Overview
Brown bears are a symbol of America’s wildlands. In the time of the Lewis and Clark
expedition
they ranged over much of North America’s mid-plains westward to California and from central Mexico to Alaska and Canada’s Northwest Territory. Today only about 1,000 brown bears are found in the lower 48 states where they have lost 99 percent of their former range
.
Of four North American brown bear species, one, California’s golden bear ( Ursus arctos californicus has been extinct
since 1922 and the northern Mexico silvery grizzly
(U. a. nelsonii) is now believed to be extinct. The populations of Alaska’s Kodiak (U. a. middendorffi) and the common brown bear in Alaska and Canada (U. a. horribilis) appear to be stable. The lower 48 states U. a. horribilis population is listed as threatened except in areas of Montana and Idaho where there are experimental (non-essential) programs for reintroduction
. Small pockets
of brown bears of various subspecies
exist in parts of Europe, Asia, and the Middle
East. Most are either threatened or endangered
.
Only about one percent of the former ranges of brown bears in the lower 48 states ais left because of encroachment of human activities such as development, logging
, mining, and road construction on their habitats
, and illegal hunting to obtain their body parts for sale in Asian markets. In the states and Canadian provinces
in which sport hunting is permitted, it is tightly regulated. For example, Alaska Fish and Game issues only 325 hunting permits
a year out of 5,000 applications
and requires that all non residents must be accompanied by a native
guide
. The cost of a non-resident license
is $10,000-15,000.
Common Names
Click on the language to view common names.
Common Names in Chinese:
Ma-Xiong, Zong-Xiong
Common Names in Dutch:
Kodiakbeer
Common Names in English:
Alaskan Brown Bear, Brown Bear, Grizzly Bear, Mexican Grizzly Bear
Common Names in Russian:
Медведь бурый
Common Names in Spanish:
Oso Pardo
Description
Physical Description
Species Ursus arctos
Brown bears have a large heavily built body, a small head
, high eye profile
, a dish-shaped face
, short thick ears, and a small tail and legs
. They have high humped shoulders
consisting of muscles that give strength for digging. The hump
occurs less often in coastal bears. Interior bears lack the neck ruffles of coastal bears and some have a spectacled facial pattern
. Claws
are long, sharp, and non-retractable.
Color varies among the subspecies
. In general coastal bears are a creamy yellow to light brown with brownish legs and underparts; Kodiak, light brown, beige, or even blond; interior, dark brown to almost black. The brown bears in the Rocky Mountains, the grizzlies, get their name
from the silver-tipped guard hairs
on their back and shoulders that give them a frosted or silvery sheen appearance
. There are white bears (not albinos
) in portions of Alberta, Montana, and south central British Columbia.
The size of these bears is dependent
on subspecies, food items and their availability, and length of feeding season
. Because of their higher protein diets
, coastal Alaska and Kodiak Island bears are significantly larger than interior and mountain bears whose diet is primarily vegetation.
Full growth is attained when these bears are 10-11 years old. Interior males weigh 136-272 kg
(300-600 lb
) and females, 91-181 kg (200-400 lb); coastal males, 363-544 kg (800-1200 lb) and females, 272-363 kg (600-800 lb); Kodiak males, 544-580 kg (1200-1500 lb) and females an average of 317 kg (700 lb). When interior bears stand on their hind feet to get a better view
of their surroundings, they may be 1.8-2.7 m
(6-9 ft
) tall whereas Kodiak bears are 2.7-3.4 m (9-11 ft) in height
Habitat
North American brown bears prefer open areas interspersed with forests
for sheltering cover
while resting. They inhabit tundra
, alpine
meadows, coastal plains
near salmon runs, and rivers
and valleys. Grizzlies range
at lower elevations
part of the year, moving to higher areas above 1800 m
(6000 ft
) where there are deep soils for digging a hibernation den
and vegetative
cover for roof support
and isolation
of it. Siberian bears are primarily forest dwellers while European bears prefer mountain woodlands.
All brown bears require an ecosystem of habitats
that includes corridors for moving from one habitat to another. They establish home ranges
to meet this need. Male ranges are about 322-805 km
(200-500 mi
) and female, 80-483 km (50-300 mi). They are territorial
only during breeding season
.
