Overview
Family : Cichlids ; Most common in blind estuaries and coastal lakes [1]. Also found in warm weedy pools of sluggish streams , canals, and ponds [2]. Mainly diurnal . May form schools. Omnivorous , feeds on almost anything from algae to insects. Can be reared under hyper-saline conditions[3]. Marketed fresh and frozen[4]. Juveniles are carnivorous , adults tend to be herbivorous[5]. Exhibits considerable plasticity in its feeding habits as well as in its reproductive biology [6].
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Near Threatened |
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Interesting Facts
Common Names
Click on the language to view common names.
Common Names in Adangme:
Logokpa
Common Names in Afrikaans:
Blou kurper, Blou-Kurper, Bloukurper, Grootbekkurper, Mosambiek-tilapia, Mosambiekkurper, Mossambiekkurper, Rooivinkurper
Common Names in Amharic:
Koroso
Common Names in Arabic:
Bulti
Common Names in Bahasa Indonesia:
Mujair, nila, Nila merah
Common Names in Batak:
Jahir, nila
Common Names in Cantonese:
Fai chau chak ue, Gam san tsak
Common Names in Chinese:
Fai chau chak ue, Gam san tsak, Wu-Kuo yu
Common Names in Creole, French:
Lapia, Petit lapia, Tilapia, Tilapya
Common Names in Creoles and Pidgins, French:
Lapia
Common Names in Czech:
Tlamoun nilský
Common Names in Danish:
Mozambique-cichlide, Mozambique-mundruger, Nil-mundruger, Nilcichlide
Common Names in Digo:
Para para
Common Names in El Molo:
Sigir orok
Common Names in English:
African mouthbrooder, African perch, Baringo tilapia, Black tilapia, Blue bream, blue tilapia, Cichlid, Common tilapia, Edward tilapia, Hawaiian Perch, Hawaiian Sunfish, Japanese fish, Java fish, Java tilapia, Kurper bream, Largemouth kurper, Largemouth tilapia, Mango fish, Mocambique Tilapia, Mojarra, Mossambique Tilapia, Mouth-Brooder, Mozambique cichlid, Mozambique mouth-breeder, Mozambique mouth-brooder, Mozambique mouthbrooder, Mozambique tilapia, Nile tilapia, Nilotica, Redfin tilapia, Suguta tilapia, Tilapia, Turkana tilapia
Common Names in Ewe:
Akpafiatsi, Gbolonu, Logokpa
Common Names in Fang:
Ekouni
Common Names in Fijian:
Malea, Maleya, Tilapia
Common Names in Finnish:
Mustatilapia, Niilintilapia, Panganintilapia, Valkoleukatilapia
Common Names in French:
carpe, Tilapia, Tilapia du Mozambique, Tilapia du Nil
Common Names in Gã:
Didee
Common Names in German:
Baringo-Nilbuntbarsch, Lake Eduard-Nilbuntbarsch, Mosambik-Maulbr, Mosambik-Maulbrüter, Mossambik-Buntbarsch, Natalbarsch, Nil-Maulbrüter, Nilbuntbarsch, Tilapie, Wei, Weißkehlbarsch
Common Names in Hausa:
Bugu, Falga, Garagaza, Gargaza, Karfasa
Common Names in Hebrew:
Amnun yeor
Common Names in Igbo:
Ifunu, Mpupa
Common Names in Ijo:
Tome, Ukuobu
Common Names in Japanese:
Chikadai, Kawasuzume, カワスズメ(テイラピア)
Common Names in Javanese:
Mujair
Common Names in Jula:
T?b?nfin, Tɛbɛnfin
Common Names in Kannada:
Jilebi
Common Names in Kanuri:
Karwa
Common Names in Khmer:
