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Biblidinae is a of nymphalid butterflies that includes the tropical brushfoots. This subfamily was sometimes merged within the Limenitidinae, but they are now recognized as quite distinct lineages. In older literature, this subfamily is sometimes called Eurytelinae.

As of 2008, there are some 340 valid species in this subfamily, placed in 38^{[verification needed]} genera. Most species of Biblidinae are neotropical, but some Old World species and genera in Biblidini and Epicaliini.

Systematics

Diaethria clymena, one of the eighty-eights formerly in "Catagramma" (now Callicorini)

The Biblidinae are a taxonomically stable monophyletic group, at least since the "wastebin genus" "Catagramma" was dismantled around 1950. Six tribes are recognized, and the monophyly of them all is likewise well supported; radical changes in taxonomy and systematics are not expected in this subfamily^{[citation needed]}. The sister group to the Biblidinae is probably the Apaturinae and/or Nymphalinae.

The tribes, in the presumed phylogenetic sequence and with notable genera and some species also listed here, are:

Red Rim (Biblis hyperia: Biblidini)

Biblidini Boisduval, 1833

• Biblis Fabricius, 1807 (= Zonaga)
• Ariadne Horsfield, 1829 (= Ergolis)
  • Ariadne ariadne – Angled Castor
  • Ariadne merione – Common Castor
• Laringa Moore, 1901
• Eurytela Boisduval, 1833
• Neptidopsis Aurivillius, 1898
• Mesoxantha Aurivillius, 1898
• Byblia Hübner, 1819 (= Hypanis Boisduval, 1833 (non Pander in Menetries, 1832: preoccupied))
  • Byblia ilithyia – Joker
• Mestra Hübner, 1825 (= Cystineura)
• Archimestra Munroe, 1949
• Vila Kirby, 1871 (= Lonia, Neptis Hübner, 1819 (non Fabricius, 1807: preoccupied), Olina Doubleday, 1848 (non Robineau-Desvoidy, 1830: preoccupied))

Epicaliini Guenée, 1865

• Catonephele
• Nessaea
• Myscelia Doubleday, 1844 (= Sagaritis Hübner, 1821 (non Billberg, 1820: preoccupied), Sea)
• Sevenia Kocak, 1996 (= Crenis Boisduval, 1833 (non Hübner, 1821: preoccupied), Sallya Hemming, 1964 (non Yochelson, 1956: preoccupied))
• Eunica
• Cybdelis Boisduval, 1836

Panacea procilla specimen underside (Ageroniini)

Ageroniini Doubleday, 1847

• Hamadryas – cracker butterflies, calico butterflies
• Ectima Doubleday, 1848
• Panacea Godman & Salvin, 1883 (= Pandora Doubleday, 1848 (non Lamarck, 1799: preoccupied))
• Batesia Felder & Felder, 1862

Epiphilini Jenkins, 1987

• Asterope
• Pyrrhogyra Hübner, 1819 (= Corybas)
• Epiphile Doubleday, 1844
• Lucinia Hübner, 1823 (= Autodea)
• Bolboneura Godman & Salvin, 1877
• Temenis Hübner, 1819 (= Callicorina, Paromia Hewitson, 1861 (non Westwood, 1851: preoccupied))
• Nica Hübner, 1826 (= Pseudonica)
• Peria Kirby, 1871 (= Pelia Doubleday, 1849 (non Bell, 1836: preoccupied))

Dynamine coenus (Eubagini)

Eubagini Burmeister, 1878

• Dynamine Hübner, 1819 (= Arisba, Eubagis, Sironia)

Callicorini Orfila, 1952 – eighty-eights and relatives

• Diaethria
• Callicore
• Perisama Doubleday, 1849 (= Orophila)
• Antigonis Felder, 1861 (= Lincoya)
• Haematera Doubleday, 1849 (= Callidula)
• Paulogramma Dillon, 1948
• Catacore Dillon, 1948

The Subfamily Biblidinae is a member of the Family Nymphalidae. Here is the complete "parentage" of Biblidinae:

• Domain: Eukaryota Whittaker & Margulis,1978 - eukaryotes
  • Kingdom: Animalia Linnaeus, 1758 - animals
    • Subkingdom: Bilateria (Hatschek, 1888) Cavalier-Smith, 1983 - bilaterians
      • Branch: Protostomia Grobben, 1908 - protostomes
        • Infrakingdom: Ecdysozoa Aguinaldo Et Al., 1997 Ex Cavalier-Smith, 1998 - ecdysozoans
          • Superphylum: Panarthropoda Cuvier
            • Phylum: Arthropoda Latreille, 1829 - Arthropods
              • Subphylum: Mandibulata Snodgrass, 1938
                • Infraphylum: Atelocerata Heymons, 1901
                  • Superclass: Panhexapoda
                    • Class: Insecta C. Linnaeus, 1758 - Insects
                      • Subclass: Dicondylia
                        • Infraclass: Pterygota - Winged Insects
                          • Cohort: Myoglossata
                            • Superorder: Amphiesmenoptera
                              • Order: Lepidoptera Linnaeus, 1758 - Butterflies and Moths
                                • Infraorder: Heteroneura - a genus of Longhorned Beetles (Cerambycidae)
                                  • Family: Nymphalidae - Brushfooted Butterflies
                                    • Subfamily: Biblidinae - Tropical Brushfoots