Typically found at an altitude of 0 to 5,386 meters (0 to 17,671 feet).[1]
Biology
Diet
These bears are omnivores
and food is a top priority
in their lives. They generally forage
in the morning and evening, resting in dense cover
during the day. Their diet
changes with the seasons
as different food sources become available. They eat roots
, fungi, insects, and small mammals at all times of the year. They use their long sharp claws
and strong
foreleg muscles to tear logs
apart and turn over
stones
to get at the insects and their larvae and to dig out the burrows of mice, ground
squirrels, and marmots. In the spring
, just out of hibernation, they seek carrion
of animals that did not survive the winter and early spring vegetation—grasses, sedges, and moss. In the summer and early autumn they gorge
on food rich in protein and fat so as to build up their fat stores to sustain them over their hibernation. These include berries
, especially huckleberries, fruits, nuts, bulbs, and tubers. Their strong shoulder muscles are an asset for roto-tilling the soil to get at tubers.
They maximize their pre-hibernation weight
gain by selecting the most nutritious portions of the food item to eat and the time of year when an item is most nutritious such as berries when they are ripe
and have the highest sugar
levels; brains of samll mammals, fish, and eggs
of salmon or trout; only the eyes of salmon after they are satiated; internal organs of deer, elk, and cattle; and “spring-greenup”, the spring and early summer grasses. xx Diet choices differ geographically. In Yellowstone National Park grizzlies fish for spawning trout and Alaskan and Canadian coastal brown bears fish for spawning salmon. Bears in Glacier National Park follow wolf packs
and mountain lions to remote
regions using them as providers of meals
of deer and elk. While 60-90% of the diet of coastal and interior bear is vegetation, that of the carnivorous
Kodiaks is much lower because of the inclusion of moose, elk, mountain sheep, and caribou in their diet.
A major food source for grizzlies in the Yellowstone area is army
cutworm moths. In the summer these insects appear in huge numbers in the mountains at elevations
above 3048 m
(10,000 ft
) where they feast on plant nectar in the early morning and then rest on rocks in swarms
. This food supply causes congregations of grizzlies from distance
and widely spread
out home ranges
to climb to the higher elevations where they each consume 10,000-20,000 moths a day. The moths are important sources of protein and fat for the bears’ weigh gain prior to hibernation.
Reproduction
Male and female brown bears become sexually mature
at 4-6 years of age but first pregnancies usually do not occur until the females are 5.5-7 years old. Males (boars) become territorial
during the mating season
although their defense of the female against other males is usually not successful with the result that both males and females mate with others during the about three week breeding season
. Consequently, the cubs in a litter
may have had different fathers.
Once the egg
is fertilized it divides several times. The embryo then remains suspended without implanting in the uterus
(delayed implantation) until the female goes into hibernation in October or November. If the sow
(female) has not eaten enough to sustain herself over the hibernation, the egg will not implant. After a 180-266 day gestation
, one to four (average two) helpless cubs are born that are toothless, almost hairless, and unable to see or hear. They weigh about 241-368 g (8.5-3 oz
) at birth.
When the family
emerges
from the den
in March to May, the sow may have lost as much as 40 percent of her weight
. In contrast her cubs now weigh 6.8-9.1 kg
(15-20 lb
). Cubs stop nursing during July to September of their first year. In the two years that the cubs are usually with the female she teaches them which foods to eat, where to find them, how to fish, and how to defend themselves. Aggressive in their defense, she may chase the cubs up a tree
as a protective measure. She also swats them if they misbehave. If she becomes pregnant again in the cubs’ second year, she drives them away and enters hibernation alone. The sibling cubs usually hibernate together. There is a high predation
of cubs by males seeking mates.
Behavior
Brown bears communicate by smell and voice. They make moaning sounds
when foraging
and spread
scent by scratching and rubbing on trees
and other landmarks to communicate reproduction
status and territorial
boundaries during breeding season
.
The exact time of brown bear hibernation is dependent
on location, weather, and condition of the individual bear. They place a bed
of dry vegetation inside a den
that they have dug on a sheltered slope
, under a large stone
, or among the roots
of a mature
tree. Males and females have separate dens and females enter hibernation first emerging last. These bears are not true hibernators. Their respiration, heartbeat, and body temperature
do not drop as low as true hibernators. They also can be awakened and may waken themselves to leave the den in search of food.
Adaptations: The size and strength of these bears puts them at the top of a food chain where they have little to fear from predators—all except humans. The long claws
, strong
leg
and shoulder
muscles of these bears are well adapted for digging dens and food. The short clawed cubs can climb but the long claws of the adults
prevent them from climbing
as black bears do.
Longevity
: As many as 50 percent of brown bears do not reach breeding age. Those that do may survive for 25 years.
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Animalia
(
)
- Linnaeus, 1758
- animals
- Subkingdom:
Bilateria
(
)
- (Hatschek, 1888) Cavalier-Smith, 1983
- Branch:
Deuterostomia
(
)
- Grobben, 1908
- Infrakingdom:
Chordonia
(
)
- (Haeckel, 1874) Cavalier-Smith, 1998
- Phylum:
Chordata
(
)
- Bateson, 1885
- Chordates
- Subphylum:
Vertebrata
(
)
- Cuvier, 1812
- Vertebrates
- Infraphylum:
Gnathostomata
(
)
- Auct.
