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Apoidea

(Superfamily)

Overview

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The Apoidea is a major group within the Hymenoptera, which includes two traditionally-recognized lineages, the "sphecoid" wasps, and the bees, who appear to be their descendants.

Nomenclature

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Bees appear in recent classifications to be a specialized lineage of crabronid wasps that switched to the use of pollen and nectar as larval food, rather than insect prey; this presumably makes Crabronidae a paraphyletic group. Accordingly, bees and sphecoids are now all grouped together in a single superfamily, and the older available name is "Apoidea" rather than "Sphecoidea" (which, like Spheciformes, has been used in the past, but also defined a paraphyletic group and has been abandoned).

As the bees themselves (not including their wasp ancestors) are still considered a monophyletic group, it is still convenient to use a grouping between superfamily and family to unify all bees. A few recent classifications have addressed this problem by lumping all bee families together into a single large family Apidae[citation needed], though this has not met with widespread acceptance. The alternative classification in more common use is to unite all bees under the name Anthophila (Engel, 2005), which is equivalent to the obsolete name Apiformes (which meant bee-like forms in Latin).

Photos

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Taxonomy

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The Superfamily Apoidea is a member of the Series Aculeata. Here is the complete "parentage" of Apoidea:

The Superfamily Apoidea is further organized into finer groupings including:

Families

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Ampulicidae

The Ampulicidae, or Cockroach wasps, is a small (approx. 200 species), primarily tropical group of wasps, all of which use various cockroaches as prey items for their larvae. They tend to have elongated jaws, a pronounced neck-like constriction behind the head, a strongly petiolate abdomen, and deep grooves on the thorax. Many are quite ant-like in appearance, though some are brilliant metallic blue or green. [more]

Andrenidae

The family Andrenidae is a large (nearly) cosmopolitan (absent in Australia) non-parasitic family, with most of the diversity in temperate and/or arid areas (warm temperate xeric), including some truly enormous genera (e.g., Andrena with over 1300 species, and Perdita with nearly 800). One of the subfamilies, Oxaeinae, are so different in appearance that they were typically accorded family status, but careful phylogenetic analysis reveals them to be an offshoot within the Andrenidae, very close to the Andreninae. [more]

Angarosphecidae

[more]

Apidae

The Apidae are a large family of , comprising the common honey bees, stingless bees (which are also cultured for honey), carpenter bees, orchid bees, cuckoo bees, bumblebees, and various other less well-known groups. The family Apidae presently includes all the genera that were previously classified in the families Anthophoridae and Ctenoplectridae, and most of these are solitary species, though a few are also cleptoparasites. The four groups that were subfamilies in the old family Apidae are presently ranked as tribes within the subfamily Apinae. This trend has been taken to its extreme in a few recent classifications that place all the existing bee families together under the name "Apidae" (or, alternatively, the non-Linnaean clade "Anthophila"), but this is not a widely-accepted practice. [more]

Baissodidae

[more]

Colletidae

Colletidae is a of bees, and are often referred to collectively as plasterer bees, due to the method of smoothing the walls of their nest cells with secretions applied with their mouthparts; these secretions dry into a cellophane-like lining. There are 5 subfamilies, 54 genera, and over 2000 species, all of them evidently solitary, though many nest in aggregations. Two of the subfamilies, Euryglossinae and Hylaeinae, lack the external pollen-carrying apparatus (the scopa) that otherwise characterizes most bees, and instead carry the pollen in their crop. These groups, and in fact most genera in this family, have liquid or semi-liquid pollen masses on which the larvae develop. [more]

Crabronidae

Crabronidae is a large family of , that includes nearly all of the species formerly comprising the now-defunct superfamily Sphecoidea. It collectively includes well over 200 genera, containing well over 9000 species. Crabronids were originally a part of Sphecidae, but the latter name is now restricted to a separate family based on what was once the subfamily Sphecinae. As this change is very recent, it seems likely that the subfamilies of Crabronidae will each eventually be treated as families in their own right, as they have been treated as such by many authorities in the past (as in the catalog linked below). [more]

Halictidae

Halictidae is a family of the order Hymenoptera consisting of small (> 4 mm) to midsize (> 8 mm) bees which are usually dark-colored and often metallic in appearance. Several species are all or partly green and a few are red; a number of them have yellow markings, especially the males, which commonly possess yellow faces, a pattern widespread among the various families of bees. They are commonly referred to as sweat bees (especially the smaller species), as they are often attracted to perspiration; when pinched, females can give a minor sting. [more]

Heterogynaidae

[more]

Megachilidae

The Megachilidae are a of (mostly) solitary bees whose pollen-carrying structure (called a scopa) is restricted to the ventral surface of the abdomen (rather than mostly or exclusively on the hind legs as in other bee families). Megachilid genera are most commonly known as mason bees and leafcutter bees, reflecting the materials they build their nest cells from (soil or leaves, respectively); a few collect plant or animal hairs and fibers, and are called carder bees. All species feed on nectar and pollen, but a few are cleptoparasites (informally called "cuckoo bees"), feeding on pollen collected by other megachilid bees. Parasitic species do not possess a scopa. The brightly colored scopa leads to a colloquial name used occasionally in North America - "Jelly-belly bees." Megachilid bees are among the world's most efficient pollinators because of their energetic swimming-like motion in the reproductive structures of flowers, which moves pollen , as needed for pollination. One of the reasons they are efficient pollinators is their frequency of visits to plants, but this is because they are extremely inefficient at gathering pollen; compared to all other bee families, megachilids require on average nearly ten times as many trips to flowers to gather sufficient resources to provision a single brood cell. [more]

Melittidae

The family Melittidae is a small family, with some 60 species in 4 genera, restricted to Africa and the northern temperate zone. Historically, the family has included the Dasypodaidae and Meganomiidae as subfamilies, but recent molecular studies indicate that Melittidae (sensu lato) was paraphyletic, so each of the three historical subfamilies is now accorded family status, with Dasypodaidae as the basal group of bees, followed by Meganomiids and Melittids, which are sister taxa.. [more]

Paleomelittidae

[more]

Sphecidae

Sphecidae (, 1802) is a cosmopolitan family of wasps that include digger wasps, mud daubers and other familiar types that all fall under the category of thread-waisted wasps. Both of the traditional definitions of the Sphecidae (the conservative one, where all the sphecoid wasps other than ampulicids and heterogynaids were in a single large family, and the more refined one, where the 7 large sphecid subfamilies were each elevated to family rank) have recently been shown to be paraphyletic, and the most recent classification is closer to the conservative scheme; the families Heterogynaidae and Ampulicidae are the sister taxa to what are now two families (instead of one), the Sphecidae and Crabronidae. Thus, the bulk of the sphecoid wasps are now placed in Crabronidae, and Sphecidae per se is a much more restricted concept, equivalent to what used to be the subfamily Sphecinae. [more]

At least 721 species and subspecies belong to the Family Sphecidae.

More info about the Family Sphecidae may be found here.

References

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Sources

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Last Revised: September 22, 2009
2009/09/22 10:23:14