Trey tilapia chhnoht, Trey tilapia khmao
Common Names in Kim:
Biering-pill, Peng-pill, Sale
Common Names in Lao:
Nin
Common Names in Laotian:
Nin
Common Names in Luo:
Ngege, Nyamami
Common Names in Mahl:
Mandapa
Common Names in Malay:
Mujair, nila, Tilapia, Tilapia Laut
Common Names in Malayalam:
Kolathile-meen, Thilapia, കൊലതിലി മീന് , തിലപിയ് , തിലാപി, തിലോപ്പി
Common Names in Mandarin Chinese:
Wu-Kuo yu, 南洋鯽仔, 南洋鲫仔, 南非非鯽, 南非非鲫, 吉里口孵非鯽, 吉里口孵非鲫, 在來吳郭魚, 在来吴郭鱼, 大鱗口孵非鯽, 大鳞口孵非鲫, 尼罗口孵非鲫(引进), 尼罗口孵鱼, 尼罗非鲫, 尼羅口孵非鯽(引進), 尼羅口孵魚, 尼羅非鯽, 巴林口孵非鯽, 巴林口孵非鲫, 柔口孵非鯽, 柔口孵非鲫, 格紋口孵非鯽, 格纹口孵非鲫, 沃氏口孵非鯽, 沃氏口孵非鲫, 潘氏口孵非鯽, 潘氏口孵非鲫, 莫三鼻口孵魚, 莫三鼻口孵鱼, 莫桑比克口孵非鯽, 莫桑比克口孵非鲫, 莫桑比克帚齒非鯽, 莫桑比克帚齿非鲫, 莫桑比克非鯽, 莫桑比克非鲫, 菲洛口孵非鯽, 菲洛口孵非鲫, 萨格口孵非鲫, 薩格口孵非鯽, 非洲仔, 魯沃口孵非鯽, 鲁沃口孵非鲫
Common Names in Maranao/Samal/Tao Sug:
Tilapiya
Common Names in Mooré:
Tegr-pere
Common Names in Nupe:
Tsokungi
Common Names in Nyanja:
Mphende
Common Names in Oromo:
Qoroosoo
Common Names in Other:
Ihere, Mahere, Matole, Ngege, Sibore, Sopore
Common Names in Polish:
Tilapia nilowa
Common Names in Portuguese:
Til, Tilápia-de-Moçambique, Tilápia-do-Nilo
Common Names in Russian:
Mozambikskaya tilapiya, мозамбикская тиляпия
Common Names in Sena:
Nkobue
Common Names in Sinhalese:
Tilapia
Common Names in Spanish:
Mojarra, Tilapia, Tilapia De Mozambique, Tilapia del Mozambique, Tilapia del Nilo, tilapia mosambica, Tilapia Moz, Tilapia mozámbica, Tilapia nilótica
Common Names in Sranan:
Tilapia
Common Names in Swahili:
Ngege, Nyamami, Perege, Sato, Uganda
Common Names in Swedish:
Mossambique-tilapia, Munruvare, Niltilapia
Common Names in Tagalog:
Pla pla, Tilapia
Common Names in Tamil:
Jelebi meen, Tilapia, ஜிலேபி மீன், திலேப்பி
Common Names in Thai:
Pla nil, ปลานิล
Common Names in Toba, Batak:
Jahir, nila
Common Names in Turkana:
Kokine, Rogene
Common Names in Venda:
Bende, Khala, Thipende, Tshena, Tshikwea
Common Names in Vietnamese:
Cá Rô Phi den, Cá Rô phi v?n, Cá Rô phi vằn, Cá Rô Phi đen
Common Names in Wolof:
Wass
Common Names in Yoruba:
Epia
Common Names in Zande:
Kpakaru
Description
Physical Description
Species Oreochromis mossambicus
28-31 vertebrae ; dorsal spines XV-XVII; total dorsal rays 26-29; 30-32 lateral line scales ; anal spines III, lower outer gill rakers 14-20; fine pharyngeal teeth; breeding males black (not in some cultured strains) with white lower parts on head ; red dorsal and caudal fin margins ; remnants of striped and barred pattern often visible in females, juveniles and non-breeding makes, as a series of mid-lateral and dorsal blotches; jaws of adult males greatly enlarged, concave dorsal head profile ; male genital papilla simple or slightly notched ; caudal fin not densely scaled .The so-called red tilapia in aquaculture is a hybrid between O. mossambicus and either O. niloticus or O. hornorum. Oreochromis mossambicus is the research subject of many physiological and biochemical studies in Asia.
Size/Age/Growth
May live as long as 11 years in the wild, 5 years in captivity.