- Jawed Vertebrates
- Superclass:
Tetrapoda
(
)
- Goodrich, 1930
- Class:
Mammalia
(
)
- C. Linnaeus, 1758
- Mammals
- Subclass:
Theriiformes
(
)
- (Rowe, 1988) M.c. Mckenna & S.k. Bell, 1997
- Infraclass:
Holotheria
(
)
- (Wible Et Al., 1995) M.c. Mckenna & S.k. Bell, 1997
- Superlegion:
Trechnotheria
(
)
- Mckenna, 1975
- Legion:
Cladotheria
(
)
- Mckenna, 1975
- Sublegion:
Zatheria
(
)
- Mckenna, 1975
- Infralegion:
Tribosphenida
(
)
- (Mckenna, 1975) M.c. Mckenna & S.k. Bell, 1997
- Supercohort:
Theria
(
)
- (Parker & Haswell, 1897) M.c. Mckenna & S.k. Bell, 1997
- Cohort:
Placentalia
(
)
- (Owen, 1837) M.c. Mckenna & S.k. Bell, 1997
- Magnorder:
Epitheria
(
)
- (Mckenna, 1975) M.c. Mckenna & S.k. Bell, 1997
- Superorder:
Preptotheria
(
)
- (Mckenna, 1975) Mckenna, in Stucky & Mckenna, in Benton, Ed., 1993
- Superorder:
Preptotheria
(
- Magnorder:
Epitheria
(
- Cohort:
Placentalia
(
- Supercohort:
Theria
(
- Infralegion:
Tribosphenida
(
- Sublegion:
Zatheria
(
- Legion:
Cladotheria
(
- Superlegion:
Trechnotheria
(
- Infraclass:
Holotheria
(
- Subclass:
Theriiformes
(
- Class:
Mammalia
(
- Superclass:
Tetrapoda
(
- Infraphylum:
Gnathostomata
(
- Subphylum:
Vertebrata
(
- Phylum:
Chordata
(
- Infrakingdom:
Chordonia
(
- Branch:
Deuterostomia
(
- Subkingdom:
Bilateria
(
- Kingdom:
Animalia
(
Notes
Name
Status: Accepted Name
. Latest taxonomic
scrutiny: October 28, 2003.
Other common names
: Spanish: Oso pardo
Similar Species
Recent DNA studies support the theory of some scientists that polar bears are not a separate species of the genus Ursus (Ursus maritimus) but a subspecies, U. arctos maritimus . The theory is reinforced by the fact that the offspring of brown/polar bear matings are fertile
Members of the genus Ursus
There are approximately 84 species in this genus:
U. abstrusus · U. americana · U. americanus (American Black Bear) · U. americanus altifrontalis · U. americanus amblyceps · U. americanus americanus (American Black Bear) · U. americanus californiensis · U. americanus carlottae · U. americanus cinnamomum · U. americanus emmonsi · U. americanus emmonsii (Glacier Bear) · U. americanus eremicus · U. americanus floridanus (Florida Black Bear) · U. americanus hamiltoni · U. americanus kermodei (Kermode Black Bear) · U. americanus luteolus (Louisiana Black Bear) · U. americanus machetes · U. americanus perniger (Kenai Black Bear) · U. americanus pugnax · U. americanus subsp · U. americanus vancouveri · U. arctos (Mexican Grizzly Bear) · U. arctos alascensis · U. arctos arctos (Mexican Grizzly Bear) · U. arctos beringianus · U. arctos californicus (Bajan Bear) · U. arctos caucasicus (Mexican Grizzly Bear) · U. arctos crowtheri (Atlas Bear) · U. arctos dalli · U. arctos gyas · U. arctos horribilis (Grizzly Bear) · U. arctos horribilus · U. arctos isabellinus · U. arctos lasiotus · U. arctos meridionalis · U. arctos middendorff · U. arctos middendorffi (Mexican Grizzly Bear) · U. arctos middendorfi · U. arctos middendorfii · U. arctos nelsoni · U. arctos pruinosus (Mexican Grizzly Bear) · U. arctos richardsoni · U. arctos shanorum · U. arctos sitkensis · U. arctos stikeenensis · U. arctos subsp · U. arctos syriacus (Sirian Brown Bear) · U. arctos yesoensis · U. arctus · U. arvernensis · U. boeckhi · U. cancrivorus · U. deningeri · U. etruscus · U. horribilis · U. lasiotus · U. luteolus · U. malayanus · U. maritimus (Polar Bear) · U. maritimus marinus (Polar Bear) · U. maritimus maritimus (Polar Bear) · U. mediterraneus · U. melanoleucus · U. middendorffi (Alaskan Brown Bear) · U. minimus · U. nasutus · U. ornatus · U. ruscinensis · U. sackdillingensis · U. spaeleus · U. spelaearctos · U. spelaeus · U. taxus · U. thibetanus (Himalayan Black Bear) · U. thibetanus gedrosianus (Baluchistan Bear) · U. thibetanus japonicus · U. thibetanus laniger · U. thibetanus thibetanus (Himalayan Black Bear) · U. thibetanus ussuricus · U. torquatus arboreus · U. ursinus · U. ursinus ursinus · U. wenzensis · U. x
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Further Reading
- A catalogue of the collection of mammals in the Field Columbian Museum. Chicago: Field Columbian Museum, 1907. ENG url p. 419, p. 660.
