Overview
Cimolesta (from Greek, literally, "White Clay Thieves") is an extinct of mammals. A few experts place the pangolins within Cimolesta, though most other experts prefer to place the pangolins within their own order, Pholidota.
Some experts place the enigmatic family Ptolemaiidae within Cimolesta, also due to similarities between dental and skull anatomies with those of Pantolesta. If the Ptolemaiids were indeed cimolestids, then the range of Cimolesta would have ranged from the Late Cretaceous to the early Miocene, when the last ptolemaiid, Kelba, disappeared in Eastern Africa.
It is part of the cohort Ferae, and is possibly a sister taxon to the Perissodactyla. The Cimolesta contains several groups that are very different from each other, and are sometimes regarded as separate orders (which makes the Cimolesta a clade between that of order and superorder): the hooved Pantodonta, the insectivorous Didelphodonta (which contains thewell-known genus Cimolestes, a possible ancestor of the Carnivores) the possibly carnivorous Tillodonta, the Taeniodonta and the Apatotheria. Several groups are believed to have descended from the Cimolesta: the Pholidota (which could be regarded a suborder of Cimolesta), the Creodonta and the Carnivora. Cimolestans had a wide variety of body shapes, dentition and livestyles. The majority of them bore superficial resemblances to rodents, weasels or opossums.
Photos
Taxonomy
The Order Cimolesta is a member of the Mirorder Eparctocyona. Here is the complete "parentage" of Cimolesta:
- Domain: Eukaryota
Whittaker & Margulis,1978 - eukaryotes
- Kingdom: Animalia
Linnaeus, 1758 - animals
- Subkingdom: Bilateria
(Hatschek, 1888) Cavalier-Smith, 1983 - bilaterians
- Branch: Deuterostomia
Grobben, 1908 - Deuterostomes
- Infrakingdom: Chordonia
(Haeckel, 1874) Cavalier-Smith, 1998
- Phylum: Chordata
Bateson, 1885 - Chordates
- Subphylum: Vertebrata
Cuvier, 1812 - Vertebrates
- Infraphylum: Gnathostomata
Auct. - Jawed Vertebrates
- Superclass: Tetrapoda
Goodrich, 1930 - Tetrapods
- Class: Mammalia
C. Linnaeus, 1758 - Mammals
- Subclass: Theriiformes
(Rowe, 1988) M.c. Mckenna & S.k. Bell, 1997
- Infraclass: Holotheria
(Wible Et Al., 1995) M.c. Mckenna & S.k. Bell, 1997
- Superlegion: Trechnotheria
Mckenna, 1975
- Legion: Cladotheria
Mckenna, 1975
- Sublegion: Zatheria
Mckenna, 1975
- Infralegion: Tribosphenida
(Mckenna, 1975) M.c. Mckenna & S.k. Bell, 1997
- Supercohort: Theria
(Parker & Haswell, 1897) M.c. Mckenna & S.k. Bell, 1997 - Therians
- Cohort: Placentalia
(Owen, 1837) M.c. Mckenna & S.k. Bell, 1997 - Placentals
- Magnorder: Epitheria
(Mckenna, 1975) M.c. Mckenna & S.k. Bell, 1997
- Superorder: Preptotheria
(Mckenna, 1975) Mckenna, in Stucky & Mckenna, in Benton, Ed., 1993
- Grandorder: Ungulata
(un-goo-LAH-tuh)
(Linnaeus, 1766) Mckenna, 1975:41
- Mirorder: Eparctocyona
Mckenna, 1975
- Order: Cimolesta Mckenna, 1975
- Mirorder: Eparctocyona
Mckenna, 1975
- Grandorder: Ungulata
(un-goo-LAH-tuh)
(Linnaeus, 1766) Mckenna, 1975:41
- Superorder: Preptotheria
(Mckenna, 1975) Mckenna, in Stucky & Mckenna, in Benton, Ed., 1993
- Magnorder: Epitheria
(Mckenna, 1975) M.c. Mckenna & S.k. Bell, 1997
- Cohort: Placentalia
(Owen, 1837) M.c. Mckenna & S.k. Bell, 1997 - Placentals
- Supercohort: Theria
(Parker & Haswell, 1897) M.c. Mckenna & S.k. Bell, 1997 - Therians
- Infralegion: Tribosphenida
(Mckenna, 1975) M.c. Mckenna & S.k. Bell, 1997
- Sublegion: Zatheria
Mckenna, 1975
- Legion: Cladotheria
Mckenna, 1975
- Superlegion: Trechnotheria
Mckenna, 1975
- Infraclass: Holotheria
(Wible Et Al., 1995) M.c. Mckenna & S.k. Bell, 1997
- Subclass: Theriiformes
(Rowe, 1988) M.c. Mckenna & S.k. Bell, 1997
- Class: Mammalia
C. Linnaeus, 1758 - Mammals
- Superclass: Tetrapoda
Goodrich, 1930 - Tetrapods
- Infraphylum: Gnathostomata
Auct. - Jawed Vertebrates
- Subphylum: Vertebrata
Cuvier, 1812 - Vertebrates
- Phylum: Chordata
Bateson, 1885 - Chordates
- Infrakingdom: Chordonia
(Haeckel, 1874) Cavalier-Smith, 1998
- Branch: Deuterostomia
Grobben, 1908 - Deuterostomes
- Subkingdom: Bilateria
(Hatschek, 1888) Cavalier-Smith, 1983 - bilaterians
- Kingdom: Animalia
Linnaeus, 1758 - animals
The Order Cimolesta is further organized into finer groupings including:
- Suborder (8): Apatotheria · Didelphodonta · Ernanodonta · Pantodonta · Pantolesta · Pholidota · Taeniodonta · Tillodontia
- Family (22): Apatemyidae · Barylambdidae · Bemalambdidae · Cimolestidae · Coryphodontidae · Cyriacotheriidae · Epoicotheriidae · Ernanodontidae · Harpyodidae · Manidae · Metacheiromyidae · Palaeoryctidae · Pantolambdidae · Pantolambdodontidae · Pantolestidae · Paroxyclaenidae · Pastoralodontidae · Ptolemaiidae · Stylinodontidae · Tillotheriidae · Titanoideidae · Wangliidae
- Species: ZipcodeZoo has pages for 140 species and subspecies in the Order Cimolesta.
Families
Apatemyidae
Barylambdidae
Bemalambdidae
Cimolestidae
Coryphodontidae
Cyriacotheriidae
Epoicotheriidae
Ernanodontidae
Harpyodidae
Manidae
Pangolins or scaly anteaters are in the order Pholidota. There is only one extant family (Manidae) and one genus (Manis) of pangolins, comprising eight species. There are also a number of extinct taxa. Pangolins have large keratin scales covering their skin and are the only mammals with this adaptation. They are found in tropical regions of Africa and Asia. The name "pangolin" derives from the Malay word pengguling ("something that rolls up"). Pangolins are nocturnal animals, using their well-developed sense of smell to find insects. The long-tailed pangolin is also active by day. Pangolins spend most of the daytime sleeping, curled up into a ball. [more]
Metacheiromyidae
Palaeoryctidae
Pantolambdidae
Pantolambdodontidae
Pantolestidae
Paroxyclaenidae
Pastoralodontidae
Ptolemaiidae
Stylinodontidae
Tillotheriidae
Titanoideidae
Wangliidae
More info about the Family Wangliidae may be found here.
References
- [1] "Ptolemaiida, a New Order of Mammalian-with Description of the Cranium of Ptolemaia grangeri"
Sources
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