Habitat
Many tilapias can live quite happily in seawater. The fact that they have not typically invaded coral reefs is perhaps due to predation by marine fishes . (Courtenay, W., pers. comm. , 2004). Oreochromis mossambicus is very hardy and tolerates the high salinities of atoll lagoons , such as that at Fanning Atoll (Lobel 1980). Thought to be ideal pond fish , they readily produce stunted stocks when overcrowded, as has been observed on Pagan in the Northern Mariana Islands (Source: SPREP).
Typically found in a lake at a mean distance from sea level of 529 meters (1,735 feet).[7]
Ecology:
Occurs in all but fast-flowing waters; thrives in standing waters
.
Further south in its range
it is most common in blind estuaries and
coastal lakes
where it tolerates brackish
and marine
environments.
Feeds
on algae, especially diatoms
, and detritus
, large individuals
also take insects and other invertebrates
. Breeds
in summer, females
raising multiple
broods every 3 to 4 weeks during a season
. Males
construct a saucer-shaped
nest
on sandy bottoms
: the female mouthbroods
the eggs
, larvae and small fry
. Juveniles
shoal in shallow water.
Prone to stunting under adverse
or crowded conditions (Skelton 2001).
[8].
List of Habitats
:
- 5 Wetlands (inland)
- 5.1 Wetlands (inland) - Permanent Rivers/Streams/Creeks (includes waterfalls )
- 5.2 Wetlands (inland) - Seasonal/Intermittent/Irregular Rivers/Streams/Creeks
- 5.4 Wetlands (inland) - Bogs , Marshes, Swamps , Fens , Peatlands
- 5.5 Wetlands (inland) - Permanent Freshwater Lakes (over 8ha)
- 5.6 Wetlands (inland) - Seasonal/Intermittent Freshwater Lakes (over 8ha)
- 5.7 Wetlands (inland) - Permanent Freshwater Marshes/Pools (under 8ha)
- 5.8 Wetlands (inland) - Seasonal/Intermittent Freshwater Marshes/Pools (under 8ha)
- 5.13 Wetlands (inland) - Permanent Inland Deltas
- 5.14 Wetlands (inland) - Permanent Saline, Brackish or Alkaline Lakes
- 5.15 Wetlands (inland) - Seasonal/Intermittent Saline, Brackish or Alkaline Lakes and Flats
- 5.16 Wetlands (inland) - Permanent Saline, Brackish or Alkaline Marshes/Pools
- 5.17 Wetlands (inland) - Seasonal/Intermittent Saline, Brackish or Alkaline Marshes/Pools
- 9 Marine Neritic
- 9.10 Marine Neritic - Estuaries
- 12 Marine Intertidal
- 12.5 Marine Intertidal - Salt Marshes (Emergent Grasses)
- 13 Marine Coastal/Supratidal
- 13.4 Marine Coastal/Supratidal - Coastal Brackish/Saline Lagoons/Marine Lakes
- 13.5 Marine Coastal/Supratidal - Coastal Freshwater Lakes
- 15 Artificial/Aquatic & Marine
- 15.1 Artificial/Aquatic - Water Storage Areas (over 8ha)
- 15.2 Artificial/Aquatic - Ponds (below 8ha)
- 15.3 Artificial/Aquatic - Aquaculture Ponds
- 15.7 Artificial/Aquatic - Irrigated Land (includes irrigation channels )
- 15.8 Artificial/Aquatic - Seasonally Flooded Agricultural Land
- 15.9 Artificial/Aquatic - Canals and Drainage Channels, Ditches [more info]
Biology
Reproduction
Egg-layer.Male builds spawning pits. Up to 1775 ripe eggs
in one
female. Hatching
after 3-5 days; fry released 10-14 days after spawning,
but mouthbrooded for about another week; more than one brood per
season
.
Size and age of sexual maturity varies according to environmental
conditions, with spawning in ponds
at 2-3 months and 6-10 cm for
females and 7-13 cm for males at intervals of 1- 5 months. In natural
conditions sexual maturity at greater age and size.