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- A checklist of the vertebrate animals of Kansas / George D. Potts, Joseph T. Collins. [Lawrence, Kan.]: Museum of Natural History, University of Kansas, 1991. ENG url p. 4.
- A guide to the quadrupeds and reptiles of Europe: with descriptions of all the species / compiled from the latest writers by Lord Clermont. London: J. Van Voorst, 1859. ENG url p. 49.
- A study of the mammals of Iran: resulting from the Street Expedition of 1962-63 [by] Douglas M. Lay. [Chicago]Field Museum of Natural History, 1967. ENG url p. 100, p. 207, p. 282, p. 75, p. 93, p. 97.
- A survey of the mammals of Afghanistan, resulting from the 1965 Street Expedition (excluding bats) [by] Jerry D. Hassinger. [Chicago]Field Museum of Natural History, 1973. ENG url p. 128, p. 130, p. x.
- A text-book of zoogeography, by Frank E. Beddard... Cambridge, University press, 1895. ENG url p. 154, p. 155, p. 90.
- Abhandlungen der Naturhistorischen Gesellschaft zu Nrnberg. Nrnberg: Conrad Geiger, 1852- GER url p. 137, p. 71.
- Allgemeine Zoologie und Abstammungslehre / von Ludwig Plate. Jena: Fischer, 1922-1924. GER url p. 297, p. 300.
- An Arctic ecosystem: the coastal tundra at Barrow, Alaska / edited by Jerry Brown [et al.]. Stroudsburg, Pa.: Dowden, Hutchinson & Ross: c1980. ENG url p. 9.
- An introduction to the study of mammals living and extinct, by William Henry Flower and Richard Lydekker London, A. and C. Black, 1891. ENG url p. 558.
- Anatomische Hefte, München [etc.]J.F. Bergmann. GER url p. 187, p. 215, p. 679, p. 733, p. 774, p. 782, p. 829, p. 830.
- Anglo-russkii biologicheskii slovar' / [avtory, I. N. Afanas'eva et al.; spetsial'nye nauch. redaktory, O. I. Chibisova i L. A. Koziar]. Moskva: Russkii iazyk, 1979. ENG url p. 65.
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Notes
Contributors
- Bear Specialist Group 1996. In IUCN 2008. 2008 IUCN Red List of Threatened Species. IUCNRedList.org. Downloaded July 20, 2008.
- Global Biodiversity Information Facility. Accessed November 12, 2007. http://www.gbif.org Mediated distribution data from 19 providers.
Data Sources
Accessed through GBIF Data Portal November 12, 2007:
- European Environment Agency, EUNIS
- Marine Science Institute, UCSB, Paleobiology Database
- Michigan State University Museum, Vertebrate specimens
- Museum of Vertebrate Zoology, Terrestrial vertebrate specimens
- Royal Ontario Museum, Mammal specimens
- Sternberg Museum of Natural History, Mammal Collection
- The Swedish Museum of Natural History
- , Mammals
- University of Alaska Museum of the North, University of Alaska Museum Mammal Collection
- University of Alaska Museum of the North, University of New Mexico Museum of Southwestern Biology Mammal Collection
Identifiers
- Biodiversity Heritage Library NamebankID: 2478082
- Catalogue of Life Accepted Name Code: ITS-202386
- Global Biodiversity Information Facility Taxonkey: 13808342
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 180543
- IUCN ID: 41688
- U.S. Fish and Wildlife Service Species Identifier: A02V
- Zipcode Zoo Species Identifier: 180032
Footnotes
- Mean = 588.090 meters (1,929.429 feet), Standard Deviation = 664.290 based on 585 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
