopher (C. gymnurus)
- Merriam?s Pocket Gopher (C. merriami)
- Genus Geomys - eastern pocket gophers; principally found in the south-western United States, east of the Sierra Nevada mountains.
- Geomys arenarius; two subspecies, the Desert and White Sands Pocket Gophers
- Attwater's Pocket Gopher (G. attwateri)
- Plains Pocket Gopher (G. bursarius); two subspecies
- Jones' Pocket Goph er (G. knoxjonesi)
- Geomys personatus; 5 subspecies including the Texas, Davis, Maritime and Carrizo Springs Pocket Gophers
- Geomys pinetis; 4 subspecies, the Southeastern, Cumberland Island, Sherman's and Goff's Pocket Gophers
- Geomys texensis; 2 subspecies, including the LLano Pocket Gopher
- Genus Orthogeomys - giant pocket gophers or taltuzas; found in Mexico, Central America and Colombia.
- Chiriqui Pocket Gopher (Orthogeomys cavator)
- Cherrie?s Pocket Gopher (O. cherriei)
- Oaxacan Pocket Gopher (O. cuniculus)
- Darien Pocket Gopher (O. dariensis)
- Giant Pocket Gopher (O. grandis)
- Variable Pocket Gopher (O. heterodus)
- Hispid Pocket Gopher (O. hispidus)
- Big Pocket Gopher (O. lanius)
- Nicaraguan Pocket Gopher (O. matagalpae)
- Thaeler?s Pocket Gopher (O. thaeleri)
- Underwood?s Pocket Gopher ( O. underwoodi)
- Genus Pappogeomys; found in Mexico.
- Alcorn?s Pocket Gopher (Pappogeomys alcorni)
- Buller?s Pocket Gopher (P. bulleri)
- Genus Thomomys - western pocket gophers; widely distributed in North America, extending into the northwestern US, Canada and the southeastern US.
- Thomomys bottae; many subspecies, including the Botta's, Fish Spring, Bonneville, Clear Lake, San Antonio, Pistol River, Mount Ellen, Guadalupe, Limpia, Mearns', Stansbury Island, Antelope Island, Cebolleta, Salinas, Skull Valley, Swasey Springs, Harquahala and Limpia Greek Pocket Gophers.
- Camas Pocket Gopher (T. bulbivorus)
- Wyoming Pocket Gopher (T. clusius)
- Idaho Pocket Gopher (T. idahoensis)
- Mazama Pocket Gopher (T. mazama); several subspecies including the Western, Gold Beach, Olympic, and Tacoma Pocket Gophers.
- Mountain Pocket Gopher (T. monticola)
- Nor thern Pocket Gopher (T. talpoides); very widely distributed; several subspecies including the Cheyenne Northern Pocket Gopher
- Townsend?s Pocket Gopher (T. townsendii)
- Southern Pocket Gopher (T. umbrinus)
- Genus Zygogeomys
- Michoacan Pocket Gopher or Tuza (Zygogeomys trichopus)
- Genus Cratogeomys; some authors treat this genus as a subgenus of Pappogeomys.
Some sources also list a genus Hypogeomys, with one species, but this genus name is normally used for the Malagasy Giant Rat, which belongs to the family Nesomyidae.
Control
Geomys spp and Thomomys spp are classed as "prohibited new organisms" under New Zealand's Hazardous Substances and New Organisms Act 1996 preventing it from being imported into the country.[3]
ur-lined, and can be turned inside out. They extend from the side of the mouth well back onto th e shoulders. They have small eyes and a short, hairy tail, which they use to feel around tunnels when they walk backwards.Behavior
All pocket gophers are burrowers. They are larder hoarders, and their cheek pouches are used for transporting food back to their burrows. Gophers can collect large hoards. Their presence is unambiguously announced by the appearance of mounds of fresh dirt about 20 cm (7.9 in) in diameter. These mounds will often appear in vegetable gardens, lawns, or farms, as gophers like moist soil (see Soil biomantle). They also enjoy feeding on vegetables. For this reason, some species are considered agricultural pests. They may also damage trees in forests. Although they will attempt to flee when threatened, they may attack other animals, including cats and humans, and can inflict serious bites with their long, sharp teeth.