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Animalia
(
)
- C. Linnaeus, 1758
- animals
- Subkingdom:
Bilateria
(
)
- (Hatschek, 1888) Cavalier-Smith, 1983
- Branch:
Deuterostomia
(
)
- Grobben, 1908
- Infrakingdom:
Chordonia
(
)
- (Haeckel, 1874) Cavalier-Smith, 1998
- Phylum:
Chordata
(
)
- Bateson, 1885
- Chordates
- Subphylum:
Vertebrata
(
)
- Cuvier, 1812
- Vertebrates
- Infraphylum:
Gnathostomata
(
)
- auct.
- Jawed Vertebrates
- Superclass:
Osteichthyes
(
)
- Huxley, 1880
- Class:
Osteichthyes
(
)
- Huxley, 1880
- Subclass:
Actinopterygii
(
)
- Infraclass:
Actinopteri
(
)
- Cohort:
Clupeocephala
(
)
- Superorder:
Acanthopterygii
(
)
- Order:
Perciformes
(
)
-
- Suborder:
Labroidei
(
)
- Family:
Cichlidae
(
)
- Genus:
Oreochromis
(
)
- Günther, 1889
- Specific name:
mossambicus
- Scientific name: - Oreochromis mossambicus (Peters, 1852)
- Specific name:
mossambicus
- Genus:
Oreochromis
(
- Family:
Cichlidae
(
- Suborder:
Labroidei
(
- Order:
Perciformes
(
- Superorder:
Acanthopterygii
(
- Cohort:
Clupeocephala
(
- Infraclass:
Actinopteri
(
- Subclass:
Actinopterygii
(
- Class:
Osteichthyes
(
- Superclass:
Osteichthyes
(
- Infraphylum:
Gnathostomata
(
- Subphylum:
Vertebrata
(
- Phylum:
Chordata
(
- Infrakingdom:
Chordonia
(
- Branch:
Deuterostomia
(
- Subkingdom:
Bilateria
(
- Kingdom:
Animalia
(
Synonyms
Chromis dumerilii • Chromis dumerilii Steindachner • Chromis mossambicus Peters • Chromis mossambicus Peters, 1852 • Chromis natalensis • Chromis natalensis Weber • Chromis niloticus (Non Linnaeus) • Chromis niloticus mossambicus Peters • Chromis vorax • Chromis vorax Pfeffer • Cromis mossambicus Peters • Oreochromis mossambica • Oreochromis mossambica (Peters • Oreochromis mossambica (Peters, 1852) • Oreochromis mossambicus • Oreochromis mozambica • Oreochromis mozambica (Peters • Oreochromis mozambica (Peters, 1852) • Oreochromis niloticus baringoensis • Oreochromis niloticus cancellatus • Oreochromis niloticus eduardianus • Oreochromis niloticus filoa • Oreochromis niloticus filoa (Trewavas 1983) • Oreochromis niloticus niloticus • Oreochromis niloticus baringoensis Trewavas, 1983 • Oreochromis niloticus cancellatus (Nichols, 1923) • Oreochromis niloticus eduardianus (Boulenger, 1912) • Oreochromis niloticus filoa Trewavas, 1983 • Oreochromis niloticus niloticus (Linnaeus, 1758) • Oreochromis niloticus sugutae Trewavas, 1983 • Oreochromis niloticus vulcani (Trewavas, 1933) • Oreochromis niloticus sugutae • Oreochromis niloticus vulcani • Oreochromis pangani girigan • Oreochromis pangani pangani • Oreochromis pangani girigan (Lowe, 1955) • Oreochromis pangani pangani (Lowe, 1955) • Oreochromis placidus placidus • Oreochromis placidus ruvumae • Oreochromis placidus placidus (Trewavas, 1941) • Oreochromis placidus ruvumae (Trewavas, 1966) • Perca nilotica • Perca nilotica Linnaeus, 1758 • Sarotherodon mossambicus (Peters • Tilapia arnoldi • Tilapia arnoldi Gilchrist & Thompson • Tilapia cancellata • Tilapia cancellata Nichols, 1923 • Tilapia dumerili (Steindachner • Tilapia dumerilii (Steindachner • Tilapia eduardiana Boulenger, 1912 • Tilapia girigan • Tilapia girigan Lowe, 1955 • Tilapia kafuensis (Non Boulenger • Tilapia mossambica (Peters • Tilapia mossambica mossambica (Peters • Tilapia mossambicus (Peters • Tilapia natalensis (Weber • Tilapia pangani • Tilapia pangani Lowe, 1955 • Tilapia placida Trewavas, 1941 • Tilapia ruvumae • Tilapia ruvumae Trewavas, 1966 • Tilapia vorax (Pfeffer • Tilapia vulcani • Tilapia vulcani Trewavas, 1933
Notes
Name
Status: Accepted Name
.
Last scrutiny: Data
last modified by FishBase 15-Dec-1995
Similar Species
Members of the genus Oreochromis
ZipcodeZoo has pages for 41 species and subspecies in this genus:
O. alcalicus (Lake Magadi Tilapia) · O. alcalicus alcalicus (Lake Natron Tilapia) · O. andersonii (Threespot Bream) · O. andersonii (Three-Spotted Tilapia) · O. aureus (Jordan St. Peter´s Fish) · O. esculentus (Singidia Tilapia) · O. hunteri (Lake Chala Tilapia) · O. jipe (Jipe Tilapia) · O. karomo (Karomo) · O. karongae (Tilapia) · O. korogwe (Korogwe Tilapia) · O. leucostictus (Blue Spotted Tilapia) · O. lidole (Green-Head Tilapia) · O. macrochir (Mozambique Mouth-Brooder) · O. macrochir macrochir (Green-Headed Bream) · O. mortimeri (Kariba Tilapia) · O. mossambicus (Mozambique Mouth-Brooder) · O. mweruensis (Mweru Tilapia) · O. niloticus (Tilapia) · O. niloticus baringoensis (Baringo Tilapia) · O. niloticus eduardianus (Nile Tilapia) · O. niloticus niloticus (Baringo Tilapia) · O. niloticus sugutae (Suguta Tilapia) · O. niloticus vulcani (Turkana Tilapia) · O. placidus (Black Tilapia) · O. placidus placidus (Rovuma Tilapia) · O. placidus rovumae (Black Tilapia) · O. rukwaensis (Lake Rukwa Tilapia) · O. saka (Tilapia) · O. salinicola (Buffalo Springs Tilapia) · O. shiranus (Ulenger) · O. shiranus chilwae (Chilwa Tilapia) · O. spilurus (Oulenger) · O. spilurus niger (Athi River Tilapia) · O. spilurus percivali (Buffalo Springs Tilapia) · O. spilurus spilurus (Sabaki Tilapia) · O. upembae (Rufigi Tilapia) · O. urolepis (Rufigi Tilapia) · O. urolepis hornorum (Wami Tilapia) · O. urolepis urolepis (Rufigi Tilapia) · O. variabilis (Victoria Tilapia)
More Info
- Search for Pictures: images.google.com
- Search for Scholarly Articles: Google Scholar
- Search using Scientific Name and Vernacular Names: All the Web | AltaVista Canada | AltaVista | Excite | Google | HotBot | Lycos
- Search using Specialized Databases: GenBank | Medline | Scirus | CISTI/CAL | Agricola Periodicals | Agricola Books
Further Reading
- A floral and faunal inventory of the Parc national de Marojejy, Madagascar: with reference to elevational variation / Steven M. Goodman, editor. 97 2000 [Chicago, Ill.]: Field Museum of Natural History, c2000. url p. 150, p. 284.
- Annual report - Western Society of Malacologists. Pomona, Calif., Western Society of Malacologists. url p. 34.
- Bulletin of the British Museum (Natural History). London: BM(NH) url p. 141.
- Directory of Wetlands of International Importance. IUCN url p. 541, p. 773.
- Fishery bulletin / U.S. Dept. of Commerce, National Oceanic and Atmospheric Administration, National Marine Fisheries Service. Washington, D.C.: The Service: url p. 1016.
- Functional design of the air ventilation apparatus and overland excurions by teleosts / Karel F. Liem. 37 1987 Chicago: Field Museum of Natural History, 1987. url fig. 14 , fig. 13, page 20, p. 21, p. iii.
- Global venting, midwater, and benthic ecological processes / Michael P. De Luca and Ivar Babb, editors. [Rockville, Md.]: U.S. Dept. of Commerce, National Oceanic and Atmospheric Administration, Oceanic and Atmospheric Research, Office of Undersea Research, [1988] url p. 344.