Pocket gophers are solitary outside of the breeding season, aggressively maintaining territories that vary in size depending on the resources available. Males and females may share some burrows and nesting chambers if their territories border each other, but in general, each pocket gopher inhabits its own individual tunnel system.
Depending on the species and local conditions, pocket gophers may have a specific annual breeding season, or may breed repeatedly through the year. Each litter typically consists of two to five young, although this may be much higher in some species. The young are born blind and helpless, and are weaned at around forty days.[2]
Classification
There has been much debate among taxonomists about which races of pocket gopher should be recognised as full species, and the following list cannot be regarded as defin itive.
- Family Geomyidae
- Genus Cratogeomys; some authors treat this genus as a subgenus of Pappogeomys.
- Yellow-faced Pocket Gopher (Cratogeomys castanops)
- Oriental Basin Pocket Gopher (C. fulvescens)
- Smoky Pocket Gopher (C. fumosus)
- Llano Pocket Gopher (C. gymnurus)
- Merriam?s Pocket Gopher (C. merriami)
- Genus Geomys - eastern pocket gophers; principally found in the south-western United States, east of the Sierra Nevada mountains.
- Geomys arenarius; two subspecies, the Desert and White Sands Pocket Gophers
- Attwater's Pocket Gopher (G. attwateri)
- Plains Pocket Gopher (G. bursarius); two subspecies
- Jones' Pocket Gopher (G. knoxjonesi)
- Geomys personatus; 5 subspecies including the Texas, Davis, Maritime and Carrizo Springs Pocket Gophers
- Geomys pinetis; 4 subspecies, the Southeastern, Cumber land Island, Sherman's and Goff's Pocket Gophers
- Geomys texensis; 2 subspecies, including the LLano Pocket Gopher
- Genus Orthogeomys - giant pocket gophers or taltuzas; found in Mexico, Central America and Colombia.
- Chiriqui Pocket Gopher (Orthogeomys cavator)
- Cherrie?s Pocket Gopher (O. cherriei)
- Oaxacan Pocket Gopher (O. cuniculus)
- Darien Pocket Gopher (O. dariensis)
- Giant Pocket Gopher (O. grandis)
- Variable Pocket Gopher (O. heterodus)
- Hispid Pocket Gopher (O. hispidus)
- Big Pocket Gopher (O. lanius)
- Nicaraguan Pocket Gopher (O. matagalpae)
- Thaeler?s Pocket Gopher (O. thaeleri)
- Underwood?s Pocket Gopher (O. underwoodi)
- Genus Pappogeomys; found in Mexico.
- Alcorn?s Pocket Gopher (Pappogeomys alcorni)
- Buller?s Pocket Gopher (P. bulleri)
- Gen
us Thomomys - western pocket gophers; widely distributed in North America, extending into the northwestern US, Canada and the southeastern US.
- Thomomys bottae; many subspecies, including the Botta's, Fish Spring, Bonneville, Clear Lake, San Antonio, Pistol River, Mount Ellen, Guadalupe, Limpia, Mearns', Stansbury Island, Antelope Island, Cebolleta, Salinas, Skull Valley, Swasey Springs, Harquahala and Limpia Greek Pocket Gophers.
- Camas Pocket Gopher (T. bulbivorus)
- Wyoming Pocket Gopher (T. clusius)
- Idaho Pocket Gopher (T. idahoensis)
- Mazama Pocket Gopher (T. mazama); several subspecies including the Western, Gold Beach, Olympic, and Tacoma Pocket Gophers.
- Mountain Pocket Gopher (T. monticola)
- Northern Pocket Gopher (T. talpoides); very widely distributed; several subspecies including the Cheyenne Northern Pocket Gopher
- Townsend?s Pocket Gopher (T. townsendii)
- Southern Pocket G opher (T. umbrinus)
- Genus Zygogeomys
- Michoacan Pocket Gopher or Tuza (Zygogeomys trichopus)
- Genus Cratogeomys; some authors treat this genus as a subgenus of Pappogeomys.
Some sources also list a genus Hypogeomys, with one species, but this genus name is normally used for the Malagasy Giant Rat, which belongs to the family Nesomyidae.
Control
Geomys spp and Thomomys spp are classed as "prohibited new organisms" under New Zealand's Hazardous Substances and New Organisms Act 1996 preventing it from being imported into the country.[3]
References
- ^ Macdonald (Ed), Professor David W. (2006). The Encyclopedia of Mammals. Oxford Univer sity Press. ISBN 0-19-920608-2.
- ^ Patton, James (1984). Macdonald, D.. ed. The Encyclopedia of Mammals. New York: Facts on File. pp. 628?631. ISBN 0-87196-871-1.
- ^ Hazardous Substances and New Organisms Act 2003 - Schedule 2 Prohibited new organisms, New Zealand Government, http://legislation.govt.nz/act/public/1996/0030/latest/DLM386556.html#DLM386556, retrieved 26 January 2012
External links
Taxonomy
The Family Geomyidae is a member of the Superfamily Geomyoidea. Here is the complete "parentage" of Geomyidae:
- Domain: Eukaryota
Whittaker & Margulis,1978 - eukaryotes
- Kingdom: Animalia
C. Linnaeus, 1758 - animals
- Subkingdom: Bilateria
(Hatschek, 1888) Cavalier-Smith, 1983 - bilaterians
- Branch: Deuterostomia
Grobben, 1908 - Deuterostomes
- Infrakingdom: Chordonia
(Haeckel, 1874) Cavalier-Smith, 1998
- Phylum: Chordata
Bateson, 1885 - Chordates
- Subphylum: Vertebrata
Cuvier, 1812 - Vertebrates
- Infraphylum: Gnathostomata
auct. - Jawed Vertebrates
- Superclass: Tetrapoda
Goodrich, 1930 - Tetrapods
- Class: Mammalia
C. Linnaeus, 1758 - Mammals
- Subclass: Theriiformes
(Rowe, 1988) M.C. McKenna & S.K. Bell, 1997
- Infraclass: Holotheria
(Wible et al., 1995) M.C. McKenna & S.K. Bell, 1997
- Superlegion: Trechnotheria
McKenna, 1975
- Legion: Cladotheria
McKenna, 1975
- Sublegion: Zatheria
McKenna, 1975
- Infralegion: Tribosphenida
(McKenna, 1975) M.C. McKenna & S.K. Bell, 1997
- Supercohort: Theria
(Parker & Haswell, 1897) M.C. McKenna & S.K. Bell, 1997 - a genus of Click Beetles (Elateridae)
- Cohort: Placentalia
(Owen, 1837) M.C. McKenna & S.K. Bell, 1997 - Placentals
- Magnorder: Epitheria
(Mckenna, 1975) M.c. Mckenna & S.k. Bell, 1997
- Superorder: Preptotheria
(McKenna, 1975) McKenna, in Stucky & McKenna, in Benton, ed., 1993
- Grandorder: Anagalida
(Szalay & McKenna, 1971) McKenna, 1975
- Mirorder: Simplicidentata