- IUCN Directory of Afrotropical Protected Areas IUCN, UNEP url p. 1026, p. 675, p. 685, p. 686, p. 697.
- IUCN Directory of Protected Areas in Oceania IUCN url p. 236.
Notes
Contributors
- Bisby, F.A., Y.R. Roskov, M.A. Ruggiero, T.M. Orrell, L.E. Paglinawan, P.W. Brewer, N. Bailly, J. van Hertum, eds (2007). Species 2000 & ITIS Catalogue of Life: 2007 Annual Checklist. Species 2000: Reading, U.K.
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 30, 2012.
- Cambray, J. & Swartz, E. 2007. Oreochromis mossambicus. In: IUCN 2011. IUCN Red List of Threatened Species. Version 2011.2. <www.iucnredlist.org>. Downloadedon 03February2012.
- FishBase 2006.
- FishBase. Release date: January 5, 2010
- Global Biodiversity Information Facility. Accessed February 25, 2007. http://www.gbif.org Mediated distribution data from provider.
- IUCN 2012. IUCN Red List of Threatened Species. Version 2011.2. . Downloaded on January 28, 2012.
- Ruggiero M., Gordon D., Bailly N., Kirk P., Nicolson D. (2011). The Catalogue of Life Taxonomic Classification, Edition 2, Part A. In: Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist (Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D., eds). DVD; Species 2000: Reading, UK.
Data Sources
Accessed through GBIF Data Portal November 15, 2007:
- Cornell University Museum of Vertebrates, Fish Collection
- Institute of Marine and Coastal Sciences, Rutgers University, South African Institute for Aquatic Biodiversity - Fish Collection
- Institute of Marine and Coastal Sciences, Rutgers University, iziko South African Museum - Fish Collection
- Museum national d'histoire naturelle, Ichtyologie
- Museum of Comparative Zoology, Harvard University, MCZ Fish Collection
- OZCAM
- Provider, Online Zoological Collections of Australian Museums
- Royal Ontario Museum, Fish specimens
- Senckenberg, Collection Pisces
- UNIBIO, IBUNAM, CNPE/Coleccion Nacional de Peces
Identifiers
- Biodiversity Heritage Library NamebankID: 133894
- Catalogue of Life Accepted Name Code: Fis-22699
- Fishbase Species ID: 3
- Global Biodiversity Information Facility Taxonkey: 13536119
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 170015
- IUCN ID: 228530
- Natural Heritage Network Species Identifier: AFCQJ10010
- Zipcode Zoo Species Identifier: 17565
Footnotes
- Blaber, S.J.M. (1997). Fish and fisheries of tropical estuaries. Fish and Fisheries Ser. 22, Chapman and Hall, London. 367 p. [back]
- Page, L.M. and B.M. Burr (1991). A field guide to freshwater fishes of North America north of Mexico. Houghton Mifflin Company, Boston. 432 p. [back]
- Robins, C.R., R.M. Bailey, C.E. Bond, J.R. Brooker, E.A. Lachner, R.N. Lea and W.B. Scott (1991). World fishes important to North Americans. Exclusive of species from the continental waters of the United States and Canada. Am. Fish. Soc. Spec. Publ. ( ... [back]
- Frimodt, C. (1995). Multilingual illustrated guide to the world's commercial warmwater fish. Fishing News Books, Osney Mead, Oxford, England. 215 p. [back]
- Otto-Infante, C. (1985). Bio-ecological aspects of the tilapia Sarotherodon mossambicus Peters 1852 (Teleostei, Perciformes, Cichlidae) in Lake Valencia, Venezuela. Acta Cient. Venez. 36(1):68-76. [back]
- Maitipe, P. and S.S. De Silva (1985). Switches between zoophagy, phytophagy and detritivory of Sarotherodon mossambicus (Peters) populations in twelve man-made Sri Lankan lakes. J. Fish Biol. 26:49-61. [back]
- Standard Deviation = 680.190 based on 861 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
- Cambray, J. & Swartz, E. 2007. Oreochromis mossambicus. In: IUCN 2011. IUCN Red List of Threatened Species. Version 2011.2. <www.iucnredlist.org>. Downloaded on 03 February 2012. [back]