(Weber, 1904) M.C. McKenna & S.K. Bell, 1997
- Order: Rodentia
Bowdich, 1821 - Rodents
- Suborder: Myomorpha
Brandt, 1855 - Mice, Rats
- Infraorder: Geomorpha
(Thaler, 1966) M.C. McKenna & S.K. Bell, 1997
- Superfamily: Geomyoidea
(Bonaparte, 1845) Weber, 1904
- Family: Geomyidae (Bonaparte, 1845) Gill, 1872 - pocket gophers
- Superfamily: Geomyoidea
(Bonaparte, 1845) Weber, 1904
- Infraorder: Geomorpha
(Thaler, 1966) M.C. McKenna & S.K. Bell, 1997
- Suborder: Myomorpha
Brandt, 1855 - Mice, Rats
- Order: Rodentia
Bowdich, 1821 - Rodents
- Mirorder: Simplicidentata
(Weber, 1904) M.C. McKenna & S.K. Bell, 1997
- Grandorder: Anagalida
(Szalay & McKenna, 1971) McKenna, 1975
- Superorder: Preptotheria
(McKenna, 1975) McKenna, in Stucky & McKenna, in Benton, ed., 1993
- Magnorder: Epitheria
(Mckenna, 1975) M.c. Mckenna & S.k. Bell, 1997
- Cohort: Placentalia
(Owen, 1837) M.C. McKenna & S.K. Bell, 1997 - Placentals
- Supercohort: Theria
(Parker & Haswell, 1897) M.C. McKenna & S.K. Bell, 1997 - a genus of Click Beetles (Elateridae)
- Infralegion: Tribosphenida
(McKenna, 1975) M.C. McKenna & S.K. Bell, 1997
- Sublegion: Zatheria
McKenna, 1975
- Legion: Cladotheria
McKenna, 1975
- Superlegion: Trechnotheria
McKenna, 1975
- Infraclass: Holotheria
(Wible et al., 1995) M.C. McKenna & S.K. Bell, 1997
- Subclass: Theriiformes
(Rowe, 1988) M.C. McKenna & S.K. Bell, 1997
- Class: Mammalia
C. Linnaeus, 1758 - Mammals
- Superclass: Tetrapoda
Goodrich, 1930 - Tetrapods
- Infraphylum: Gnathostomata
auct. - Jawed Vertebrates
- Subphylum: Vertebrata
Cuvier, 1812 - Vertebrates
- Phylum: Chordata
Bateson, 1885 - Chordates
- Infrakingdom: Chordonia
(Haeckel, 1874) Cavalier-Smith, 1998
- Branch: Deuterostomia
Grobben, 1908 - Deuterostomes
- Subkingdom: Bilateria
(Hatschek, 1888) Cavalier-Smith, 1983 - bilaterians
- Kingdom: Animalia
C. Linnaeus, 1758 - animals
The Family Geomyidae is further organized into finer groupings including:
- Subfamily (2): Geomyinae · Heteromyinae
- Tribe (5): Dipodomyini · Geomyini · Heteromyini · Perognathini · Thomomyini
- Genus (46): Balantiomys · Chaetodipus · Cheatodipus · Cratogeomys · Cupidinimus · Dikkomys · Dipodomys · Diprionomys · Entoptychus · Geomys · Grangerimus · Gregorymys · Griphomys · Harrymys · Heliscomys · Heterogeomys · Heteromys · Lignimus · Liomys · Macrogeomys · Microdipodops · Mioheteromys · Mojavemys · Mookomys · Nerterogeomys · Oregonomys · Orthogeomys · Pappogeomys · Parapliosaccomys · Peridiomys · Perognathoides · Perognathus · Phelosaccomys · Pleurolicus · Pliogeomys · Pliosaccomys · Prodipodomys · Progeomys · Proheteromys · Reynoldsomys · Schizodontomys · Stratimus · Tenudomys · Thomomys · Ziamys · Zygogeomys
- Species: ZipcodeZoo has pages for 870 species and subspecies in the Family Geomyidae.
Genera
Balantiomys
Chaetodipus
Chaetodipus is a genus of pocket mouse containing 19 species endemic to the United States and Mexico. Like other members of their family such as pocket mice in the genus Perognathus, they are more closely related to pocket gophers than to true mice. [more]
Cheatodipus
Cratogeomys
A Genus in the Kingdom Animalia.[1] [more]
Cupidinimus
Dikkomys
Dipodomys
Kangaroo rats, genus Dipodomys, are small rodents native to North America. The common name derives from their bipedal form: as they hop in a manner similar to the much larger kangaroo, although they are not related. It has been noted that they are not properly characterized as "rats" at all. [more]
Diprionomys
Entoptychus
Geomys
The Geomys contains nine species of pocket gophers often collectively referred to as the eastern pocket gophers. Like all pocket gophers, members of this genus are fossorial herbivores. [more]
Grangerimus
Gregorymys
Griphomys
Harrymys
Heliscomys
Heterogeomys
A Genus in the Kingdom Animalia.[2] [more]
Heteromys
Heteromys is a genus of in the Heteromyidae family. It contains the following species: [more]
Lignimus
Liomys
Liomys is a genus of in the Heteromyidae family. It contains the following species: [more]
Macrogeomys
A Genus in the Kingdom Animalia.[3] [more]
Microdipodops
A kangaroo mouse is either one of the two species of jumping mouse (genus Microdipodops) native to the deserts of the Southwestern United States, predominantly found in the state of Nevada. Although native to the deserts in the southwest they have been plentiful in Missouri and Pennsylvania. The name "kangaroo mouse" refers to the species' extraordinary jumping ability, as well as its habit of bipedal locomotion. The two species are: [more]
Mioheteromys
Mojavemys
Mookomys
Nerterogeomys
Oregonomys
Orthogeomys
Orthogeomys is a genus of in the Geomyidae family, found in Mexico, Central America and Colombia. It contains the following species: [more]
Pappogeomys
Pappogeomys is a genus of in the Geomyidae family. It contains the following species: [more]
Parapliosaccomys
Peridiomys
Perognathoides
Perognathus
Perognathus is a genus of pocket mouse. Like other members of their family they are more closely related to pocket gophers than to true mice. [more]
Phelosaccomys
Pleurolicus
Pliogeomys
Pliosaccomys
Prodipodomys
Progeomys
Proheteromys
Reynoldsomys
Schizodontomys
Stratimus
Tenudomys
Thomomys
The smooth-toothed pocket gophers, genus Thomomys, are so called because they are among the only pocket gophers without grooves on their incisors. They are also called the Western pocket gophers because they are distributed in western North America. They are considered distinct enough from other pocket gophers to be recognized as a separate subfamily or tribe. [more]
Ziamys
Zygogeomys
The Michoacan Pocket Gopher (Zygogeomys trichopus) is a species of in the Geomyidae family. It is monotypic within the genus Zygogeomys. It is endemic to Mexico. Its natural habitat is temperate forests. [more]
More info about the Genus Zygogeomys may be found here.
References
- ^ Macdonald (Ed), Professor David W. (2006). The Encyclopedia of Mammals. Oxford University Press. ISBN 0-19-920608-2.
- ^ Patton, James (1984). Macdonald, D.. ed. The Encyclopedia of Mammals. New York: Facts on File. pp. 628?631. ISBN 0-87196-871-1.
- ^ Hazardous Substances and New Organisms Act 2003 - Schedule 2 Prohibited new organisms, New Zealand Government, http://legislation.govt.nz/act/public/1996/0030/latest/DLM386556.html#DLM386556, retrieved 26 January 2012
Footnotes
- http://www.ubio.org/browser/details.php?namebankID=112603
- http://www.ubio.org/browser/details.php?namebankID=112852
- http://www.ubio.org/browser/details.php?namebankID=112998
Sources
- The text on this page is licensed under the GNU Free Documentation License. It includes material from Wikipedia retrieved Wednesday, April 25, 2012.
- The distribution map on the Distribution tab comes from the Global Biodiversity Information Facility and is used with permission.
- Photographs on this page are copyrighted by individual photographers, and individual copyrights apply.
- The technology underlying this page, including the controls behind Keep Exploring, is owned by the BayScience Foundation. All rights are reserved.
